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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L182	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Coleura kibomalandy	Coleura kibomalandy	Coleura kibomalandy	Coleura kibomalandy	Coleura kibomalandy	Coleura kibomalandy	Coleura kibomalandy	Coleura kibomalandy	Coleura kibomalandy		[HMW] Coleura kibomalandy Goodman et aL, 2012, “ Madagascar : Province d’Antsiranana , Parc National d’Ankarana, 2.2 km ESE d’Amboandriky, Grotte d’Ambatoharanana (Crocodile Cave) , 12.9883°S , 49.0217°E , 20 m asl.” This representative of Coleura was first found on Madagascar in 2004 and originally identified as C. afra by S. M. Goodman and colleagues in 2008. On the basis of new specimens and morphological and molecular data, the Madagascar population was shown to be distinct from C. afra and described as a new endemic species. Monotypic.; [batnames2022] None.; [IUCN] This genus was first found on Madagascar in 2004 and originally identified as Coleura afra (Goodman et al. 2008). On the basis of new specimen material and morphological and molecular data the Madagascar population was shown to be distinct from C. afra and described as a new species endemic to the island (Goodman et al. 2012).; [batnames2023] None.; [batnames2025_1.7] None.														kibomalandy	This genus was first found on Madagascar in 2004 and originally identified as Coleura afra (Goodman et al. 2008). On the basis of new specimen material and morphological and molecular data the Madagascar population was shown to be distinct from C. afra and described as a new species endemic to the island (Goodman et al. 2012).			kibomalandy	kibomalandy			kibomalandy S. M. Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, M. C. Schoeman, Stanley, & E. C. Teeling, 2012						N/A																																								NA																											03D587F2FFC14C0AFF113AAFF5B6F633	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	363	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFC14C0AFF113AAFF5B6F633.xml	Coleura kibomalandy	Emballonuridae	Coleura	kibomalandy	Goodman et al.	2012	Madagascar Sheath-tailed Bat @en | Emballonure àventre blanc @fr | Madagaskar-Freischwanzfledermaus @de | Embalonuro de Madagascar @es | German @en	Coleura kibomalandy Goodman et aL, 2012, “ Madagascar : Province d’Antsiranana , Parc National d’Ankarana, 2.2 km ESE d’Amboandriky, Grotte d’Ambatoharanana (Crocodile Cave) , 12.9883°S , 49.0217°E , 20 m asl.” This representative of Coleura was first found on Madagascar in 2004 and originally identified as C. afra by S. M. Goodman and colleagues in 2008. On the basis of new specimens and morphological and molecular data, the Madagascar population was shown to be distinct from C. afra and described as a new endemic species. Monotypic.	Endemic to NW Madagascar, known only from Ankarana Special Reserve and Tsingy de Namoroka National Park.	Head—body c.63— 64 mm, tail 11-18 mm, ear 12-18 mm, hindfoot 6-9 mm, forearm 48-52 mm; weight 8-4—12-5 g. Long, shaggy dorsal fur of the Madagascar Sheath-tailed Bat is blackish brown. Venter is grayish cream to pure white, often with underfur having gray bases. Wing membranes are light to medium brown, often with relatively large translucent patches. As with other species of Coleura , Madagascar Sheath-tailed Bats completely lack radio-metacarpal and gular throat pouches.	Dry deciduous forests on lowland limestone karst at known elevations of 15—115 m.	The Madagascar Sheath-tailed Bat forages for insects in open areas or close to forest edges.	Female Madagascar Sheath-tailed Bats produce one young perhaps twice a year, as suggested by synchronous lactation in females at Ankarana in early July and late-term pregnancy in November. Adult males in this population had enlarged testes with convoluted epididymides at start of the rainy season in early November.	The Madagascar Sheath-tailed Bat is crepuscular. Limestone caves including those with running water are shelters for roosting by day. It produces LDC echolocation calls, with prominent QCF componentwhere most energy is concentrated in narrow frequency band of second harmonic and small FM portion. Echolocation search calls consist of sound pulse sequences, with first harmonic peak frequencies of 30-6-35-9 kHz, narrow bandwidth, and short durations of 2-5-5-3 milliseconds. Thus, search calls are designed for detection of flying insects at relatively long distances.	Madagascar Sheath-tailed Bats are gregarious, with colonies up to 500 individuals in Grotte d’Ambatoharanana in Ankarana National Park .	Classified as Data Deficient on The IUCN Red List. Current known distribution of the Madagascar Sheath-tailed Bat is within two national parks: Ankarana and Tsingy de Namoroka. Threats and population trends are unknown, and strict protection of known roosts is essential. Although extensive bat surveys have been conducted in the region without finding additional roosts outside the two parks, vast areas of limestone karst remain largely unstudied in western Madagascar where additional local populations might be found in the future.	