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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L177	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Coelops frithi	Coelops frithi	Coelops frithii	Coelops frithi	Coelops frithi	Coelops frithii	Coelops frithii	Coelops frithii	Coelops frithii	Coelops frithii	Coelops frithii	Coelops frithii	Coelops frithii	Coelops frithii	Coelops frithii		[MSW3] Reviewed in part by Bates and Harrison (1997). Malay material has not been allocated to subspecies. Sometimes spelled frithi (e.g., Koopman, 1993).; [HMW] Coelops jrithii Blyth, 1848 , “Soonderbuns [= Sunderbans] of Lower Bengal,” India . Taxonomic revision of this species is needed, as diagnostic and geographical limits of each taxon are still unclear. Five subspecies are currendy recognized.; [batnames2022] Reviewed in part by Bates and Harrison (1997). Malay material has not been allocated to subspecies. Sometimes spelled frithi (e.g.,Koopman, 1993).; [IUCN] The genus Coelops requires taxonomic revision; and C. frithii might actually be two species.; [batnames2023] Reviewed in part by Bates and Harrison (1997). Malay material has not been allocated to subspecies. Sometimes spelled frithi (e.g.,Koopman, 1993).; [batnames2025_1.7] Reviewed in part by Bates and Harrison (1997). Malay material has not been allocated to subspecies. Sometimes spelled frithi (e.g.,Koopman, 1993).						bernsteini, formosanus, inflatus, sinicus.	frithi, sinicus, inflatus, formosanus, bernsteini	frithii , bernsteini , formosanus , inflatus , sinicus		frithii, bemsteini, formosanus, inflatus, sinicus		frithii, bernsteini, formosanus, inflatus, sinicus		frithii, bernsteini, formosanus, inflatus, sinicus	The genus Coelops requires taxonomic revision; and C. frithii might actually be two species.	frithii, bernsteini, formosanus, inflatus, sinicus		frithii, bernsteini, formosanus, inflatus, sinicus 	frithii, bernsteinii, formosanus, inflatus, sinicus, frithi, bernsteini 	bernsteini, formosanus, frithii, inflatus, sinicus		frithii E. Blyth, 1848|bernsteinii W. C. H. Peters, 1862|formosanus Horikawa, 1928|inflatus G. S. Miller, 1928|sinicus G. M. Allen, 1928|frithi Corbet & J. Edwards Hill, 1980 [incorrect subsequent spelling]|bernsteini Koopman, 1994 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Tail-less leaf-nosed bat	India – Java, Bali, Taiwan	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Coelops frithi	India, Bengal, Sunderbans.	Blyth	1848	J. Asiat. Soc. Bengal, 17:251.	Distribu tion: Range same as for the genus except for Bor neo and the Philippines.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Tail-less leaf-nosed bat	E India, S China – Java, Bali, Taiwan	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Blyth	1848	J. Asiat. Soc. Bengal, 17:251.			India, Bengal, Sunderbans. distribution; NE India to S China and Vietnam, south to W Malaysia, Taiwan, and Java and Bali.		BLYTH	1848	Lappets projecting from the supplementary leaflets flanking the an terior noseleaf narrow and elongate. Narial part of the anterior noseleaf not depressed nor sharply demarcated from the intermediate noseleaf, the intervening ridges low and indefinite. Posterior part of the upper surface of the rostrum sloping anteriorly, rostral inflations low. A definite recess within the maxillary root of the zygoma. Upper toothrows convergent anteriorly. Posterior lower premolar elongate and narrow. Lower molars with prominent horizontal external cingula. Size medium (forearm length, 35-47 mm).	Distribu tion: Range same as for the genus except for Bor neo and the Philippines.	Five subspecies are here recognized:	C.f. frithi (northeastern India to Thailand), C.f. sinicus (southwestern China), C.f. inflatus (southeastern mainland China to Vietnam and Thailand), C.f. formosanus (Taiwan), C.f. bernsteini (Java, Bali). Malay material has not been allocated to subspecies.	