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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1672	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Eptesicus douglasi	Eptesicus douglasi	Pipistrellus douglasorum	Eptesicus douglasorum	Pipistrellus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum	Vespadelus douglasorum		[MSW2] Subgenus Vespadelus. Originally described as douglasi but emended by Kitchener et al. (1987).; [MSW3] caurinus species group. Originally described as douglasi but emended by Kitchener et al. (1987).; [HMW] Eptesicus douglasi [sic] Kitchener, 1976 , “Tunnel Creek, Napier Range, Western Australia ( 17°37'S , 125°09'E ),” Australia . Vespadelus seems to be sister to a clade including Chalinolobus and Nyctophilus . The specific name was originally given as “douglas?” but was emended to “ douglasorum ” by D. J. Kitchener and colleagues in 1987 as the taxon name was dedicated to both Mr. and Mrs. Douglas. As currently constituted, V. douglasorum may comprise more than one species. Monotypic.; [batnames2022]  caurinus species group. Originally described as douglasi but emended by Kitchener et al. (1987).; [IUCN] There is the possibility of a cryptic species resembling V. douglasorum in the Kimberley (Armstrong and Kitchener 2008; Armstrong 2011).; [batnames2023]  caurinus species group. Originally described as douglasi but emended by Kitchener et al. (1987).; [MDD2023] the original spelling of the specific epithet was 'douglasi', but was emended to 'douglasorum' later; normally, this would be considered an unjustified emendation, but the spelling 'douglasorum' is treated as in prevailing use given that almost no publications have used douglasi since the spelling emendation; [MDD2025_2.0] the original spelling of the specific epithet was 'douglasi', but was emended to 'douglasorum' later; normally, this would be considered an unjustified emendation, but the spelling 'douglasorum' is treated as in prevailing use given that almost no publications have used douglasi since the spelling emendation; [batnames2025_1.7] caurinusspecies group. Originally described as douglasi but emended by Kitchener et al. (1987).; [MDD2025_2.2] the original spelling of the specific epithet was 'douglasi', but was emended to 'douglasorum' later; normally, this would be considered an unjustified emendation, but the spelling 'douglasorum' is treated as in prevailing use given that almost no publications have used douglasi since the spelling emendation														douglasorum	There is the possibility of a cryptic species resembling V. douglasorum in the Kimberley (Armstrong and Kitchener 2008; Armstrong 2011).			douglasorum	douglasorum, douglasorum			douglasorum (D. J. Kitchener, 1976) [as emended]|douglasorum (D. J. Kitchener, B. Jones, & Caputi, 1987) [justified emendation]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		W Australia; ref. 4.87	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Eptesicus douglasi	Australia, Western Australia, Kimberley, Napier Ranges, Tunnel Creek.	Kitchener	1976	Rec. W. Aust. Mus., 4:295, 296.	Distribution: Confined to extreme northern parts of Western Australia and the Northern Territory.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Yellow-lipped bat	NW Australia	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Kitchener	1976	Rec. W. Aust. Mus., 4:295, 296.	Subgenus Vespadelus. Originally described as douglasi but emended by Kitchener et al. (1987).	Kimberley (N Western Australia).	Australia, Western Australia, Kimberley, Napier Ranges, Tunnel Creek.		KITCHENER	1976	Size fairly small (forearm length, 34-38 mm), but larger than pumilus where sympatric with it. Forehead relatively concave. Rostrum relatively broad. Head, foot, and forearm yellowish.	Distribution: Confined to extreme northern parts of Western Australia and the Northern Territory.	No subspecies.		116	species	P. douglasorum	KITCHENER	1976	Vespadelus	subgenus	Pipistrellus douglasorum				Size fairly small (forearm length, 34-38 mm), but larger than pumilus where sympatric with it. Forehead relatively concave. Rostrum relatively broad. Head, foot, and forearm yellowish.	