Goodman (2017a) | Goodman & Ramasindrazana (2013) | Goodman, Cardiff & Ratrimomanarivo (2008) | Goodman, Puechmaille, Hizem et al. (2012)	https://zenodo.org/record/3747958/files/figure.png	31 . Madagascar Sheath-tailed Bat Coleura kibomalandy French: Emballonure à ventre blanc / German : Madagaskar-Freischwanzfledermaus / Spanish: Embalonuro de Madagascar Taxonomy . Coleura kibomalandy Goodman et aL, 2012, “ Madagascar : Province d’Antsiranana , Parc National d’Ankarana, 2.2 km ESE d’Amboandriky, Grotte d’Ambatoharanana (Crocodile Cave) , 12.9883°S , 49.0217°E , 20 m asl.” This representative of Coleura was first found on Madagascar in 2004 and originally identified as C. afra by S. M. Goodman and colleagues in 2008. On the basis of new specimens and morphological and molecular data, the Madagascar population was shown to be distinct from C. afra and described as a new endemic species. Monotypic. Distribution. Endemic to NW Madagascar, known only from Ankarana Special Reserve and Tsingy de Namoroka National Park. Descriptive notes. Head—body c.63— 64 mm, tail 11-18 mm, ear 12-18 mm, hindfoot 6-9 mm, forearm 48-52 mm; weight 8-4—12-5 g. Long, shaggy dorsal fur of the Madagascar Sheath-tailed Bat is blackish brown. Venter is grayish cream to pure white, often with underfur having gray bases. Wing membranes are light to medium brown, often with relatively large translucent patches. As with other species of Coleura , Madagascar Sheath-tailed Bats completely lack radio-metacarpal and gular throat pouches. Habitat . Dry deciduous forests on lowland limestone karst at known elevations of 15—115 m. Food and Feeding . The Madagascar Sheath-tailed Bat forages for insects in open areas or close to forest edges. Breeding . Female Madagascar Sheath-tailed Bats produce one young perhaps twice a year, as suggested by synchronous lactation in females at Ankarana in early July and late-term pregnancy in November. Adult males in this population had enlarged testes with convoluted epididymides at start of the rainy season in early November. Activity patterns. The Madagascar Sheath-tailed Bat is crepuscular. Limestone caves including those with running water are shelters for roosting by day. It produces LDC echolocation calls, with prominent QCF componentwhere most energy is concentrated in narrow frequency band of second harmonic and small FM portion. Echolocation search calls consist of sound pulse sequences, with first harmonic peak frequencies of 30-6-35-9 kHz, narrow bandwidth, and short durations of 2-5-5-3 milliseconds. Thus, search calls are designed for detection of flying insects at relatively long distances. Movements, Home range and Social organization. Madagascar Sheath-tailed Bats are gregarious, with colonies up to 500 individuals in Grotte d’Ambatoharanana in Ankarana National Park . Status and Conservation . Classified as Data Deficient on The IUCN Red List. Current known distribution of the Madagascar Sheath-tailed Bat is within two national parks: Ankarana and Tsingy de Namoroka. Threats and population trends are unknown, and strict protection of known roosts is essential. Although extensive bat surveys have been conducted in the region without finding additional roosts outside the two parks, vast areas of limestone karst remain largely unstudied in western Madagascar where additional local populations might be found in the future. Bibliography. Goodman (2017a), Goodman & Ramasindrazana (2013), Goodman, Cardiff & Ratrimomanarivo (2008), Goodman, Puechmaille, Hizem eta/. (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Coleura kibomalandy	Coleura		kibomalandy	Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, Schoeman, Stanley & Teeling	2012	0	J. Mamm.	93(6): 1446	Madagascar Sheath-tailed bat	None.	Madagascar: Province d&#39;Antsiranana, Parc National d&#39;Ankarana, 2.2 km ESE d&#39;Amboandriky, Grotte d&#39;Ambatoharanana (Crocodile Cave)	N Madagascar.	Not listed.	Data Deficient	None.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Coleura kibomalandy	23	Madagascar Sheath-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	EMBALLONURINAE	EMBALLONURINI	Coleura	NA	kibomalandy	Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, Schoeman, Stanley, & Teeling	2012	0	Coleura_kibomalandy	Goodman, S. M., Puechmaille, S. J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M. C., Stanley, W. T. & E. C. Teeling. (2012). Phylogeny of the Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy, 93(6), 1446.	https://academic.oup.com/jmammal/article/93/6/1440/910878?login=true	FMNH 213598		"Madagascar: Province d'Antsiranana, Parc National d'Ankarana, 2.2 km ESE d'Amboandriky, Grotte d'Ambatoharanana (Crocodile Cave), 12.9883°S, 49.0217°E, 20 m asl."	