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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L171	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Eptesicus bottae [synonym of]	N/A	Eptesicus bottae [synonym of]	Eptesicus bottae ognevi	Eptesicus bottae ognevi	Eptesicus ognevi	Eptesicus ognevi	Eptesicus ognevi	Eptesicus ognevi	Eptesicus ognevi	Eptesicus ognevi	Cnephaeus ognevi	Cnephaeus ognevi	Cnephaeus ognevi		[HMW] Eptesicus ognevi Bobrinski, 1918 , Bokhara district, Soviet Turkestan . Eptesicus ognevi is generally recognized as a subspecies of E. bottae , but genetic data support recognition of them as distinct. Morphological data are less conclusive because LE. ognevi greatly resembles E. bottae . Phylogenetic relationship of E. ognevi to other species of Eptesicus is still unresolved. Using mitochondrial genes, E. ognevi was sister to a clade including E. serotinus (specifically subspecies mirza and turcomanus) and E. bottae , but nuclear genes showed it to be sister to a clade including E. hottentotus , E. anatolicus , and E. bottae . It consistently formed a monophyletic group separate from E. bottae , supporting its status as a distinct species. Nevertheless, I. V. Artyushin and colleagues in 2018 found a similarly conflicting dataset that supported the traditional view that E. bottae includes E. ognevi and E. anatolicus based on nuclear genes. Eptesicus ognevi and E. anatolicus are included as distinct species here, pending a more complete genetic and morphological study encompassing the full distribution of all taxa in E. bottae sensu lato . Specimens in Afghanistan and Pakistan need to be examined further because they occur at much higher elevations than most other specimens reported for E. ognevi . Distribution of E. ognevi might overlap with E. anatolicus in the southern Caucasus. Monotypic.; [batnames2022] Subgenus Cnephaeus . Distinct from bottae; see Juste et al. (2012). May include additional cryptic forms - see Juste et al. (2012).; [MDD2022] split from E. bottae; [IUCN] <p>Previously included in ;E. ;<span class="SpellE">bottae , this taxon ;has now been raised</span> ;to species level (see ;Juste ;et al. 2013, ;Artyushin</span> ;et al. 2018).</p>; [batnames2023] Subgenus Cnephaeus . Distinct from bottae; see Juste et al. (2012). May include additional cryptic forms - see Juste et al. (2012).; [MDD2023] split from E. bottae; [MDD2025_2.0] split from E. bottae; moved from Eptesicus to Cnephaeus; [batnames2025_1.7] Distinct from bottae; see Juste et al. (2012). May include additional cryptic forms - see Juste et al. (2012).; [MDD2025_2.2] split from E. bottae; moved from Eptesicus to Cnephaeus														ognevi	<p>Previously included in ;E. ;<span class="SpellE">bottae , this taxon ;has now been raised</span> ;to species level (see ;Juste ;et al. 2013, ;Artyushin</span> ;et al. 2018).</p>			ognevi	ognevi			ognevi (Bobrinski, 1918)						N/A																																								NA																											4C3D87E8FFA16A1EFF4A97B51C0AB09C	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	848	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFA16A1EFF4A97B51C0AB09C.xml	Eptesicus ognevi	Vespertilionidae	Eptesicus	ognevi	Bobrinski	1918	Sérotine d'Ognev @fr | Ognev-Breitfligelfledermaus @de | Eptesicus de Ognev @es	Eptesicus ognevi Bobrinski, 1918 , Bokhara district, Soviet Turkestan . Eptesicus ognevi is generally recognized as a subspecies of E. bottae , but genetic data support recognition of them as distinct. Morphological data are less conclusive because LE. ognevi greatly resembles E. bottae . Phylogenetic relationship of E. ognevi to other species of Eptesicus is still unresolved. Using mitochondrial genes, E. ognevi was sister to a clade including E. serotinus (specifically subspecies mirza and turcomanus) and E. bottae , but nuclear genes showed it to be sister to a clade including E. hottentotus , E. anatolicus , and E. bottae . It consistently formed a monophyletic group separate from E. bottae , supporting its status as a distinct species. Nevertheless, I. V. Artyushin and colleagues in 2018 found a similarly conflicting dataset that supported the traditional view that E. bottae includes E. ognevi and E. anatolicus based on nuclear genes. Eptesicus ognevi and E. anatolicus are included as distinct species here, pending a more complete genetic and morphological study encompassing the full distribution of all taxa in E. bottae sensu lato . Specimens in Afghanistan and Pakistan need to be examined further because they occur at much higher elevations than most other specimens reported for E. ognevi . Distribution of E. ognevi might overlap with E. anatolicus in the southern Caucasus. Monotypic.	