68	species	C. frithi	BLYTH	1848	Coelops	genus	Coelops frithi				Lappets projecting from the supplementary leaflets flanking the an terior noseleaf narrow and elongate. Narial part of the anterior noseleaf not depressed nor sharply demarcated from the intermediate noseleaf, the intervening ridges low and indefinite. Posterior part of the upper surface of the rostrum sloping anteriorly, rostral inflations low. A definite recess within the maxillary root of the zygoma. Upper toothrows convergent anteriorly. Posterior lower premolar elongate and narrow. Lower molars with prominent horizontal external cingula. Size medium (forearm length, 35-47 mm).	Five subspecies are here recognized:		1. C. frithi BLYTH 1848.	1	_C. f. bernsteinii_ Peters, 1862; _C. f. formosanus_ Horikawa, 1928; _C. f. frithii_ Blyth, 1848; _C. f. inflatus_ Miller, 1928; _C. f. sinicus_ Allen, 1928			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Coelops frithii	Coelops		frithii	Blyth		1848		J. Asiat. Soc. Bengal	17		251		East Asian Tailless Leaf-nosed Bat	Bangladesh, Sunderbans.	Bangladesh and NE India to S China, Thailand, Burma, Laos, Vietnam, south to W Malaysia, Sumatra, and Java and Bali; Taiwan.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	bernsteini Peters, 1862; formosanus Horikawa, 1928; inflatus Miller, 1928; sinicus Allen, 1928.	Reviewed in part by Bates and Harrison (1997). Malay material has not been allocated to subspecies. Sometimes spelled frithi (e.g., Koopman, 1993).	03BD87A2C67EA20CFF5BFDBBFB7848A4	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	235	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C67EA20CFF5BFDBBFB7848A4.xml	Coelops frithii	Hipposideridae	Coelops	frithii	Blyth	1848	Phyllorhine de rith @fr | Ostasiatische Schwanzlose Blattnase @de | Coelops de rith @es | East Asian Tailless Roundleaf Bat @en | Tailless Leaf-nosed Bat @en	Coelops jrithii Blyth, 1848 , “Soonderbuns [= Sunderbans] of Lower Bengal,” India . Taxonomic revision of this species is needed, as diagnostic and geographical limits of each taxon are still unclear. Five subspecies are currendy recognized.	C. f. jrithii Blyth, 1848 -NE India , Bangladesh , Myanmar , and possibly N Thailand . C. f. bemsteini Peters, 1862 — Sumatra , Java , and Bali . C.f. formosanus Horikawa, 1928 — Taiwan I. C. f. inflatus G. S. Miller, 1928 -SE & S China ( Fujian , Guangdong , Guangxi , and Hainan I), Thailand , Laos , Vietnam , and Cambodia . C. f. sinicus G. M. Allen, 1928 — C China ( Sichuan ) Specimens from Malay Peninsula are not yet assigned to any subspecies.	Head—body 38-50 mm, forearm 34 44 mm ; weight 3-7 g. Ears of the East Asian Tailless Leaf-nosed Bat are very large and rounded with antitragus at base. Noseleaf is distinct, with a large anterior leaf that covers muzzle, and divided by deep median emargination. There are two well developed supplementary lappets, rounded and elongated, projecting forwards under anterior leaf and extending beyond muzzle. Intermediate leaf possesses moderately developed median process. Pelage is long and soft, dark gray, or brown to chestnut-brown on dorsal side; ventral side is paler, brownish or ashy gray. Tail and interfemoral membrane are poorly developed. Skull is small; braincase is bulbous. In contrast to Hipposideros species , rostral swellings are flattened. Zygoma is without dorsal process. C1 is large, with well-developed posterior cusp. P2 is small and lies within the tooth row. Baculum is very small, with a bilobed base and slightly expanded tip. Dental formula for all members of Coelops is I 1/2, C 1/1, P 2/2, M 3/3 (x2) = 30. Chromosomal complement has 2n = 30 and FN = 56.	Associated with forest habitats. Recorded from sea level up to 1370 m .	