No subspecies.		49. P. douglasorum KITCHENER 1976.	49	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Vespadelus douglasorum	Vespadelus		douglasorum	Kitchener	y	1976		Rec. West. Aust. Mus.	4		295, 296		Yellow-lipped Bat	Australia, Western Australia, Kimberley, Napier Range, Tunnel Creek.	Kimberley (N Western Australia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt) as Eptesicus douglasorum.		caurinus species group. Originally described as douglasi but emended by Kitchener et al. (1987).	4C3D87E8FFDA6A66FA8B9FA41A19B22A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	791	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFDA6A65FF7997FC1A19B09C.xml	Vespadelus douglasorum	Vespertilionidae	Vespadelus	douglasorum		1976	Vespertilion des Douglas @fr | Gelblippen-Waldfledermaus @de | Vespadela de Douglas @es | Kimberley Cave Bat @en | Yellow-lipped Bat @en	Eptesicus douglasi [sic] Kitchener, 1976 , “Tunnel Creek, Napier Range, Western Australia ( 17°37'S , 125°09'E ),” Australia . Vespadelus seems to be sister to a clade including Chalinolobus and Nyctophilus . The specific name was originally given as “douglas?” but was emended to “ douglasorum ” by D. J. Kitchener and colleagues in 1987 as the taxon name was dedicated to both Mr. and Mrs. Douglas. As currently constituted, V. douglasorum may comprise more than one species. Monotypic.	W Kimberley region of N Western Australia along with a few offshore Is (including Adolphus, Augustus, Bigge, Boongaree, Middle Osborn, Storr, and Wulalam).	Head-body 35-3-43-5 mm, tail 35-2—42-1 mm, ear 11-3-12-5 mm, forearm 34-3-37-8 mm; weight 4-5-6-1 g. Dorsal pelage is pale gray to olive buff, whereas ventral pelage is somewhat paler; there is generally a deep yellow to orange tinge to fur around mouth, head, shoulders, feet, and forearms; amount ofyellow or orange on body varies largely throughout the year. Face is mostly naked and similarly pale cinnamon to orange buff. Face has larger glandular swellings than in Northern Cave Bat (V. caurinus ). Ears are small and rounded triangular with a smoothly convex anterior edge; tragusis narrow, anteriorly straight or slightly concave, posteriorly convex, and with a rounded tip and slight posterobasal lobe. Wings and uropatagium are brown; uropatagium extends to tip of tail and is sparsely haired on proximal two-thirds. Glans penishas laterally compressed and rod-shaped head; dorsal surface has deep longitudinal groove, and ventral surface is covered in a number of small spines; urethra is covered by narrow strip of skin projecting upward from ventral tip. Baculum is moderately long (mean 3-2 mm) with rod-shaped shaft,slightly bifurcated base, and bow shape in lateral view. Skull is moderately long; lambdoidalcrest is weakly to moderately developed; anterior narial notch is generally V-shaped, occasionally U-shaped; rostrum is short.	Found in tropical woodlands, often captured along waterways lined by Melaleuca ( Myrtaceae ) and Pandanus ( Pandanaceae ), and in adjacent open woodland. All known recordsof the species are in regions with high rainfall (over 800 mmyear).	No information.	Pregnant females have been recorded in June, August, October, and November, and births in November and December. Litter size seems to be one.	Yellow-lipped Cave Bats are often found roosting in caves in limestone regions. They enter a torpid state during the day while roosting. Call shape is FM/QCF with a peak frequency recorded at 52-8 kHz. Average peak frequency was recorded at 51-2 kHz and the call duration at 5-13 milliseconds.	Yellow-lipped Cave Bats form mixed-sex colonies of up to 200 individuals. A colony with twelve individuals was found roosting in an abandoned building. Torpid individuals have been observed roosting in small clusters of 4-6 individuals in both cool and dry conditions at 20°C and 33% humidity in July; they were not close to the opening but in the same cave, there were more alert individuals deeper in the cave at 27°C and 48% humidity. This cave was shared with Orange Diamond-faced Bats (Rhinonicteris aurantia), which were even further into the cave. The species occasionally shares caves with the Northern Cave Bat, the Dusky Leat-nosed Bat ( Hipposideros ater ), and the Common Sheath-tailed Bat (Taphozous georgianus).	