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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L3	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis	Acerodon celebensis		[MSW2] Includes arquatus and Sulawesi specimens formerly in Pteropus argentatus (see Musser et al., 1982<i); also see P. argentatus.; [MSW3] Includes arquatus and Sulawesi specimens formerly in Pteropus argentatus (see Musser et al., 1982a). Also see Flannery (1995b).; [HMW] Pteropus celebensis Peters, 1867 , “ Celebes [= Sulawesi ],” Indonesia . Acerodon celebensis includes Sulawesian specimens formerly identified as Pleropus arquatus and P. argentatus . Monotypic.; [batnames2022] Includes arquatus and Sulawesi specimens formerly included in Pteropus  argentatus (see Musser et al. 1982a). Also see Flannery (1995b).; [IUCN] This species includes arquatus and Sulawesi specimens formerly assigned to Pteropus argentatus (Musser et al . 1982, Flannery 1995).; [batnames2023] Includes arquatus and Sulawesi specimens formerly included in Pteropus  argentatus (see Musser et al. 1982a). Also see Flannery (1995b).; [batnames2025_1.7] Includes arquatus and Sulawesi specimens formerly included in Pteropus argentatus (see Musser et al. 1982a). Also see Flannery (1995b).					(arqua t us)	arquatus.			arquatus			celebensis	celebensis - arquatus	celebensis, arquatus	(arquatus)	celebensis	celebensis - arquatus	celebensis, arquatus	celebensis, arquatus	celebensis	celebensis - arquatus	celebensis (W. C. H. Peters, 1867)|arquatus (G. S. Miller & Hollister, 1921)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Celebes flying fox	Celebes	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Acerodon celebensis	Indonesia, Sulawesi.	Peters	1867	Monatsb. Preuss. Akad. Wiss. Berlin, p. 333.	Distribution: Confined to Ce lebes and surrounding small islands.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Sulawesi flying fox	Sulawesi; ref. 4.16	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1867	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1867:333.	Includes arquatus and Sulawesi specimens formerly in Pteropus argentatus (see Musser et al., 1982<i); also see P. argentatus.	Sulawesi, Saleyer Isl, and Sula Mangoli (Indonesia).	Indonesia, Sulawesi.		PETERS	1867	No antero-internal cusp on the middle lower premolar. Ears relatively long and pointed (29 mm). Dentition relatively weak. Size relatively small (forearm length, 125-142 mm).	Distribution: Confined to Ce lebes and surrounding small islands.	No sub species.		26	species	A. celebensis	PETERS	1867	Acerodon	genus	Acerodon celebensis				No antero-internal cusp on the middle lower premolar. Ears relatively long and pointed (29 mm). Dentition relatively weak. Size relatively small (forearm length, 125-142 mm).	No sub species.		1. A. celebensis (PETERS 1867 ) ( = arquatus MILLER & HOLLISTER 1921).	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Acerodon celebensis	Acerodon		celebensis	Peters	y	1867		Monatsb. K. Preuss. Akad. Wiss. Berlin	1867		333		Sulawesi Fruit Bat	Indonesia, Sulawesi.	Sulawesi, Saleyer Isl, Sangihe Isls, Sula Isls (Indonesia).	CITES – Appendix II. IUCN/SSC Action Plan (1992) – No data. IUCN 2003 – Lower Risk (nt).	arquatus Miller and Hollister, 1921.	Includes arquatus and Sulawesi specimens formerly in Pteropus argentatus (see Musser et al., 1982a). Also see Flannery (1995b).	03AD87FAFF88F66689AE3716F82AFEC7	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	134	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF88F66689AE3716F82AFEC7.xml	Acerodon celebensis	Pteropodidae	Acerodon	celebensis		1867	Acérodon de Sulawesi @fr | Sulawesi-Flughund @de | Zorro volador de Célebes @es | Sulawesi Acerodon @en | Sulawesi Fruit Bat @en	Pteropus celebensis Peters, 1867 , “ Celebes [= Sulawesi ],” Indonesia . Acerodon celebensis includes Sulawesian specimens formerly identified as Pleropus arquatus and P. argentatus . Monotypic.	Endemic to Sulawesi (including nearby Sangihe, Siau, Togian, Buton, and Selayar Is), Banggai Is (Peleng), and Sula Is (Mangole and Sanana).	