The East Asian Tailless Leaf-nosed Bat is insectivorous. Its highfrequency echolocation suggests that its prey is probably small insects, captured within cluttered space of forest habitats; prey also captured very close to water surface.	No information.	The East Asian Tailless Leaf-nosed Bat roosts in caves or hollow trees. Echolocation is of very high frequency, 350 kHz.	East Asian Tailless Leaf-nosed Bats are known to roost in small colonies of 16 individuals or less.	Classified as Least Concern on T/ie IUCN Red List. The East Asian Tailless Leaf-nosed Bat is a relatively widespread species but rather rare.	Bates, Bumrungsri, Francis, Csorba & Molur (2008) | Bogdanowicz & Owen (1998) | Corbet & Hill (1992) | Francis (2008a) | Lekagul & McNeely (1988) | Simmons (2005)	https://zenodo.org/record/3749079/files/figure.png	21. East Asian Tailless Leaf-nosed Bat Coelops frithii French: Phyllorhine de rith / German: Ostasiatische Schwanzlose Blattnase / Spanish: Coelops de rith Other common names: East Asian Tailless Roundleaf Bat , Tailless Leaf-nosed Bat Taxonomy. Coelops jrithii Blyth, 1848 , “Soonderbuns [= Sunderbans] of Lower Bengal,” India . Taxonomic revision of this species is needed, as diagnostic and geographical limits of each taxon are still unclear. Five subspecies are currendy recognized. Subspecies and Distribution. C. f. jrithii Blyth, 1848 -NE India , Bangladesh , Myanmar , and possibly N Thailand . C. f. bemsteini Peters, 1862 — Sumatra , Java , and Bali . C.f. formosanus Horikawa, 1928 — Taiwan I. C. f. inflatus G. S. Miller, 1928 -SE & S China ( Fujian , Guangdong , Guangxi , and Hainan I), Thailand , Laos , Vietnam , and Cambodia . C. f. sinicus G. M. Allen, 1928 — C China ( Sichuan ) Specimens from Malay Peninsula are not yet assigned to any subspecies. Descriptive notes. Head—body 38-50 mm, forearm 34 44 mm ; weight 3-7 g. Ears of the East Asian Tailless Leaf-nosed Bat are very large and rounded with antitragus at base. Noseleaf is distinct, with a large anterior leaf that covers muzzle, and divided by deep median emargination. There are two well developed supplementary lappets, rounded and elongated, projecting forwards under anterior leaf and extending beyond muzzle. Intermediate leaf possesses moderately developed median process. Pelage is long and soft, dark gray, or brown to chestnut-brown on dorsal side; ventral side is paler, brownish or ashy gray. Tail and interfemoral membrane are poorly developed. Skull is small; braincase is bulbous. In contrast to Hipposideros species , rostral swellings are flattened. Zygoma is without dorsal process. C1 is large, with well-developed posterior cusp. P2 is small and lies within the tooth row. Baculum is very small, with a bilobed base and slightly expanded tip. Dental formula for all members of Coelops is I 1/2, C 1/1, P 2/2, M 3/3 (x2) = 30. Chromosomal complement has 2n = 30 and FN = 56. Habitat. Associated with forest habitats. Recorded from sea level up to 1370 m . Food and Feeding. The East Asian Tailless Leaf-nosed Bat is insectivorous. Its highfrequency echolocation suggests that its prey is probably small insects, captured within cluttered space of forest habitats; prey also captured very close to water surface. Breeding. No information. Activity patterns. The East Asian Tailless Leaf-nosed Bat roosts in caves or hollow trees. Echolocation is of very high frequency, 350 kHz. Movements, Home range and Social organization. East Asian Tailless Leaf-nosed Bats are known to roost in small colonies of 16 individuals or less. Status and Conservation. Classified as Least Concern on T/ie IUCN Red List. The East Asian Tailless Leaf-nosed Bat is a relatively widespread species but rather rare. Bibliography. Bates, Bumrungsri, Francis, Csorba & Molur (2008), Bogdanowicz & Owen (1998), Corbet & Hill (1992), Francis (2008a), Lekagul & McNeely (1988), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Coelops frithii	Coelops		frithii	Blyth	1848	0	J. Asiat. Soc. Bengal	0.8826	East Asian Tailless Leaf-nosed Bat	<b> bernsteini </b>Peters, 1862;<b> formosanus </b>Horikawa, 1928;<b> inflatus </b>Miller, 1928; <b> sinicus </b>Allen, 1928.	