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(1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common 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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1644	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Tylonycteris robustula [synonym of]	N/A	Tylonycteris robustula [synonym of]	Tylonycteris robustula malayana	Tylonycteris robustula malayana	Tylonycteris malayana	Tylonycteris malayana	Tylonycteris malayana	Tylonycteris robustula [synonym of]?	Tylonycteris malayana	Tylonycteris malayana	Tylonycteris malayana	Tylonycteris malayana	Tylonycteris malayana		[HMW] 7ylonycteris malayana Chasen, 1940 , “Jor, Batang Padang district, Perak , Malay States [= Malaysia ].” Previously considered a subspecies of 1. robustula , but elevated to species status as a result of recent phylogenetic studies. 1. malayana is sister species to 1. tonkinensis. The exact distributional limit between the two is currently uncertain. The recently described subspecies eremtaga is sister to all other populations of 7. malayana . Two subspecies recognized.; [batnames2022] Previously included in robustula , but distinct, see Tu et al. (2017).; [MDD2022] split from T. robustula; [batnames2023] Previously included in robustula , but distinct, see Tu et al. (2017).; [MDD2023] split from T. robustula; [MDD2025_2.0] split from T. robustula; [batnames2025_1.7] Previously included in robustula, but distinct, see Tu et al. (2017).; [MDD2025_2.2] split from T. robustula										malayana, eremtaga		malayana, eremtaga		malayana, eremtaga		malayana, eremtaga		malayana, eremtaga	malayana, eremtaga	eremtaga, malayana 		malayana Chasen, 1940|eremtaga C. Srinivasulu, A. Srinivasulu, B. Srinivasulu, & G. Jones, 2018						N/A																																								_T. m. eremtaga_ Srinivasulu, Srinivasulu, Srinivasulu & Jones, 2018; _T. m. malayana_ Chasen, 1940																											4C3D87E8FFE66A59FF8B929D1D1DB7FD	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	787	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFE66A5AFA939CFC1B22B34D.xml	Tylonycteris malayana	Vespertilionidae	Tylonycteris	malayana	Chasen	1940	Vespertilion de Malaisie @fr | Malaya-Bambusfledermaus @de | Tilonicteriode Malasia @es | Malayan Bamboo Bat @en	7ylonycteris malayana Chasen, 1940 , “Jor, Batang Padang district, Perak , Malay States [= Malaysia ].” Previously considered a subspecies of 1. robustula , but elevated to species status as a result of recent phylogenetic studies. 1. malayana is sister species to 1. tonkinensis. The exact distributional limit between the two is currently uncertain. The recently described subspecies eremtaga is sister to all other populations of 7. malayana . Two subspecies recognized.	T.m.malayanaChasen,1940—NEIndia(Mizoram),andmainlandSEAsia;possiblyalsoChina(Yunnan)butthesepopulationsneedtobeconfirmed. T. m. eremtaga C. Srinivasulu et al., 2018 — North Andaman I in the Andaman Is.	Head—body 36—-37-3 mm, tail 26-3-27-8 mm, ear 10-2 mm, hindfoot 5-8-6-2 mm, forearm 23-3-27-5 mm; weight 6:5-8-5 g. The Malayan Greater Bamboo Batis the largest species of Tylonycteris . Head is dorso-ventrally flattened and broadened. Dorsal pelage is glossy dark brown to grayish brown (subspecies eremtaga) with light-tipped hairs (giving the glossy appearance); ventral pelage is paler grayish brown to sandy brown. Membranes are dark brown. Ears are subtriangular, relatively long with a broadly rounded tip; tragus is short and blunt. Base of thumbs and soles of hindfeet have well-developed rounded fleshy pads for gripping smooth surfaces. Wing membrane attaches at base of metatarsus; uropatagium extends to tip oftail, and calcar is over halfway to tail from ankle. Baculum of eremtaga is located at tip of short penis (which has a globular glans); base is broad, flared, knob like, and