Head-body 190- 225 mm (tailless), ear 30-34 mm , hindfoot 37-44 mm , forearm 130-145 mm ; weight 374 g (mean). Greatest lengths of skulls are 62-63 mm . Males are larger than females and have glandular neck tufts with strong musk that are rich tawny at base and produce thick ocherous orange band around shoulder. Muzzle of the Sulawesi Flying Fox is short and narrow, rhinarium is black with large nostrils, and philtrum has side furrow. Eyes are large, with brown irises. Ears are longer than muzzle from front of eye to tip of nose, wide at base, and pointed. Pelage on crown, mantle, and underside is generally golden or yellowish buffy, with chocolate-brown backside and small patch of brown fur on throat with sprinkles of longer silvery whitish hairs. Belly is slightly lighter. Fur is short and thin around crown and mantle to expose buffy and brown head and muzzle. Sides of head are sprinkled with long whitish silvery hairs. Golden buffy tips of fur generally hide brown bases. Tibia is dark and not furred; femur is furred. Fur is woollier on underside of lateral membrane between humerus and femur. Genitals are black and concealed in fur. Uropatagium is reduced; calcar is small. Wing membranes are black and originate from sides of body,slightly above medial plane. Skull is typical pteropine and moderately robust, with short narrow rostrum, small orbits, low and broad sagittal crest, and rather thick mandible. Dental formula for most species of Acerodonis 12/2, C1/1,P3/3,M2/3 (x2) =34.1, and last lower molars are small, and P' is early deciduous. Canine cingula and posterior basal ledges on premolars and molars are strong, especially compared with congeners. Caninesare slightly larger in males. Dentition is weak in general; P? lacks posterointernal cusp. Anterointernal basal tubercle is more sharply obvious in M, than in P,, and there is no distinct anterointernal cusp in P,. Basal ledge of P,, M,, and M, do not extend as far forward on innerside ofteeth as in other species of Acerodon .	Coastal mangrove forests on offshore islands and villages and bamboo on coast and inland from sea level up to elevations of ¢. 1500 m .	The Sulawesi Flying Fox eats fruits of native and introduced species. Fruits in diets include multiple native Ficus species ( Moraceae ), Duabanga moluccana ( Lythraceae ), and Palaquium sp. ( Sapotaceae ). It has also been observed visiting durian flowers ( Durio zibethinus, Malvaceae ). Recent studies suggest it is a key pollinator and seed disperser. Diet is not as broad as that of co-occurring species of flying foxes, but it does contain more native fruits.	Reproductive cycle of the Sulawesi Flying Fox is probably seasonal, with births occurring in February-March. Females have been seen with dependent young in August-October.	Sulawesi Flying Foxes are nocturnal and remain in roosts during the day. They engage in typical behaviors such as fanning their wings and territorial defense. Colonies leave roosts a little before dusk (c.19:00 h) to go to foraging sites.	The Sulawesi Flying Fox is generally gregarious and roosts in large colonies in mangroves with low levels of disturbance. Individuals and small groups have been found in areas with higher levels of disturbance (e.g. nearvillages or roadsides), but it is unclearif these roosts are temporary or not. It often co-roosts with other species offlying foxes, particularly the Black Flying Fox ( Pteropus alecto ).	CITES Appendix II. Classified as Vulnerable on The IUCN Red List. The Sulawesi Flying Fox is endemic and increasingly pressured by hunting for bushmeat. It has been locally extirpated from North Sulawesi , where bushmeattrade is centralized, with hunting causing an inferred population reduction greater than 30% in the past three generations. Known colonies are under constantly threat of being extirpated, but there are no legal protections. Mangrove forests have declined due to conversion for shrimp aquaculture. As mangroves disappear, large roosting colonies will probably decrease because they are some of the only habitats left with low levels of human disturbance. Deforestation and logging also threaten stability of foraging sites.	