Bangladesh, Sunderbans.	Nepal, NE India, and Bangladesh to S China, Thailand, Burma, Laos, Vietnam, south to W Malaysia, Sumatra, and Java and Bali; Taiwan	Not listed.	Near Threatened	Reviewed in part by Bates and Harrison (1997). Malay material has not been allocated to subspecies. Sometimes spelled frithi (e.g.,Koopman, 1993).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Coelops frithii	23	East Asian Tailless Leaf-nosed Bat	East Asian Tailless Roundleaf Bat|Tailless Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Coelops	NA	frithii	Blyth	1848	0						"Soonderbuns [= Sunderbans] of Lower Bengal," India.			frithii Blyth, 1848|bernsteini W. Peters, 1862|formosanus Horikawa, 1928|inflatus G. S. Miller, 1928|sinicus G. M. Allen, 1928	NA	NA	India|Bangladesh|Myanmar|China|Taiwan|Thailand|Laos|Vietnam|Cambodia|Malaysia|Indonesia	Asia	Indomalaya	NT	0	0	0	Coelops_frithii	0	sciname match	Coelops_frithii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	5074	Coelops frithii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Coelops	frithii	Blyth, 1848	The genus Coelops requires taxonomic revision; and C. frithii might actually be two species.	20000000	Coelops frithii	Near Threatened	A4c	2019	2018-08-27 00:00:00 UTC	3.1	English	The species is listed as Near Threatened, approaching criterion A4c, because the global population has likely declined at a rate between 20-25% over the past five years due to a decline in its extent of occurrence (EOO), area of occupancy (AOO), and the loss and disturbance of forest and cave systems across its range. The declines are expected to continue for at least another 10 years (three generations in total; GL = 5 years, Pacifici et al. 2013).	The species has been recorded from various types of forest, including karst forest, lowland secondary forest, subtropical forest, submontane forest, and mangrove (Molur et al. 2002, Lee et al. 2007, Bates et al. 2008, Furey et al. 2010, Loi and Thong). It has also been observed flying near surface of backwater near forest (Abramov et al. 2009).  It is primarily a cave-dweller and uses several types of roosting habitats such as cave, tree hollow (Molur et al. 2002), man-made tunnel, abandoned mining cave (Fang and Cheng 2011), and beneath of big rock (Tsai, W.-Y. pers. comm.). In Taiwan, roost sites are highly associated with forest. It has been recorded using the same caves with Rhinolophus monoceros , R. formosae , Hipposideros armiger , Myotis laniger and Miniopterus fuliginosus (Fang and Cheng 2011). In Vietnam, this species is known from the same cave with Hipposideros alongensis (Thong et al. 2012) and with Rhinolophis affinis , Rh. pusillus , Hipposideros larvatus and Myotis muricola in other caves (Son, unpublished data). In Indonesia, this species roosts in the cave, together with Hipposideros larvatus (Wiantoro, S. pers. Com.). Several maternity colonies with 100-200 mature individuals have been reported from artificial caves and abandoned tunnels in mountain areas with elevation up to 900m asl (Ho, Y.-Y. and Huang, J. C.-C., unpub. data).  Pregnant females were observed between March to May and gave birth of one offspring in May and June. Young bats were able to fly by 40 days after the birth. No evidence shows it hibernates or torpors in a year. In Taiwan, the species remains active in winter, when most of the other cave bat species hibernate (Fang and Cheng 2011).  It is generally a lowland species, of which most occurrence reports were from elevation less than 1,000 m asl. The species is found foraging mainly in clutter vegetation close to the ground in forest understory (Fang, Y.