Classified as Least Concern on The IUCN Red List. The Yellowlipped Cave Bat is found in a remote region of Australia , which limits persecution due to human interaction. Burning for fire management may represent the greatest threat, as it can reduce habitat quality at specific localities. Overall, the speciesis rare and seldom recorded.	Armstrong (2011) | Armstrong & Kitchener (2008) | Armstrong, Burbidge & Woinarski (2017) | Churchill (2008) | Kitchener (1976b) | Kitchener et al. (1987) | McKenzie & Bullen (2012)	https://zenodo.org/record/6397902/files/figure.png	62. Yellow-lipped Cave Bat Vespadelus douglasorum French: Vespertilion des Douglas / German: Gelblippen-Waldfledermaus / Spanish: Vespadela de Douglas Other common names: Kimberley Cave Bat , Yellow-lipped Bat Taxonomy. Eptesicus douglasi [sic] Kitchener, 1976 , “Tunnel Creek, Napier Range, Western Australia ( 17°37'S , 125°09'E ),” Australia . Vespadelus seems to be sister to a clade including Chalinolobus and Nyctophilus . The specific name was originally given as “douglas?” but was emended to “ douglasorum ” by D. J. Kitchener and colleagues in 1987 as the taxon name was dedicated to both Mr. and Mrs. Douglas. As currently constituted, V. douglasorum may comprise more than one species. Monotypic. Distribution. W Kimberley region of N Western Australia along with a few offshore Is (including Adolphus, Augustus, Bigge, Boongaree, Middle Osborn, Storr, and Wulalam). Descriptive notes. Head-body 35-3-43-5 mm, tail 35-2—42-1 mm, ear 11-3-12-5 mm, forearm 34-3-37-8 mm; weight 4-5-6-1 g. Dorsal pelage is pale gray to olive buff, whereas ventral pelage is somewhat paler; there is generally a deep yellow to orange tinge to fur around mouth, head, shoulders, feet, and forearms; amount ofyellow or orange on body varies largely throughout the year. Face is mostly naked and similarly pale cinnamon to orange buff. Face has larger glandular swellings than in Northern Cave Bat (V. caurinus ). Ears are small and rounded triangular with a smoothly convex anterior edge; tragusis narrow, anteriorly straight or slightly concave, posteriorly convex, and with a rounded tip and slight posterobasal lobe. Wings and uropatagium are brown; uropatagium extends to tip of tail and is sparsely haired on proximal two-thirds. Glans penishas laterally compressed and rod-shaped head; dorsal surface has deep longitudinal groove, and ventral surface is covered in a number of small spines; urethra is covered by narrow strip of skin projecting upward from ventral tip. Baculum is moderately long (mean 3-2 mm) with rod-shaped shaft,slightly bifurcated base, and bow shape in lateral view. Skull is moderately long; lambdoidalcrest is weakly to moderately developed; anterior narial notch is generally V-shaped, occasionally U-shaped; rostrum is short. I? is bicuspid and I? is unicuspid; P* is much smaller than C' and within tooth row; crista linking base of metacone and hypocone on M' is moderate, and slight to large on M2 Dental formula for species of Vespadelusis12/3,C 1/1, P1/2,M 3/3 (x2) = 32, Habitat. Found in tropical woodlands, often captured along waterways lined by Melaleuca ( Myrtaceae ) and Pandanus ( Pandanaceae ), and in adjacent open woodland. All known recordsof the species are in regions with high rainfall (over 800 mmyear). Food and Feeding. No information. Breeding. Pregnant females have been recorded in June, August, October, and November, and births in November and December. Litter size seems to be one. Activity patterns. Yellow-lipped Cave Bats are often found roosting in caves in limestone regions. They enter a torpid state during the day while roosting. Call shape is FM/QCF with a peak frequency recorded at 52-8 kHz. Average peak frequency was recorded at 51-2 kHz and the call duration at 5-13 milliseconds. Movements, Home range and Social organization. Yellow-lipped Cave Bats form mixed-sex colonies of up to 200 individuals. A colony with twelve individuals was found roosting in an abandoned building. Torpid