Andersen (1912b) | Bergmans & Rozendaal (1988) | Corbet & Hill (1992) | Flannery (1995a) | Musser et al. (1982) | Tsang & Sheherazade (2016)		130. Sulawesi Flying Fox Acerodon celebensis French: Acérodon de Sulawesi / German: Sulawesi-Flughund / Spanish: Zorro volador de Célebes Other common names: Sulawesi Acerodon , Sulawesi Fruit Bat Taxonomy. Pteropus celebensis Peters, 1867 , “ Celebes [= Sulawesi ],” Indonesia . Acerodon celebensis includes Sulawesian specimens formerly identified as Pleropus arquatus and P. argentatus . Monotypic. Distribution. Endemic to Sulawesi (including nearby Sangihe, Siau, Togian, Buton, and Selayar Is), Banggai Is (Peleng), and Sula Is (Mangole and Sanana). Descriptive notes. Head-body 190- 225 mm (tailless), ear 30-34 mm , hindfoot 37-44 mm , forearm 130-145 mm ; weight 374 g (mean). Greatest lengths of skulls are 62-63 mm . Males are larger than females and have glandular neck tufts with strong musk that are rich tawny at base and produce thick ocherous orange band around shoulder. Muzzle of the Sulawesi Flying Fox is short and narrow, rhinarium is black with large nostrils, and philtrum has side furrow. Eyes are large, with brown irises. Ears are longer than muzzle from front of eye to tip of nose, wide at base, and pointed. Pelage on crown, mantle, and underside is generally golden or yellowish buffy, with chocolate-brown backside and small patch of brown fur on throat with sprinkles of longer silvery whitish hairs. Belly is slightly lighter. Fur is short and thin around crown and mantle to expose buffy and brown head and muzzle. Sides of head are sprinkled with long whitish silvery hairs. Golden buffy tips of fur generally hide brown bases. Tibia is dark and not furred; femur is furred. Fur is woollier on underside of lateral membrane between humerus and femur. Genitals are black and concealed in fur. Uropatagium is reduced; calcar is small. Wing membranes are black and originate from sides of body,slightly above medial plane. Skull is typical pteropine and moderately robust, with short narrow rostrum, small orbits, low and broad sagittal crest, and rather thick mandible. Dental formula for most species of Acerodonis 12/2, C1/1,P3/3,M2/3 (x2) =34.1, and last lower molars are small, and P' is early deciduous. Canine cingula and posterior basal ledges on premolars and molars are strong, especially compared with congeners. Caninesare slightly larger in males. Dentition is weak in general; P? lacks posterointernal cusp. Anterointernal basal tubercle is more sharply obvious in M, than in P,, and there is no distinct anterointernal cusp in P,. Basal ledge of P,, M,, and M, do not extend as far forward on innerside ofteeth as in other species of Acerodon . Habitat. Coastal mangrove forests on offshore islands and villages and bamboo on coast and inland from sea level up to elevations of ¢. 1500 m . Food and Feeding. The Sulawesi Flying Fox eats fruits of native and introduced species. Fruits in diets include multiple native Ficus species ( Moraceae ), Duabanga moluccana ( Lythraceae ), and Palaquium sp. ( Sapotaceae ). It has also been observed visiting durian flowers ( Durio zibethinus, Malvaceae ). Recent studies suggest it is a key pollinator and seed disperser. Diet is not as broad as that of co-occurring species of flying foxes, but it does contain more native fruits. Breeding. Reproductive cycle of the Sulawesi Flying Fox is probably seasonal, with births occurring in February-March. Females have been seen with dependent young in August-October. Activity patterns. Sulawesi Flying Foxes are nocturnal and remain in roosts during the day. They engage in typical behaviors such as fanning their wings and territorial defense. Colonies leave roosts a little before dusk (c.19:00 h) to go to foraging sites. Movements, Home range and Social organization. The Sulawesi Flying Fox is generally gregarious and roosts in large colonies in mangroves with low levels of disturbance. Individuals and small groups have been found in areas with higher levels of disturbance (e.g. nearvillages or roadsides), but it is unclearif these roosts are temporary or not. It often co-roosts with other species offlying foxes, particularly the Black Flying Fox ( Pteropus alecto ). Status and Conservation. CITES Appendix II. Classified as Vulnerable on The IUCN Red List. The Sulawesi Flying Fox is endemic and increasingly pressured by hunting for bushmeat. It has been locally extirpated from North Sulawesi , where bushmeattrade is centralized, with hunting causing an inferred population reduction greater than 30% in the past three generations. Known colonies are under constantly threat of being extirpated, but there are no legal protections. Mangrove forests have declined due to conversion for shrimp aquaculture. As mangroves disappear, large roosting colonies will probably decrease because they are some of the only habitats left with low levels of human disturbance. Deforestation and logging also threaten stability of foraging sites. Bibliography. Andersen (1912b), Bergmans & Rozendaal (1988), Corbet & Hill (1992), Flannery (1995a), Musser et al. (1982), Tsang & Sheherazade (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Acerodon celebensis	Acerodon		celebensis	Peters	1867	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:33:00	Sulawesi Fruit Bat	 arquatus Miller and Hollister, 1921.	