Georgia , Armenia , Azerbaijan , NW & NE Iran , S Kazakhstan , Uzbekistan , Turkmenistan , W Kyrgyzstan , Tajikistan , NW Afghanistan , N Pakistan , and NW India ( Jammu and Kashmir ).	Head—body 62-65 mm, tail 45-52 mm, ear 17-9-20 mm, hindfoot 7mm (one specimen), forearm 42-5-44-6 mm. Ognev’s Serotineis externally undistinguishable from Botta’s Serotine ( E. bottae ). Dorsal pelage is light to dark creamy buff, with grayish beige to rusty brown tinge (hairs tricolored, with dark bases); ventral pelage is creamy to grayish beige (hairs bicolored, with white tips and pale grayish brown bases). Bare muzzle, ears, and membranes are pale grayish brown to dark brown. Ears are relatively short and subtriangular, with rounded tips; tragus is about one-half the ear length and is the same width from base to rounded tip. Wings and uropatagium are semi-translucent, and tail protrudes c.3-5 mm past uropatagium; postcalcarial lobe is well developed. Craniodental features are similar to Botta’s Serotine overall but can be distinguished by some slight differences, such as having relatively longer and narrowerrostral part of skull and relatively narrower rostrum in its mesial and distal parts. Chromosomal complement has 2n = 50 and FNa = 48 (Central Asia).	Variety of arid and semiarid deserts and steppe in primarily lowland areas, recorded at elevations of 895 m in Afghanistan and at 3200 m in Pakistan (typically found at lower elevations).	Ognev’s Serotine captures various insects by slow hawking. Seventeen fecal samples from Khorasan , Iran , contained Hymenoptera (mainly Formicoidea with some Ichneumonoidea), Coleoptera ( Carabidae and Elateridae ), Heteroptera, Auchenorrhyncha, Lepidoptera , and Neuroptera ( Hemerobiidae and Chrysopidae ).	In Iran , a pregnant Ognev’s Serotine with two well-developed fetuses was caught in mid-May, suggesting that births occur in late May or early June. In Turkmenistan , births reportedly occur from mid-May to early June, and in south-eastern Kazakhstan , a pregnant female with two fetuses was caught in early and late May. Pregnancy and births occur weeks later in Azerbaijan where pregnant females are known from mid-May to mid-June.	Ognev’s Serotine is nocturnal and probably roosts in rock crevices.	No information.	Not assessed on The IUCN Red List. Ognev’s Serotine 1s included under Botta’s Serotine, which is classified as Least Concern. Ognev’s Serotine is widespread but does not seem to be very common; little 1s known about its threats and ecology.	Artyushin et al. (2018) | Aulagnier et al. (2008) | Bates & Harrison (1997) | Benda & Gaisler (2015) | Benda, Dietz et al. (2008) | Benda, Faizolahi et al. (2012) | Benda, Hank & Cerveny (2011) | Hanak & Gaisler (1971) | Zima & Horadek (1981)	https://zenodo.org/record/6398210/files/figure.png	191. Ognev’s Serotine Eptesicus ognevi French: Sérotine d'Ognev / German: Ognev-Breitfligelfledermaus / Spanish: Eptesicus de Ognev Taxonomy. Eptesicus ognevi Bobrinski, 1918 , Bokhara district, Soviet Turkestan . Eptesicus ognevi is generally recognized as a subspecies of E. bottae , but genetic data support recognition of them as distinct. Morphological data are less conclusive because LE. ognevi greatly resembles E. bottae . Phylogenetic relationship of E. ognevi to other species of Eptesicus is still unresolved. Using mitochondrial genes, E. ognevi was sister to a clade including E. serotinus (specifically subspecies mirza and turcomanus) and E. bottae , but nuclear genes showed it to be sister to a clade including E. hottentotus , E. anatolicus , and E. bottae . It consistently formed a monophyletic group separate from E. bottae , supporting its status as a distinct species. Nevertheless, I. V. Artyushin and colleagues in 2018 found a similarly conflicting dataset that supported the traditional view that E. bottae includes E. ognevi and E. anatolicus based on nuclear genes. Eptesicus ognevi and E. anatolicus are included as distinct species here, pending a more complete genetic and