individuals have been observed roosting in small clusters of 4-6 individuals in both cool and dry conditions at 20°C and 33% humidity in July; they were not close to the opening but in the same cave, there were more alert individuals deeper in the cave at 27°C and 48% humidity. This cave was shared with Orange Diamond-faced Bats (Rhinonicteris aurantia), which were even further into the cave. The species occasionally shares caves with the Northern Cave Bat, the Dusky Leat-nosed Bat ( Hipposideros ater ), and the Common Sheath-tailed Bat (Taphozous georgianus). Status and Conservation. Classified as Least Concern on The IUCN Red List. The Yellowlipped Cave Bat is found in a remote region of Australia , which limits persecution due to human interaction. Burning for fire management may represent the greatest threat, as it can reduce habitat quality at specific localities. Overall, the speciesis rare and seldom recorded. Bibliography. Armstrong (2011), Armstrong & Kitchener (2008), Armstrong, Burbidge & Woinarski (2017), Churchill (2008), Kitchener (1976b), Kitchener et al. (1987), McKenzie & Bullen (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Vespadelus douglasorum	Vespadelus		douglasorum	Kitchener	1976	1	Rec. West. Aust. Mus.	4: 295, 296	Yellow-lipped Bat	None.	Australia, Western Australia, Kimberley, Napier Range, Tunnel Creek.	Kimberley (N Western Australia).	Not listed.	Least Concern	 caurinus species group. Originally described as douglasi but emended by Kitchener et al. (1987).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Vespadelus douglasorum	23	Yellow-lipped Cave Bat	Kimberley Cave Bat|Yellow-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Vespadelus	NA	douglasorum	Kitchener	1976	1	Eptesicus_douglasi	Kitchener, D. J. (1976). Eptesicus douglasi, a new Vespertilionid bat from Kimberley, Western Australia. Records of the Western Australian Museum, 4(3), 296.	http://museum.wa.gov.au/research/records-supplements/records/eptesicus-douglasi-new-vespertilionid-bat-kimberley-western-aus	WAM M50910		"Tunnel Creek, Napier Range, Western Australia (17°37'S, 125°09'E)," Australia.	-17.62	125.15	douglasorum (Kitchener, 1976)	NA	NA	Australia	Oceania	Australasia/Oceania	LC	0	0	0	Vespadelus_douglasorum	0	sciname match	Vespadelus_douglasorum	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	7923	Vespadelus douglasorum	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Vespadelus	douglasorum	(Kitchener, 1976)	There is the possibility of a cryptic species resembling V. douglasorum in the Kimberley (Armstrong and Kitchener 2008; Armstrong 2011).	200000000	Vespadelus douglasorum	Least Concern		2021	2016-07-31 00:00:00 UTC	3.1	English	<p>Listed as Least Concern given its apparently wide distribution, use of a broad range of habitats, presumed large population size, occurrence in protected areas, and the absence of significant key threats or evidence for a decline.</p>	<p>Vespadelus douglasorum roosts in caves, but they have also been collected from buildings. They occur mainly in woodlands, and can be encountered close to waterbodies in sandstone ranges, but do not utilise the extensive area of mangal habitat in the Kimberley (McKenzie and Bullen 2012). Little is known about their breeding, but females give birth to a single young and there is a record of lactating females in a maternity roost in March, so breeding probably occurs during the wet season (Armstrong and Kitchener 2008; Woinarski et al. 2014).</p>	<p>Vespadelus douglasorum lives in remote areas of Australia, so the risk of human disturbance, including at roost sites, is low. However, fire management and the frequency, timing and scale of burning is a significant issue in the Kimberley, and could have implications for the reduction of habitat quality for local occurrences (Woinarski et al . 2014). ; </p>	<p>This species is rare and seldom recorded, but there have been essentially no targeted surveys. Colonies of up to 80 individuals have been observed (Armstrong and Kitchener 2008), though this may not be typical of every roost.