slightly concave or notched; shaft is long, straight, and has slight ventral concavity beneath broadly rounded tip; shaft when viewed laterally is long and straight with flared base (unlike in nominate, where shaft curves dorsally along its glans). Skull is small, flattened and broad; rostrum is short and broad; there is no sagittal crest; lambdoid crests are weakly developed. P? is large, narrow, and bicuspid; P* is unicuspid and subequal in height to second cusp of P?; C! has distinct second cusp; P* has crown area about half that of M!; M! slightly exceeds M? in breadth and has distinct protocone; three lowerincisors are tricuspid; lower premolars are subequalin size. Chromosomal complement has 2n = 32 and FN = 56 (Peninsular Malaysia ).	Both intact and disturbed forest habitats with abundant bamboo. Recorded at elevations up to 1000 m in India .	The Malayan Greater Bamboo Batis insectivorous,taking a variety of insect prey. It has been recorded feeding on termites; more specific data are not available.	Breeding takes place once a year, and females usually give birth to twins in April. Weaning occurs by late May in Malaysia , when young become volant and independent. Males and females become sexually mature and breed in their first year. Males become fertile from mid-October to mid-March; females enter estrus from mid-November to when ovulation occurs in mid-January. Gestation appears to last 84-91 days.	Malayan Greater Bamboo Bats are nocturnal, leaving their roosts around dusk to forage. They roost inside hollow bamboo internodes and some roosts have also been found in rock crevices. Call shapeis a steep FM/QCF sweep with average maximum frequency of 105-7 kHz, minimum frequency 43-2 kHz, peak frequency 52-3 kHz, and call duration 3-6 milliseconds, in Singapore . In Thailand , average peak frequency was recorded at 51 kHz, maximum frequency 137-2 kHz, minimum frequency 39-7 kHz, duration 3-3 milliseconds, and pulse interval 12-8 milliseconds.	The Malayan Greater Bamboo Bat has been recorded roosting with the Indomalayan Lesser Bamboo Bat (7. fulvida), although only rarely. Males typically roost alone, whereas females are more gregarious. Colonies with up to 32 bats have been recorded. Both sexes switch roosts often.	Not assessed as a distinct species on The IUCN Red List, but makes up most of what was formerly recognized as T. robustula , which was listed as Least Concern.	Huang Chujing et al. (2014) | Hughes et al. (2011) | Medway (1972, 1973) | Medway & Marshall (1970) | Pottie et al. (2005) | Tu Vuong Tan, Csorba et al. (2017) | Srinivasulu et al. (2018) | Yong et al. (1971)	https://zenodo.org/record/6397886/files/figure.png	55. Malayan Greater Bamboo Bat Tylonycteris malayana French: Vespertilion de Malaisie / German: Malaya-Bambusfledermaus / Spanish: Tilonicterio de Malasia Other common names: Malayan Bamboo Bat Taxonomy. 7ylonycteris malayana Chasen, 1940 , “Jor, Batang Padang district, Perak , Malay States [= Malaysia ].” Previously considered a subspecies of 1. robustula , but elevated to species status as a result of recent phylogenetic studies. 1. malayana is sister species to 1. tonkinensis. The exact distributional limit between the two is currently uncertain. The recently described subspecies eremtaga is sister to all other populations of 7. malayana . Two subspecies recognized. Subspecies and Distribution. T.m.malayanaChasen,1940—NEIndia(Mizoram),andmainlandSEAsia;possiblyalsoChina(Yunnan)butthesepopulationsneedtobeconfirmed. T. m. eremtaga C. Srinivasulu et al., 2018 — North Andaman I in the Andaman Is. Descriptive notes. Head—body 36—-37-3 mm, tail 26-3-27-8 mm, ear 10-2 mm, hindfoot 5-8-6-2 mm, forearm 23-3-27-5 mm; weight 6:5-8-5 g. The Malayan Greater Bamboo Batis the largest species of Tylonycteris . Head is dorso-ventrally flattened and broadened. Dorsal pelage is glossy