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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1658	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Vampyressa pusilla [synonym of]	N/A	Vampyressa pusilla [synonym of]	Vampyressa pusilla thyone	Vampyressa thyone	Vampyressa thyone	Vampyressa thyone	Vampyressa thyone	Vampyressa thyone	Vampyressa thyone	Vampyressa thyone	Vampyressa thyone	Vampyressa thyone	Vampyressa thyone		[MSW3] Subgenus Vampyressa. Previously included in pusilla, but clearly distinct; see Lim et al. (2003). Much of the account of V. pusilla provided in Lewis and Wilson (1987) actually applies to thyone.; [HMW] Vampyressa thyone Thomas, 1909 , “Chimbo, near Guayaquil [Bolivar], Ecuador . Alt. 1000° [= 305 m ].” Vampyressa thyone was associated with V. minuta by G. S. Miller, Jr. in 1912 (from Panama ) and V. venilla by O. Thomas in 1924 (from northern Peru ); however, it more commonly has been treated as a synonym or subspecies of V. pusilla . B. K. Lim and collaborators in 2003 confirmed that it is a distinct species. Monotypic.; [batnames2022] Previously included in pusilla , but clearly distinct; see Lim et al. (2003). Much of the account of V. pusilla provided in Lewis and Wilson (1987) actually applies to thyone .; [IUCN] This is a species complex of at least three species (Tavares pers. comm.), which was previously included in V. pusilla , and separated by Lim et al. (2003).; [batnames2023] Previously included in pusilla , but clearly distinct; see Lim et al. (2003). Much of the account of V. pusilla provided in Lewis and Wilson (1987) actually applies to thyone .; [batnames2025_1.7] Previously included in pusilla, but clearly distinct; see Lim et al. (2003). Much of the account of V. pusilla provided in Lewis and Wilson (1987) actually applies to thyone. Specimens from SW Mexico are attributed to the new species villai by Garbino et al. (2024).									minuta, venilla			thyone 	thyone - minuta, venilla	thyone, minuta, venilla	This is a species complex of at least three species (Tavares pers. comm.), which was previously included in V. pusilla , and separated by Lim et al. (2003).	thyone 	thyone - minuta, venilla	thyone, minuta, venilla	thyone, minuta, venilla	thyone 	thyone - minuta, venilla	thyone O. Thomas, 1909|minuta G. S. Miller, 1912|venilla O. Thomas, 1924						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Stenodermatinae	Stenodermatini	Vampyressa thyone	Vampyressa	Vampyressa	thyone	Thomas		1909		Ann. Mag. Nat. Hist., ser. 8	4		231		Northern Little Yellow-eared Bat	Ecuador, Bolívar, Chimbo; 1000 ft (305 m).	Oaxaca and Veracruz (Mexico) to Bolivia, Peru, Venezuela, Guyana, and French Guiana.	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).	minuta Miller, 1912; venilla Thomas, 1924.	Subgenus Vampyressa. Previously included in pusilla, but clearly distinct; see Lim et al. (2003). Much of the account of V. pusilla provided in Lewis and Wilson (1987) actually applies to thyone.	03A687BCFFF6FFF61341F600F8F7FA1D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	559	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFFAFFFA13BDF4E2F80FFA2B.xml	Vampyressa thyone	Phyllostomidae	Vampyressa	thyone	Thomas	1909	Vampyresse thyoné @fr | Nordliche Kleine Gelbohrfledermaus @de | Vampiresa de Thyone @es	Vampyressa thyone Thomas, 1909 , “Chimbo, near Guayaquil [Bolivar], Ecuador . Alt. 1000° [= 305 m ].” Vampyressa thyone was associated with V. minuta by G. S. Miller, Jr. in 1912 (from Panama ) and V. venilla by O. Thomas in 1924 (from northern Peru ); however, it more commonly has been treated as a synonym or subspecies of V. pusilla . B. K. Lim and collaborators in 2003 confirmed that it is a distinct species. Monotypic.	From S Mexico ( Veracruz , Oaxaca , Tabasco , and Chiapas ) through Central America (except dry Pacific slope and Yucatan Peninsula) to W Colombia , Venezuela , Guyana , French Guiana , and W Ecuador , and through the Amazon Basin of Colombia , Venezuela , N & W Brazil , Ecuador , Peru , and Bolivia .	