Indonesia, Sulawesi.	Sulawesi, Saleyer Isl, Sangihe Isls, Sula Isls (Indonesia).	Appendix II	Vulnerable	Includes arquatus and Sulawesi specimens formerly included in Pteropus  argentatus (see Musser et al. 1982a). Also see Flannery (1995b).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Acerodon celebensis	23	Sulawesi Flying Fox	Sulawesi Acerodon|Sulawesi Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTEROPODINI	Acerodon	NA	celebensis	W. Peters	1867	1	Pteropus_celebensis	Peters, W. C. H. (1867). Ãœber die Flederhunde, Pteropi, und insbesondere über die Arten der Gattung Pteropus s. s. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1867, 333.	https://www.biodiversitylibrary.org/item/112419#page/343/mode/1up	ZMB 4012 [lectotype]		"Celebes [= Sulawesi]," Indonesia.			celebensis (W. Peters, 1867)|arquatus (G. S. Miller & Hollister, 1921)	NA	NA	Indonesia	Asia	Australasia/Oceania	VU	0	0	0	Acerodon_celebensis	0	sciname match	Acerodon_celebensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	137	Acerodon celebensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Acerodon	celebensis	Peters, 1867	This species includes arquatus and Sulawesi specimens formerly assigned to Pteropus argentatus (Musser et al . 1982, Flannery 1995).	20000000	Acerodon celebensis	Vulnerable	A2d	2016	2016-01-08 00:00:00 UTC	3.1	English	This species is endemic to the Sulawesi subregion. It has been locally extirpated from North Sulawesi, the centre of the bushmeat trade at unsustainable rates (Sheherazade and Tsang 2015). Known colonies are under constant threat from hunting pressure, and none have official government protection. Hunting has caused an inferred population reduction of &gt;30% over the past three generations (13.5 years; Pacifici et al.  2013), and is unlikely to stop completely. The species is therefore listed as Vulnerable under criterion A2d. Large colonies primarily occur in mangrove forests, which have drastically declined due to conversion to shrimp aquacultures. Continued hunting and loss of mangroves over the next decade will likely result in the species being uplisted to Endangered.	This bat roosts in trees in villages, and also in bamboo and mangroves. It is commonly found inland and along the coast and shows a medium tolerance of human disturbance. However, large colonies of A. celebensis are only currently known from mangrove forests, roosting with Pteropus alecto on lower branches (S.M. Tsang pers. obs). Individuals are commonly seen feeding on breadfruit trees and coconut (Flannery 1995). Neither of two adult females examined by Flannery in November 1991 were pregnant or lactating, but suggested seasonal breeding, with births occurring in February to March. In south Sulawesi females were observed with pups in August, September, and October (S. Heinrichs pers. comm. 2012).	Acerodon celebensis is commonly hunted and is found in bush meat markets for local and regional trade, which is a major factor in its population decline. They are hunted throughout their entire range and are now locally extirpated in North Sulawesi. Known roost sites are at constant threat from hunting pressures, some sites have been decimated due to over hunting and lack of protection. An example is Soppeng village, where the flying foxes have been hunted and major roosting trees have been cut down (S. Heinrichs pers. comm. 2012). Deforestation due to logging and agriculture is another major threat for this species.	It is likely the population is in decline due to unsustainable levels of hunting (Sheherazade and Tsang 2015). Known roost sites are at constant threat from hunting pressures, some sites have been decimated due to over hunting and lack of protection (S. Heinrichs pers. comm. 2012).	