dark brown to grayish brown (subspecies eremtaga) with light-tipped hairs (giving the glossy appearance); ventral pelage is paler grayish brown to sandy brown. Membranes are dark brown. Ears are subtriangular, relatively long with a broadly rounded tip; tragus is short and blunt. Base of thumbs and soles of hindfeet have well-developed rounded fleshy pads for gripping smooth surfaces. Wing membrane attaches at base of metatarsus; uropatagium extends to tip oftail, and calcar is over halfway to tail from ankle. Baculum of eremtaga is located at tip of short penis (which has a globular glans); base is broad, flared, knob like, and slightly concave or notched; shaft is long, straight, and has slight ventral concavity beneath broadly rounded tip; shaft when viewed laterally is long and straight with flared base (unlike in nominate, where shaft curves dorsally along its glans). Skull is small, flattened and broad; rostrum is short and broad; there is no sagittal crest; lambdoid crests are weakly developed. P? is large, narrow, and bicuspid; P* is unicuspid and subequal in height to second cusp of P?; C! has distinct second cusp; P* has crown area about half that of M!; M! slightly exceeds M? in breadth and has distinct protocone; three lowerincisors are tricuspid; lower premolars are subequalin size. Chromosomal complement has 2n = 32 and FN = 56 (Peninsular Malaysia ). Habitat. Both intact and disturbed forest habitats with abundant bamboo. Recorded at elevations up to 1000 m in India . Food and Feeding. The Malayan Greater Bamboo Batis insectivorous,taking a variety of insect prey. It has been recorded feeding on termites; more specific data are not available. Breeding. Breeding takes place once a year, and females usually give birth to twins in April. Weaning occurs by late May in Malaysia , when young become volant and independent. Males and females become sexually mature and breed in their first year. Males become fertile from mid-October to mid-March; females enter estrus from mid-November to when ovulation occurs in mid-January. Gestation appears to last 84-91 days. Activity patterns. Malayan Greater Bamboo Bats are nocturnal, leaving their roosts around dusk to forage. They roost inside hollow bamboo internodes and some roosts have also been found in rock crevices. Call shapeis a steep FM/QCF sweep with average maximum frequency of 105-7 kHz, minimum frequency 43-2 kHz, peak frequency 52-3 kHz, and call duration 3-6 milliseconds, in Singapore . In Thailand , average peak frequency was recorded at 51 kHz, maximum frequency 137-2 kHz, minimum frequency 39-7 kHz, duration 3-3 milliseconds, and pulse interval 12-8 milliseconds. Movements, Home range and Social organization. The Malayan Greater Bamboo Bat has been recorded roosting with the Indomalayan Lesser Bamboo Bat (7. fulvida), although only rarely. Males typically roost alone, whereas females are more gregarious. Colonies with up to 32 bats have been recorded. Both sexes switch roosts often. Status and Conservation. Not assessed as a distinct species on The IUCN Red List, but makes up most of what was formerly recognized as T. robustula , which was listed as Least Concern. Bibliography. Huang Chujing et al. (2014), Hughes et al. (2011), Medway (1972, 1973), Medway & Marshall (1970), Pottie et al. (2005), Tu Vuong Tan, Csorba et al. (2017), Srinivasulu et al. (2018), Yong et al. (1971).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Tylonycteris malayana	Tylonycteris		malayana	Chasen	1940	0	Bull. Raffles Mus. Singapore	15:52	Malayan Bamboo Bat	</b> eremtaga </b> Srinivasulu, Srinivasulu, Srinivasulu & Jones, 2018.	Peninsular Malaysia	NE India, Bangladesh, Bhutan, Burma, Thailand, Cambodia, S Vietnam, peninsular Malaysia	Not listed	Least Concern under Tylonycteris robustula 	Previously included in robustula , but distinct, see Tu et al. (2017).