-P. pers. com.), which is likely associated with its slow flight (Lai 2000) and low duty-cycled echolocation strategy (Ho et al. 2013). In Taiwan, the species has been reported to feed mainly on spiders and some insects, including adult lepidopterans, coleopterans, dipterans and homopterans (Fang and Cheng 2011).	Overall the species is most likely threatened by habitat loss and degradation associated with ongoing deforestation and disturbances to roosts in large area of its distribution range in Southeast Asia, South Asia and China. Declining of population is expected at the global scale. In Taiwan, decreasing gene flows among geographic populations with increasing geographic distance were reported (Fang and Cheng 2011), suggesting the species is a poor disperser. In account of the association with forest and slow flight behavior, it is presumably vulnerable to loss and fragmentation of forest. The expansion of road systems could also threat the species. Two roadkill incidences are reported from lowland in Taiwan where forest is highly fragmented by road systems (<a rel="nofollow" target="_blank" href="https://roadkill.tw/">https://roadkill.tw/</a>). Roost disturbances and destruction likely threaten the species. Loss of roost sites, especially large maternity roosts, is expected to interrupt gene flows among the nearby colonies. Light pollution might be also a threat since individuals are observed avoiding artificial light by cave entrance (Huang, J. C.-C., pers. obs.). In South Asia, some populations are threatened by deforestation, generally resulting from logging operations and the conversion of land to agriculture and settlements especially in the Indian northeast (Molur et al. 2002). In Bangladesh, cyclones represent an additional threat to the habitat. In Indonesia, especially Java Island, limestone mining is become the major threat on their roosting sites (Wiantoro S, pers. obs).	The species is thought to undergoing a past and continuing decline across its range. Although the species is widespread, it does not appear to be common. This may indicate its natural rarity or reflect difficulties in surveying the species. It is usually found in groups of less than 20 or fewer individuals in South Asia (Molur et al.   2002), Thailand (Bates et al.   2008) and Taiwan (Fang and Cheng 2011), except a few colonies of 50~350 animals in Taiwan (Fang and Cheng 2011, Ying-Yi Ho pers. obs.).	Decreasing	This widespread species ranges from South Asia eastwards into southeastern China, Taiwan and Southeast Asia. In South Asia this species is presently known from Bangladesh Sunderbans (Khulna division) and India (Meghalaya and West Bengal) (Molur et al. 2002). In China, it has been recorded from the provinces of Fujian, Guangdong, Guangxi, Hainan, Yunnan, Sichuan and the island of Hainan (Smith and Xie 2013). In Southeast Asia it ranges from Myanmar, through all countries in the region, to Thailand and Indonesia (including the islands of Sumatra, Java and Bali). The presence of this species in Cambodia is previously uncertain (Bates et al. 2008), but recently confirmed from the northern part of the country (Sarak et al. 2013). In Taiwan, it has been recorded from elevations ranging from lowland to 1900 m asl, while most known records were from below 300 m (Fang and Cheng 2011). In Vietnam, it is recorded from lowland to 700 m asl, however, they are found from the karst areas of the north and center areas (Kruskop 2013, Loi and Thong 2017). In Indonesia, this species has been recorded from two limestone areas, which are Tuban karst area in East Java and Gunungsewu karst area in Central Java.		