Head-body 43-52 mm (tailless), ear 11-15 mm, hindfoot 7-10 mm, forearm 29-34 mm; weight 6-10 g. The Northern Little Yellow-eared Bat is very small, among the smallest in the family. Dorsal pelage is pale brown; hairs are tricolored, with brown bases, central portion creamy-white, and paler tips. There is no median back stripe. Facial stripes are indistinct, most often looking like small paler spots above and below eyes. Tragus, entire base of ear, and upper edges of ears are yellow; noseleafis mostly brown, with yellow edges. Uropatagium is short, with light fringe of yellowish hairs along middle posterior border. Legs and feet are sparsely haired. Forearms, fingers, and thumbs are brown. Skull has short, broad rostrum, and postorbital processes are weakly developed. I? are elongated and roughly evenly bilobate, converging distally but not in contact at tips, and more than twice as high as I*. Upper molars are broader than long; lower incisors are small, completely filling space between canines, and their cutting edges are faintly trilobate. P® is one-half or less the size of P*, and the latteris triangular, with well-developed cingulum. M, is quadrate-shaped, with large protoconid and small hypoconid. M, has large metaconid and blade-like entoconid of about the same length as metaconid. M, is absent. Dental formula is I 2 /2, C 1/1, P 2/2, M 2/2 ( x2 ) = 28. There are three chromosomal complements: 2n = 18 and FN = 20 in Honduras and Nicaragua ; 2n = 23/24 and FN = 22 in Colombia and 2n = 22/23 and FN = 22 in northern Peru , with a sex-chromosome system XXXXXX.Y, with the Y-chromosome translocated to the autosomes.	Mostly well-preserved humid forests and also disturbed secondary forests, rarely croplands, modified habitats, and dry habitats at elevations up to 1500 m (most commonly below 800 m ).	Fruits in diets of Northern Little Yellow-eared Bats include a high proportion of Ficus spp. ( Moraceae ) and other small fruits; it qualifies as a fig specialist in Panama . Samples from Costa Rica showed a cultivated fruit-bearing tree as a source of food. Most stomach contents had fruit pulp.	Northern Little Yellow-eared Bats reproduce twice a year, with young born late in dry season and middle of wet season. Pregnant and lactating females have been recorded year-round, suggesting bimodal polyestry.	Greatest activity of Northern Little Yellow-eared Bats occurs during the first two hours after sunset, but they can be active throughout the night. They are usually caught in nets set across streams, preferring to fly in forest subcanopies rather than understories. Known roosting sites include bushes and tree branches. They make tents from Philodendron and other aroids. Availability of suitable leaves to build tents could determine distribution and local abundance.	Roosting groups of Little Yelloweared Bats are composed of one to five individuals.	Classified as Least Concern on The IUCN Red List. Although the Northern Little Yellow-eared Bat is not locally common, its distribution is very wide, diminishing potential threats to its overall population.	Arroyo-Cabrales (2008b) | Baker et al. (1973) | Bonaccorso (1979) | Davis & Dixon (1976) | Gardner (1977a, 1977b) | Gomes et al. (2016) | Miller (1912) | Lewis & Wilson (1987) | Lim, B.K., Pedro & Passos (2003) | Reid (2009) | Thomas (1924a) | Timm (1984) | Wilson (1979)	https://zenodo.org/record/6458922/files/figure.png	160. Northern Little Yellow-eared Bat Vampyressa thyone French: Vampyresse thyoné / German: Nordliche Kleine Gelbohrfledermaus / Spanish: Vampiresa de Thyone Taxonomy. Vampyressa thyone Thomas, 1909 , “Chimbo, near Guayaquil [Bolivar], Ecuador . Alt. 1000° [= 305 m ].” Vampyressa thyone was associated with V. minuta by G. S. Miller, Jr. in 1912 (from Panama ) and V. venilla by O. Thomas in 1924 (from northern Peru ); however, it more commonly has been treated as a synonym or subspecies of V. pusilla . B. K. Lim and collaborators in 2003 confirmed that it is a distinct species. Monotypic. Distribution. From S Mexico ( Veracruz , Oaxaca , Tabasco , and Chiapas ) through Central America (except dry Pacific slope and Yucatan Peninsula) to W Colombia , Venezuela , Guyana , French Guiana , and W Ecuador , and through the Amazon Basin of Colombia , Venezuela , N & W Brazil , Ecuador , Peru , and Bolivia . Descriptive notes. Head-body 43-52 mm (tailless), ear 11-15 mm, hindfoot 7-10 mm, forearm 29-34 mm; weight 6-10 g. The Northern Little Yellow-eared Bat is very small, among the smallest in the family. Dorsal pelage is pale brown; hairs are tricolored, with brown bases, central portion creamy-white, and paler tips. There is no median back stripe. Facial stripes are indistinct, most often looking like small paler spots above and below eyes. Tragus, entire base of ear, and upper edges of ears are yellow; noseleafis mostly brown, with yellow edges. Uropatagium is short, with light fringe of yellowish hairs along middle posterior border. Legs and feet are sparsely haired. Forearms, fingers, and thumbs are brown. Skull has short, broad rostrum, and postorbital processes are weakly developed. I? are elongated and roughly evenly bilobate, converging distally but not in contact at tips, and more than twice as high as I*. Upper molars are broader than long; lower incisors are small, completely filling space between canines, and their