Decreasing	<p>This species is endemic to the Sulawesi subregion, on Sulawesi, Butan, only recorded from Talenge (but likely from all of the Togeran Islands), Mangole, Sanana, Siau, Sangihe, and Selayar, in Indonesia. It is primarily a lowland species, occurring from sea level up to 1,500 m asl.</p>	The species is locally hunted.	Terrestrial	The species is listed on CITES Appendix II. ; No large colonies are located within protected areas nor granted formal protection. The species is locally extirpated in North Sulawesi, the centre of the bushmeat trade. In some areas (such as Watansoppeng in South Sulawesi) they are protected, because local people believe this bat brings good fortune, particularly with rains. However, even in locally protected roosting/foraging areas, outside hunters have been known to take bats for trade in the wildlife market. More studies on the species populations trends are needed, including population size and distribution through its range. Also, needed are studies of the species habitat requirements to better understand the effects of forest conversion and land use change. Finally, studies are needed to determine the amount of hunting of this species and the impact of the bushmeat trade on population persistence. Similar to most threatened flying foxes, local capacity building for conservation managers ; and education and awareness within local communities are sorely needed to support conservation efforts.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Acerodon		celebensis	Peters	1867	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:33:00	Sulawesi Fruit Bat	 arquatus Miller and Hollister, 1921.	Indonesia, Sulawesi.	Sulawesi, Saleyer Isl, Sangihe Isls, Sula Isls (Indonesia).	Appendix II	Vulnerable	Includes arquatus and Sulawesi specimens formerly included in Pteropus  argentatus (see Musser et al. 1982a). Also see Flannery (1995b).	Acerodon celebensis	1004442	23	Sulawesi Flying Fox	Sulawesi Acerodon|Sulawesi Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTEROPODINI	Acerodon	NA	celebensis	W. Peters	1867	1	Pteropus_celebensis	Peters, W. C. H. (1867). Ãœber die Flederhunde, Pteropi, und insbesondere über die Arten der Gattung Pteropus s. s. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1867, 333.	https://www.biodiversitylibrary.org/item/112419#page/343/mode/1up	ZMB 4012 [lectotype]		"Celebes [= Sulawesi]," Indonesia.			celebensis (W. Peters, 1867)|arquatus (G. S. Miller & Hollister, 1921)	NA	NA				Indonesia	Asia	Australasia/Oceania	VU	0	0	0	Acerodon_celebensis	0	sciname match	Acerodon_celebensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Acerodon_celebensis	1004442	23	Sulawesi Flying Fox	Sulawesi Acerodon|Sulawesi Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteropodini	Acerodon	NA	celebensis	W. C. H. Peters	1	Pteropus celebensis	Peters, W.C.H. 1867. Ãœber die Fiederhunde, _Pteropi_, und insbesondere über die Arten der Gattung _Pteropus_ s. s. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1867:319-333.	https://www.biodiversitylibrary.org/page/36510596	ZMB 4012	lectotype		"Celebes [= Sulawesi]," Indonesia.			NA	NA				Indonesia	Asia	Australasia	VU	0	0	0	Acerodon_celebensis	0	sciname match	Acerodon_celebensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Acerodon		celebensis	Peters	1867	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1872:33:00	Sulawesi Fruit Bat	arquatus Miller and Hollister, 1921.	Indonesia, Sulawesi.	Sulawesi, Saleyer Isl, Sangihe Isls, Sula Isls (Indonesia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Appendix II</a>	<a href='https://www.iucnredlist.org/species/137/220501878/' target='_blank'>Vulnerable</a>	Includes arquatus and Sulawesi specimens formerly included in Pteropus argentatus (see Musser et al. 1982a). Also see Flannery (1995b).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Acerodon celebensis; A. celebensis; Acerodon celebensis; Acerodon celebensis; Acerodon celebensis; Acerodon celebensis; Acerodon celebensis; arquatus; arquatus; celebensis; arquatus; Acérodon de Sulawesi; Sulawesi-Flughund; Zorro volador de Célebes; Sulawesi Acerodon; Sulawesi Fruit Bat; Sulawesi Flying Fox; Sulawesi Acerodon; Sulawesi Fruit Bat; Sulawesi Fruit Bat; Sulawesi Fruit Bat; Acerodon celebensis; Acerodon celebensis celebensis; Acerodon celebensis arquatus; Pteropus argentatus; Pteropus celebensis; Pteropus arquatus; Pteropus argentatus; A. celebensis