-12.99	49.022	kibomalandy Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, Schoeman, Stanley, & Teeling, 2012	NA	NA	Madagascar	Africa	Neotropic	DD	0	0	0	Coleura_kibomalandy	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Coleura kibomalandy	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Coleura	kibomalandy	Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, Schoeman, Stanley, &; Teeling 2012	This genus was first found on Madagascar in 2004 and originally identified as Coleura afra (Goodman et al. 2008). On the basis of new specimen material and morphological and molecular data the Madagascar population was shown to be distinct from C. afra and described as a new species endemic to the island (Goodman et al. 2012).	100000000	Coleura kibomalandy	Data Deficient		2017	2016-05-02 00:00:00 UTC	3.1	English	This newly described species is known from two sites in the western half of Madagascar, both within existing protected areas. Even though extensive bat surveys have been conducted in the region of one of these sites, vast areas of limestone karst remain largely unstudied in western Madagascar and we await further field data to properly assess this taxon.	This is a species that uses caves for day roost sites and at both sites; it is known from these are surrounded by dry deciduous forests. It presumably ;forages in more open areas or close to the forest edge.	Threats to this species are currently unknown.	Population size and trend is not known for this species. In July 2004, Cardiff estimated that about 500 individuals of this species occupied the Grotte d’Ambatoharanana in Ankarana.	Unknown	This bat is only known from Ankarana and Namoroka (Madagascar) and across an elevational range from 20 to 115 m (Goodman and Ramasindrazana 2013).		Terrestrial	The two sites this species is known from include the Ankarana National Park and Namoroka National Park.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Coleura		kibomalandy	Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, Schoeman, Stanley & Teeling	2012	0	J. Mamm.	93(6): 1446	Madagascar Sheath-tailed bat	None.	Madagascar: Province d&#39;Antsiranana, Parc National d&#39;Ankarana, 2.2 km ESE d&#39;Amboandriky, Grotte d&#39;Ambatoharanana (Crocodile Cave)	N Madagascar.	Not listed.	Data Deficient	None.	Coleura kibomalandy	1004800	23	Madagascar Sheath-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	EMBALLONURINAE	EMBALLONURINI	Coleura	NA	kibomalandy	Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, Schoeman, Stanley, & Teeling	2012	0	Coleura_kibomalandy	Goodman, S. M., Puechmaille, S. J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M. C., Stanley, W. T. & E. C. Teeling. (2012). Phylogeny of the Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy, 93(6), 1446.	https://academic.oup.com/jmammal/article/93/6/1440/910878?login=true	FMNH 213598		"Madagascar: Province d'Antsiranana, Parc National d'Ankarana, 2.2 km ESE d'Amboandriky, Grotte d'Ambatoharanana (Crocodile Cave), 12.9883°S, 49.0217°E, 20 m asl."	-12.9883	49.0217	kibomalandy Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, Schoeman, Stanley, & Teeling, 2012	NA	NA				Madagascar	Africa	Neotropic	DD	0	0	0	Coleura_kibomalandy	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Coleura_kibomalandy	1004800	23	Madagascar Sheath-tailed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Emballonurinae	Emballonurini	Coleura	NA	kibomalandy	S. M. Goodman, Puechmaille, Friedli-Weyeneth, Gerlach, Ruedi, M. C. Schoeman, Stanley, & E. C. Teeling	0	Coleura kibomalandy	Goodman, S.M., Puechmaille, S.J., Friedli-Weyeneth, N., Gerlach, J., Ruedi, M., Schoeman, M.C., Stanley, W.T. and Teeling, E.C. 2012-12-17. Phylogeny of the Emballonurini (Emballonuridae) with descriptions of a new genus and species from Madagascar. Journal of Mammalogy 93(6):1440-1455.	https://doi.org/10.1644/11-MAMM-A-271.1	FMNH:Mamm:213598	holotype	https://collections-zoology.fieldmuseum.org/catalogue/1964904	"Madagascar: Province d'Antsiranana, Parc National d'Ankarana, 2.2 km ESE d'Amboandriky, Grotte d'Ambatoharanana (Crocodile Cave), 12.9883°S, 49.0217°E, 20 m asl."	-12.9883	49.0217	NA	NA				Madagascar	Africa	Afrotropic	DD	0	0	0	Coleura_kibomalandy	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Coleura		kibomalandy	Goodman, Puechmaille, Friedli-Weyeneth, Gerlach,  Ruedi, Schoeman, Stanley & Teeling	2012	0	J. Mamm.	93(6): 1446	Madagascar Sheath-tailed bat	None.	Madagascar: Province d&#39;Antsiranana, Parc  National d&#39;Ankarana, 2.2 km ESE d&#39;Amboandriky, Grotte  d&#39;Ambatoharanana (Crocodile Cave)	N Madagascar.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/80221085/95642170/' target='_blank'>Data Deficient</a>	None.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Coleura kibomalandy; Coleura kibomalandy; Coleura kibomalandy; Coleura kibomalandy; Coleura kibomalandy; kibomalandy; Madagascar Sheath-tailed Bat; Emballonure àventre blanc; Madagaskar-Freischwanzfledermaus; Embalonuro de Madagascar; German; Madagascar Sheath-tailed Bat; Madagascar Sheath-tailed bat; C. kibomalandy