cutting edges are faintly trilobate. P® is one-half or less the size of P*, and the latteris triangular, with well-developed cingulum. M, is quadrate-shaped, with large protoconid and small hypoconid. M, has large metaconid and blade-like entoconid of about the same length as metaconid. M, is absent. Dental formula is I 2 /2, C 1/1, P 2/2, M 2/2 ( x2 ) = 28. There are three chromosomal complements: 2n = 18 and FN = 20 in Honduras and Nicaragua ; 2n = 23/24 and FN = 22 in Colombia and 2n = 22/23 and FN = 22 in northern Peru , with a sex-chromosome system XXXXXX.Y, with the Y-chromosome translocated to the autosomes. Habitat. Mostly well-preserved humid forests and also disturbed secondary forests, rarely croplands, modified habitats, and dry habitats at elevations up to 1500 m (most commonly below 800 m ). Food and Feeding. Fruits in diets of Northern Little Yellow-eared Bats include a high proportion of Ficus spp. ( Moraceae ) and other small fruits; it qualifies as a fig specialist in Panama . Samples from Costa Rica showed a cultivated fruit-bearing tree as a source of food. Most stomach contents had fruit pulp. Breeding. Northern Little Yellow-eared Bats reproduce twice a year, with young born late in dry season and middle of wet season. Pregnant and lactating females have been recorded year-round, suggesting bimodal polyestry. Activity patterns. Greatest activity of Northern Little Yellow-eared Bats occurs during the first two hours after sunset, but they can be active throughout the night. They are usually caught in nets set across streams, preferring to fly in forest subcanopies rather than understories. Known roosting sites include bushes and tree branches. They make tents from Philodendron and other aroids. Availability of suitable leaves to build tents could determine distribution and local abundance. Movements, Home range and Social organization. Roosting groups of Little Yelloweared Bats are composed of one to five individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List. Although the Northern Little Yellow-eared Bat is not locally common, its distribution is very wide, diminishing potential threats to its overall population. Bibliography. Arroyo-Cabrales (2008b), Baker et al. (1973), Bonaccorso (1979), Davis & Dixon (1976), Gardner (1977a, 1977b), Gomes et al. (2016), Miller (1912), Lewis & Wilson (1987), Lim, B.K., Pedro & Passos (2003), Reid (2009), Thomas (1924a), Timm (1984), Wilson (1979).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Vampyressa thyone	Vampyressa	Vampyressa	thyone	Thomas	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 231	Northern Little Yellow-eared Bat	 minuta Miller, 1912; venilla Thomas, 1924.	Ecuador, BolÃ­var, Chimbo; 1000 ft (305 m).	Oaxaca and Veracruz (Mexico) to Bolivia, Peru, Venezuela, Guyana, and French Guiana.	Not listed.	Least Concern	Previously included in pusilla , but clearly distinct; see Lim et al. (2003). Much of the account of V. pusilla provided in Lewis and Wilson (1987) actually applies to thyone .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Vampyressa thyone	23	Northern Little Yellow-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STENODERMATINI	Vampyressa	NA	thyone	O. Thomas	1909	0	Vampyressa_thyone	Thomas, O. (1909). Notes on some South-American mammals, with descriptions of new species. Annals and Magazine of Natural History, ser. 8, 4, 231.	https://www.biodiversitylibrary.org/item/71907#page/261/mode/1up	BM 1897.11.7.77		"Chimbo, near Guayaquil [BolÃ­var], Ecuador. Alt. 1000' [= 305 m]."			thyone O. Thomas, 1909|minuta G. S. Miller, 1912|venilla O. Thomas, 1924	NA	NA	Mexico|Belize|Guatemala|Honduras|Nicaragua|Costa Rica|Colombia|Venezuela|Guyana|French Guiana|Ecuador|Peru|Bolivia|Brazil	North America|South America	Nearctic|Neotropic	LC	0	0	0	Vampyressa_thyone	0	sciname match	Vampyressa_thyone	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136671	Vampyressa thyone	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Vampyressa	thyone	Thomas, 1909	This is a species complex of at least three species (Tavares pers. comm.), which was previously included in V. pusilla , and separated by Lim et al. (2003).	20000000	Vampyressa thyone	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern as it is widespread, can be locally common and is unlikely to be declining rapidly enough to qualify for a threatened category. It is also adaptable to habitat change. This species should be reassessed following taxonomic revision.	This bat is found in semi-deciduous and evergreen forest. ;It is found in a variety of habitat types. ;Its roosts in tents ;(Timm 1984, Kunz ;et al . 