morphological study encompassing the full distribution of all taxa in E. bottae sensu lato . Specimens in Afghanistan and Pakistan need to be examined further because they occur at much higher elevations than most other specimens reported for E. ognevi . Distribution of E. ognevi might overlap with E. anatolicus in the southern Caucasus. Monotypic. Distribution. Georgia , Armenia , Azerbaijan , NW & NE Iran , S Kazakhstan , Uzbekistan , Turkmenistan , W Kyrgyzstan , Tajikistan , NW Afghanistan , N Pakistan , and NW India ( Jammu and Kashmir ). Descriptive notes. Head—body 62-65 mm, tail 45-52 mm, ear 17-9-20 mm, hindfoot 7mm (one specimen), forearm 42-5-44-6 mm. Ognev’s Serotineis externally undistinguishable from Botta’s Serotine ( E. bottae ). Dorsal pelage is light to dark creamy buff, with grayish beige to rusty brown tinge (hairs tricolored, with dark bases); ventral pelage is creamy to grayish beige (hairs bicolored, with white tips and pale grayish brown bases). Bare muzzle, ears, and membranes are pale grayish brown to dark brown. Ears are relatively short and subtriangular, with rounded tips; tragus is about one-half the ear length and is the same width from base to rounded tip. Wings and uropatagium are semi-translucent, and tail protrudes c.3-5 mm past uropatagium; postcalcarial lobe is well developed. Craniodental features are similar to Botta’s Serotine overall but can be distinguished by some slight differences, such as having relatively longer and narrowerrostral part of skull and relatively narrower rostrum in its mesial and distal parts. Chromosomal complement has 2n = 50 and FNa = 48 (Central Asia). Habitat. Variety of arid and semiarid deserts and steppe in primarily lowland areas, recorded at elevations of 895 m in Afghanistan and at 3200 m in Pakistan (typically found at lower elevations). Food and Feeding. Ognev’s Serotine captures various insects by slow hawking. Seventeen fecal samples from Khorasan , Iran , contained Hymenoptera (mainly Formicoidea with some Ichneumonoidea), Coleoptera ( Carabidae and Elateridae ), Heteroptera, Auchenorrhyncha, Lepidoptera , and Neuroptera ( Hemerobiidae and Chrysopidae ). Breeding. In Iran , a pregnant Ognev’s Serotine with two well-developed fetuses was caught in mid-May, suggesting that births occur in late May or early June. In Turkmenistan , births reportedly occur from mid-May to early June, and in south-eastern Kazakhstan , a pregnant female with two fetuses was caught in early and late May. Pregnancy and births occur weeks later in Azerbaijan where pregnant females are known from mid-May to mid-June. Activity patterns. Ognev’s Serotine is nocturnal and probably roosts in rock crevices. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN Red List. Ognev’s Serotine 1s included under Botta’s Serotine, which is classified as Least Concern. Ognev’s Serotine is widespread but does not seem to be very common; little 1s known about its threats and ecology. Bibliography. Artyushin et al. (2018), Aulagnier et al. (2008), Bates & Harrison (1997), Benda & Gaisler (2015), Benda, Dietz et al. (2008), Benda, Faizolahi et al. (2012), Benda, Hank & Cerveny (2011), Hanak & Gaisler (1971), Zima & Horadek (1981).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Eptesicus ognevi	Eptesicus	Cnephaeus	ognevi	Bobrinskii	1918	0	In: Materials for the Knowledge of the Fauna and Flora of the Russian Empire: Zoological Department	15:12	Ognev's Serotine	None.		Jammu and Kasmir (India).	Not listed.	Least Concern	Subgenus Cnephaeus . Distinct from bottae; see Juste et al. (2012). May include additional cryptic forms - see Juste et al. (2012).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Eptesicus ognevi	23	Ognev's Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Eptesicus	Cnephaeus	ognevi	Bobrinski	1918	0	Eptesicus_ognevi		https://zmmu.msu.ru/dbs/pdf/bobrinskoi1915.pdf	ZMMU S-96374		Bokhara district, Soviet Turkestan.			ognevi Bobrinski, 1918	split from E. bottae	Juste, J., Benda, P., Garcia-Mudarra, J. L., & Ibanez, C. (2013). Phylogeny and systematics of Old World serotine bats (genus Eptesicus, Vespertilionidae, Chiroptera): an integrative approach. Zoologica Scripta, 42(5), 441-457.	