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Tylonycteris malayana	23	Malayan Greater Bamboo Bat	Malayan Bamboo Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Tylonycteris	NA	malayana	Chasen	1940	0						"Jor, Batang Padang district, Perak, Malay States [= Malaysia]."			malayana Chasen, 1940|eremtaga C. Srinivasulu, A. Srinivasulu, B. Srinivasulu, & G. Jones, 2018	split from T. robustula	Tu, V. T., Csorba, G., Ruedi, M., Furey, N. M., Son, N. T., Thong, V. D., ... & Hassanin, A. (2017). Comparative phylogeography of bamboo bats of the genus Tylonycteris (Chiroptera, Vespertilionidae) in Southeast Asia. European Journal of Taxonomy, 274, 1-38.	India|Andaman Islands|China?|Myanmar|Vietnam|Thailand|Laos|Cambodia|Malaysia|Singapore	Asia	Indomalaya	NA	0	0	0	Tylonycteris_malayana	0	unmatched	NA	1																																			Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Tylonycteris		malayana	Chasen	1940	0	Bull. Raffles Mus. Singapore	15:52	Malayan Bamboo Bat	</b> eremtaga </b> Srinivasulu, Srinivasulu, Srinivasulu & Jones, 2018.	Peninsular Malaysia	NE India, Bangladesh, Bhutan, Burma, Thailand, Cambodia, S Vietnam, peninsular Malaysia	Not listed	Least Concern under Tylonycteris robustula 	Previously included in robustula , but distinct, see Tu et al. (2017).	Tylonycteris malayana	1005778	23	Malayan Greater Bamboo Bat	Malayan Bamboo Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Tylonycteris	NA	malayana	Chasen	1940	0						"Jor, Batang Padang district, Perak, Malay States [= Malaysia]."			malayana Chasen, 1940|eremtaga C. Srinivasulu, A. Srinivasulu, B. Srinivasulu, & G. Jones, 2018	split from T. robustula	Tu, V. T., Csorba, G., Ruedi, M., Furey, N. M., Son, N. T., Thong, V. D., ... & Hassanin, A. (2017). Comparative phylogeography of bamboo bats of the genus Tylonycteris (Chiroptera, Vespertilionidae) in Southeast Asia. European Journal of Taxonomy, 274, 1-38.				India|Andaman Islands|China?|Myanmar|Vietnam|Thailand|Laos|Cambodia|Malaysia|Singapore	Asia	Indomalaya	NA	0	0	0	Tylonycteris_malayana	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Tylonycteris_malayana	1005778	23	Malayan Greater Bamboo Bat	Malayan Bamboo Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Tylonycteris	NA	malayana	Chasen	0	Tylonycteris malayana	Chasen, F.N. 1940-04. A handlist of Malaysian mammals. Bulletin of Raffles Museum 15:1-209.	http://rmbr.nus.edu.sg/rbz/biblio/15/15brmiii-209.pdf	BMNH:Mamm:1947.1433	holotype	https://data.nhm.ac.uk/object/5d16b915-b486-4071-a0fb-37d7adb45873	"Jor, Batang Padang district, Perak, Malay States [= Malaysia]."	4.366667	101.3333	split from T. robustula	Tu, V. T., Csorba, G., Ruedi, M., Furey, N. M., Son, N. T., Thong, V. D., ... & Hassanin, A. (2017). Comparative phylogeography of bamboo bats of the genus Tylonycteris (Chiroptera, Vespertilionidae) in Southeast Asia. European Journal of Taxonomy, 274, 1-38.				India|Andaman and Nicobar Islands|China?|Myanmar|Vietnam|Thailand|Laos|Cambodia|Malaysia|Singapore	Asia	Indomalaya	NE	0	0	0	Tylonycteris_malayana	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Tylonycteris		malayana	Chasen	1940	0	Bull. Raffles Mus. Singapore	15:52	Malayan Bamboo Bat	eremtaga Srinivasulu, Srinivasulu, Srinivasulu & Jones, 2018.	Peninsular Malaysia	NE India, Bangladesh, Bhutan, Burma, Thailand, Cambodia, S Vietnam, peninsular Malaysia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	Not Evaluated	Previously included in robustula, but distinct, see Tu et al. (2017).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Tylonycteris malayana; Tylonycteris malayana; Tylonycteris malayana; Tylonycteris malayana; malayana; eremtaga; eremtaga; malayana; eremtaga; Vespertilion de Malaisie; Malaya-Bambusfledermaus; Tilonicteriode Malasia; Malayan Bamboo Bat; Malayan Greater Bamboo Bat; Malayan Bamboo Bat; Malayan Bamboo Bat; T. malayana