1994) ;made from philodendron and other heart-shaped leaves. The basal nerves and midrib are chewed near the leaf stem, causing the sides and tip of the leaf to fold down. Tents are occupied by one to five individuals. This species has the greatest activity during the first two hours after sunset. ;It is a frugivore and figs and other small fruits are eaten. Reproduction may occur twice annually, with young born late in the dry season and in the middle of the wet season (Kunz et al.  1994, Reid 1997). ;As with other members of the genus, the ecology is not well known.	There are no major threats known as this species is adaptable to human land-uses. Habitat loss is a threat in some parts of the range.	This species is uncommon but widespread (Reid 1997) in Mesoamerica, but is common throughout its range in South America. It can be locally common (Tavares pers. comm.).	Unknown	This species occurs throughout Oaxaca and Veracruz (Mexico) to Bolivia, Peru, Venezuela, Guyana and French Guiana (Simmons 2005). It also occurs in northern and western Brazil (Tavares pers. comm.). It occurs up to 1,000 m asl (Molinari pers. comm.) in the Andes. There is a record from Chiapas, Mexico (Reid pers. comm.). ;The last record is from Amazonian Brazil (Tavares pers. comm.)	This species is not used.	Terrestrial	The recommended conservation action is to reduce habitat loss. Further studies of the basic ecology, threats and taxonomy are required.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Vampyressa	Vampyressa	thyone	Thomas	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 231	Northern Little Yellow-eared Bat	 minuta Miller, 1912; venilla Thomas, 1924.	Ecuador, BolÃ­var, Chimbo; 1000 ft (305 m).	Oaxaca and Veracruz (Mexico) to Bolivia, Peru, Venezuela, Guyana, and French Guiana.	Not listed.	Least Concern	Previously included in pusilla , but clearly distinct; see Lim et al. (2003). Much of the account of V. pusilla provided in Lewis and Wilson (1987) actually applies to thyone .	Vampyressa thyone	1005064	23	Northern Little Yellow-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STENODERMATINI	Vampyressa	NA	thyone	O. Thomas	1909	0	Vampyressa_thyone	Thomas, O. (1909). Notes on some South-American mammals, with descriptions of new species. Annals and Magazine of Natural History, ser. 8, 4, 231.	https://www.biodiversitylibrary.org/item/71907#page/261/mode/1up	BM 1897.11.7.77		"Chimbo, near Guayaquil [BolÃ­var], Ecuador. Alt. 1000' [= 305 m]."			thyone O. Thomas, 1909|minuta G. S. Miller, 1912|venilla O. Thomas, 1924	NA	NA				Mexico|Belize|Guatemala|Honduras|Nicaragua|Costa Rica|Colombia|Venezuela|Guyana|French Guiana|Ecuador|Peru|Bolivia|Brazil	North America|South America	Nearctic|Neotropic	LC	0	0	0	Vampyressa_thyone	0	sciname match	Vampyressa_thyone	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Vampyressa_thyone	1005064	23	Northern Little Yellow-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Stenodermatini	Vampyressa	NA	thyone	O. Thomas	0	Vampyressa thyone	Thomas, O. 1909-09-01. Notes on some South-American mammals, with descriptions of new species. Annals and Magazine of Natural History (8)4(21):230-242.	https://www.biodiversitylibrary.org/page/22097610	BMNH:Mamm:1897.11.7.77	holotype	https://data.nhm.ac.uk/object/a8100d37-6c40-405c-acfe-caeb9d7f9611	"Chimbo, near Guayaquil [BolÃ­var], Ecuador. Alt. 1000' [= 305 m]."			NA	NA				Mexico|Belize|Guatemala|Honduras|Nicaragua|Costa Rica|Colombia|Venezuela|Guyana|French Guiana|Ecuador|Peru|Bolivia|Brazil	North America|South America	Nearctic|Neotropic	LC	0	0	0	Vampyressa_thyone	0	sciname match	Vampyressa_thyone	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Vampyressa	Vampyressa	thyone	Thomas	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 231	Northern Little Yellow-eared Bat	minuta Miller, 1912; venilla Thomas, 1924.	Ecuador, BolÃ­var, Chimbo; 1000 ft (305 m)	Veracruz (Mexico) to Bolivia, Peru, Venezuela, Guyana, and French Guiana	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136671/21989318/' target='_blank'>Least Concern</a>	Previously included in pusilla, but clearly distinct; see Lim et al. (2003). Much of the account of V. pusilla provided in Lewis and Wilson (1987) actually applies to thyone. Specimens from SW Mexico are attributed to the new species villai by Garbino et al. (2024).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Vampyressina	Vampyressa thyone; Vampyressa thyone; Vampyressa thyone; Vampyressa thyone; Vampyressa thyone; Vampyressa thyone; minuta; venilla; minuta; venilla; thyone; minuta; venilla; Vampyresse thyoné; Nordliche Kleine Gelbohrfledermaus; Vampiresa de Thyone; Northern Little Yellow-eared Bat; Northern Little Yellow-eared Bat; Northern Little Yellow-eared Bat; V. thyone