Terrestrial	The species is considered vulnerable and protected in China (Smith and Xie 2013) and Taiwan (Cheng et al 2017), most likely due to its rarity. It has been recorded from several protected areas in Southeast Asia and Taiwan. It might be present in protected areas in South Asia and China. Within South Asia there is a need for programmes to increase public awareness about the species and bats in general (Molur et al. 2002). Protection of roost sites and the surrounding forest, especially for those support large maternity colonies and connect different subpopulations, are critical to maintain the population and gene diversity for the species. Additional research to monitor the key roosts, understand the influences of roost disturbances, habitat degradation and development of road systems are needed.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Coelops		frithii	Blyth	1848	0	J. Asiat. Soc. Bengal	0.882639	East Asian Tailless Leaf-nosed Bat	<b> bernsteini </b>Peters, 1862;<b> formosanus </b>Horikawa, 1928;<b> inflatus </b>Miller, 1928; <b> sinicus </b>Allen, 1928.	Bangladesh, Sunderbans.	Nepal, NE India, and Bangladesh to S China, Thailand, Burma, Laos, Vietnam, south to W Malaysia, Sumatra, and Java and Bali; Taiwan	Not listed.	Near Threatened	Reviewed in part by Bates and Harrison (1997). Malay material has not been allocated to subspecies. Sometimes spelled frithi (e.g.,Koopman, 1993).	Coelops frithii	1004563	23	East Asian Tailless Leaf-nosed Bat	East Asian Tailless Roundleaf Bat|Tailless Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Coelops	NA	frithii	Blyth	1848	0						"Soonderbuns [= Sunderbans] of Lower Bengal," India.			frithii Blyth, 1848|bernsteini W. Peters, 1862|formosanus Horikawa, 1928|inflatus G. S. Miller, 1928|sinicus G. M. Allen, 1928	NA	NA				India|Bangladesh|Myanmar|China|Taiwan|Thailand|Laos|Vietnam|Cambodia|Malaysia|Indonesia	Asia	Indomalaya	NT	0	0	0	Coelops_frithii	0	sciname match	Coelops_frithii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Coelops_frithii	1004563	23	East Asian Tailless Roundleaf Bat	East Asian Tailless Roundleaf Bat|Tailless Leaf-nosed Bat|East Asian Tailless Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Coelops	NA	frithii	E. Blyth	0	Coelops Frithii	Blyth, E. 1848. Report of Curator Zoological Department. Journal of the Asiatic Society of Bengal 17(1):247-255.	https://www.biodiversitylibrary.org/page/40129684		holotype		"Soonderbuns [= Sunderbans] of Lower Bengal," India.			NA	NA				India|Bangladesh|Myanmar|China|Taiwan|Thailand|Laos|Vietnam|Cambodia|Malaysia|Indonesia	Asia	Indomalaya	NT	0	0	0	Coelops_frithii	0	sciname match	Coelops_frithii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Coelops		frithii	Blyth	1848	0	J. Asiat. Soc. Bengal	0.882639	East Asian Tailless Leaf-nosed Bat	bernsteini Peters, 1862; formosanus Horikawa, 1928; inflatus Miller, 1928; sinicus Allen, 1928.	Bangladesh, Sunderbans.	Nepal, NE India, and Bangladesh to S China, Thailand, Burma, Laos, Vietnam, south to W Malaysia, Sumatra, and Java and Bali; Taiwan	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/5074/22030377/' target='_blank'>Near Threatened</a>	Reviewed in part by Bates and Harrison (1997). Malay material has not been allocated to subspecies. Sometimes spelled frithi (e.g.,Koopman, 1993).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Coelops frithii; Coelops frithii; Coelops frithii; Coelops frithii; Coelops frithii; Coelops frithii; frithii ; bernsteini ; formosanus ; inflatus ; sinicus; frithii; bemsteini; formosanus; inflatus; sinicus; bernsteini; formosanus; inflatus; sinicus; frithii; bernsteini; formosanus; inflatus; sinicus; Phyllorhine de rith; Ostasiatische Schwanzlose Blattnase; Coelops de rith; East Asian Tailless Roundleaf Bat; Tailless Leaf-nosed Bat; East Asian Tailless Leaf-nosed Bat; East Asian Tailless Roundleaf Bat; Tailless Leaf-nosed Bat; East Asian Tailless Leaf-nosed Bat; East Asian Tailless Leaf-nosed Bat; Coelops frithi ; C. frithii