Georgia|Armenia|Azerbaijan|Iran|Kazakhstan|Uzbekistan|Turkmenistan|Kyrgyzstan|Tajikistan|Afghanistan|Pakistan|India	Asia	Palearctic	LC	0	0	0	Eptesicus_ognevi	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Eptesicus ognevi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Eptesicus	ognevi	Bobrinskii, 1918	<p>Previously included in ;E. ;<span class="SpellE">bottae , this taxon ;has now been raised</span> ;to species level (see ;Juste ;et al. 2013, ;Artyushin</span> ;et al. 2018).</p>	90000000	Eptesicus ognevi	Least Concern		2020	2020-07-05 00:00:00 UTC	3.1	English	This species ;is assessed ;as Least Concern due to its wide distribution and no known major threats. In its eastern range, the species is naturally not abundant.	Found in a wide range of arid and semi-arid habitats including steppe in lowland areas and ;rocky mountains. A crevice-dwelling species, it inhabits buildings, ruins (including tombs), and natural rock crevices throughout the year.	Little is known about threats to the species; ;however ;the habitats in some parts of its range are degrading. As this species lives in crevices in ruins, tourism-related developmental activities and disturbance will have negative impact on their populations.	Appears to be generally rare to uncommon although reported as locally common.	Unknown	A palaearctic species, occurring in central Asia to India.	Nothing ;is known ;about its use and trade.	Terrestrial		Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Eptesicus	Cnephaeus	ognevi	Bobrinskii	1918	0	In: Materials for the Knowledge of the Fauna and Flora of the Russian Empire: Zoological Department	15:12	Ognev's Serotine	None.		Jammu and Kasmir (India).	Not listed.	Least Concern	Subgenus Cnephaeus . Distinct from bottae; see Juste et al. (2012). May include additional cryptic forms - see Juste et al. (2012).	Eptesicus ognevi	1005527	23	Ognev's Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Eptesicus	Cnephaeus	ognevi	Bobrinski	1918	0	Eptesicus_ognevi		https://zmmu.msu.ru/dbs/pdf/bobrinskoi1915.pdf	ZMMU S-96374		Bokhara district, Soviet Turkestan.			ognevi Bobrinski, 1918	split from E. bottae	Juste, J., Benda, P., Garcia-Mudarra, J. L., & Ibanez, C. (2013). Phylogeny and systematics of Old World serotine bats (genus Eptesicus, Vespertilionidae, Chiroptera): an integrative approach. Zoologica Scripta, 42(5), 441-457.				Georgia|Armenia|Azerbaijan|Iran|Kazakhstan|Uzbekistan|Turkmenistan|Kyrgyzstan|Tajikistan|Afghanistan|Pakistan|India	Asia	Palearctic	LC	0	0	0	Eptesicus_ognevi	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Cnephaeus_ognevi	1005527	23	Ognev's Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Cnephaeus	NA	ognevi	Bobrinski	1	Eptesicus ognevi	Bobrinski, N.A. 1918. Заметки о летучих мышах, добытых в Бухарском ханстве. Материалы к познанию природы Российской Империи 15:1-22.	https://zmmu.msu.ru/dbs/pdf/bobrinskoi1915.pdf	ZMMU S-96374	holotype	https://zmmu.msu.ru/dbs/list_record.php?id=S-96374	Bokhara district, Soviet Turkestan.			split from E. bottae; moved from Eptesicus to Cnephaeus	Juste, J., Benda, P., Garcia-Mudarra, J. L., & Ibanez, C. (2013). Phylogeny and systematics of Old World serotine bats (genus Eptesicus, Vespertilionidae, Chiroptera): an integrative approach. Zoologica Scripta, 42(5), 441-457.|Cláudio, V. C., Novaes, R. L., Gardner, A. L., Nogueira, M. R., Wilson, D. E., Maldonado, J. E., ... & Moratelli, R. (2023). Taxonomic re-evaluation of New World Eptesicus and Histiotus (Chiroptera: Vespertilionidae), with the description of a new genus. Zoologia (Curitiba), 40, e22029.				Georgia|Armenia|Azerbaijan|Iran|Kazakhstan|Uzbekistan|Turkmenistan|Kyrgyzstan|Tajikistan|Afghanistan|Pakistan|India	Asia	Palearctic	LC (as Eptesicus ognevi)	0	0	0	Eptesicus_ognevi	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Cnephaeus		ognevi	Bobrinski	1918	1	Fauna and Flora of Russia	15: 2, 12	Ognev's Serotine	None.	Tajikistan [Emirate of Bukhara, Hisar prefecture, Sokhta-Chinar, village of Gisar (Tajik: Hisar) Bekstvo]	Jammu and Kasmir (India).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136598/21996784/' target='_blank'>Least Concern as Eptesicus ognevi</a>	Distinct from bottae; see Juste et al. (2012). May include additional cryptic forms - see Juste et al. (2012).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Eptesicus ognevi; Eptesicus ognevi; Eptesicus ognevi; Eptesicus ognevi; Eptesicus ognevi; ognevi; Sérotine d'Ognev; Ognev-Breitfligelfledermaus; Eptesicus de Ognev; Ognev's Serotine; Ognev's Serotine; E. ognevi