</p>	Unknown	<p>This species is endemic to the Kimberley region of Western Australia. All records are from the high rainfall areas of the north-western Kimberley and the Devonian Reef limestone ranges in the southern Kimberley (Woinarski et al . 2014). It has also been detected on seven Kimberley offshore islands: Adolphus, Augustus, Bigge, Boongaree, Middle Osborn, Storr, and Wulalam (McKenzie and Bullen 2012).</p>		Terrestrial	<p>This species has been recorded from several protected areas, but there is no specific management action that considers it. ;There is a requirement for further studies into basic biology, foraging and roost habitat preference, and to consider this species in broad-scale regional management strategies given its restriction to the Kimberley region of Australia.</span></p>	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Vespadelus		douglasorum	Kitchener	1976	1	Rec. West. Aust. Mus.	4: 295, 296	Yellow-lipped Bat	None.	Australia, Western Australia, Kimberley, Napier Range, Tunnel Creek.	Kimberley (N Western Australia).	Not listed.	Least Concern	 caurinus species group. Originally described as douglasi but emended by Kitchener et al. (1987).	Vespadelus douglasorum	1005786	23	Yellow-lipped Cave Bat	Kimberley Cave Bat|Yellow-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Vespadelus	NA	douglasorum	Kitchener	1976	1	Eptesicus_douglasi	Kitchener, D. J. (1976). Eptesicus douglasi, a new Vespertilionid bat from Kimberley, Western Australia. Records of the Western Australian Museum, 4(3), 296.	http://museum.wa.gov.au/research/records-supplements/records/eptesicus-douglasi-new-vespertilionid-bat-kimberley-western-aus	WAM M50910		"Tunnel Creek, Napier Range, Western Australia (17°37'S, 125°09'E)," Australia.	-17.6167	125.15	douglasorum (Kitchener, 1976)	the original spelling of the specific epithet was 'douglasi', but was emended to 'douglasorum' later; normally, this would be considered an unjustified emendation, but the spelling 'douglasorum' is treated as in prevailing use given that almost no publications have used douglasi since the spelling emendation	NA				Australia	Oceania	Australasia/Oceania	LC	0	0	0	Vespadelus_douglasorum	0	sciname match	Vespadelus_douglasorum	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Vespadelus_douglasorum	1005786	23	Yellow-lipped Cave Bat	Kimberley Cave Bat|Yellow-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Vespadelus	NA	douglasorum	D. J. Kitchener	1	Eptesicus douglasi	Kitchener, D.J. 1976-10-15. _Eptesicus douglasi_, a new vespertilionid bat from Kimberley, Western Australia. Records of the Western Australian Museum 4(3):295-301.	https://www.biodiversitylibrary.org/page/52806351	WAM M50190 (= WAM M3405c)	holotype	https://museum.wa.gov.au/catalogues-beta/digitised-types/mammalogy/vespadelus-douglasorum	"Tunnel Creek, Napier Range, Western Australia (17°37'S, 125°09'E)," Australia.	-17.6167	125.15	the original spelling of the specific epithet was 'douglasi', but was emended to 'douglasorum' later; normally, this would be considered an unjustified emendation, but the spelling 'douglasorum' is treated as in prevailing use given that almost no publications have used douglasi since the spelling emendation	NA				Australia	Oceania (Continent)	Australasia	LC	0	0	0	Vespadelus_douglasorum	0	sciname match	Vespadelus_douglasorum	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Vespadelus		douglasorum	Kitchener	1976	1	Rec. West. Aust. Mus.	4: 295, 296	Yellow-lipped Bat	None.	Australia, Western Australia, Kimberley, Napier Range, Tunnel Creek.	Kimberley (N Western Australia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/7923/209538760/' target='_blank'>Least Concern</a>	caurinusspecies group. Originally described as douglasi but emended by Kitchener et al. (1987).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Vespadelus douglasorum; Vespadelus douglasorum; Vespadelus douglasorum; Vespadelus douglasorum; Vespadelus douglasorum; Vespadelus douglasorum; douglasorum; Vespertilion des Douglas; Gelblippen-Waldfledermaus; Vespadela de Douglas; Kimberley Cave Bat; Yellow-lipped Bat; Yellow-lipped Cave Bat; Kimberley Cave Bat; Yellow-lipped Bat; Yellow-lipped Bat; Yellow-lipped Bat; V. douglasorum