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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1642	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Micronycteris nicefori	Micronycteris nicefori	Micronycteris nicefori	Micronycteris nicefori	Micronycteris nicefori	Triaenops rufus	Trinycteris nicefori	Trinycteris nicefori	Trinycteris nicefori	Triaenops rufus	Trinycteris nicefori	Trinycteris nicefori	Trinycteris nicefori	Trinycteris nicefori	Trinycteris nicefori		[MSW2] Subgenus Trinycteris.; [HMW] Micronycteris (Trinycteris) nicefori Sanborn, 1949 , “Cucuta, Colombia .” This species is monotypic.; [batnames2022] See Simmons and Voss (1998).; [IUCN] This taxon was recognized as a full species by Peterson et al. (1995).; [batnames2023] See Simmons and Voss (1998).; [batnames2025_1.7] See Simmons and Voss (1998).						hypoleuca.			humbolti					nicefori	This taxon was recognized as a full species by Peterson et al. (1995).			nicefori	nicefori			nicefori (Sanborn, 1949)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Nicaragua – NE Peru,	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Micronycteris nicefori	Colombia, Norte de Santander, Cucuta.	Sanborn	1949	Fieldiana Zool., 31:230.	Distribution: Same as for subgenus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Niceforo's big-eared bat	S Mexico – NE Peru, Brazil, Trinidad	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Sanborn	1949	Fieldiana Zool., 31:230.	Subgenus Trinycteris.	Belize to N Colombia, Venezuela, Guianas, Amazonian Brazil, and Peru; Trinidad.	Colombia, Norte de Santander, Cucuta.		SANBORN	1949	Size fairly small (forearm length, 35 -41mm; condylobasal length, 18-20 mm).	Distribution: Same as for subgenus.	No subspecies.		73	species	M. nicefori	SANBORN	1949	Trinycteris	subgenus	Micronycteris nicefori				Size fairly small (forearm length, 35 -41mm; condylobasal length, 18-20 mm).	No subspecies.		4. M. nicefori SANBORN 1949.	4	NA																											4C3D87E8FFE76A59FF579A181D28BF49	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	539	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF86FF861343F61BF639F74C.xml	Trinycteris nicefori	Phyllostomidae	Trinycteris	nicefori		1949	Trinyctere de Niceforo @fr | Niceforo-Gro Rohrblattnase @de | Trinicterio de Niceforo @es | Niceforo's Bat @en	Micronycteris (Trinycteris) nicefori Sanborn, 1949 , “Cucuta, Colombia .” This species is monotypic.	S Mexico ( Chiapas ), Belize , Guatemala , S Nicaragua , Costa Rica , Panama , Colombia , Venezuela , the Guianas, W, N & SE Brazil , E Ecuador , E Peru , and NW Bolivia ; also on Trinidad I.	Head—body 55-68 mm, tail 9-13 mm, ear 16-20 mm, hindfoot 11-14 mm, forearm 36—41 mm ; weight 7-8-9 g. Dorsal fur of Niceforo’s Big-eared Bat can be grayish or orange-brown. Underparts are dark. Dorsal hairs have four bands: narrow and inconspicuous pale basal band followed by dark band, pale middle band, and dark tip. Pale median dorsalstripe can be present. Ears are pointed, whitish at bases, and blackish on remaining length. Noseleatis blackish, with base of horseshoe fused to upperlip. Chin has pair of longitudinal fleshy pads, converging ventrally. Wing membranes and uropatagium are blackish. Uropatagium is relatively long, naked, and V-shaped posteriorly. Calcar is smaller than foot. Tail perforates uropatagium dorsally. Rostrum is small, less than one-half the length of braincase. Zygomatic arches are slender. Basisphenoid pits are deep. Palate is relatively broad and long. Mandible condyle is above tooth row. Dental formula is 12,2. C1/1.,P 2/3. M 3/3 ( x2 ) = 34. I! isabout the same size as C'. P? touches C' and is separated from P* by a small gap. M' and M* are dilambdodont (W-shaped). P, is much smaller than and P,. Chromosomal complement has 2n = 28 and FN = 52, with ten pairs of metacentric or submetacentric and three pairs of subtelocentric autosomes. X-chromosome is submetacentric, and Y-chromosome is acrocentric.	Mature and secondary moist tropical forests, captured at elevations of 24— 460 m in Venezuela . In Mexico and Central America, Niceforo’s Big-eared Bat occurs along the humid Caribbean coast.	In Panama , 87-5% of diets of Niceforo’s Big-eared Bats was arthropods, and the remaining was fruit from species of Piper ( Piperaceae ).	No information.	Niceforo’s Big-eared Bat roosts in tree-cavities. It also uses artificial shelters; specimens were captured in a concrete building in Panama and an artificial tunnel. Echolocation calls recorded on Trinidad Island are characterized by multiharmonic FM sweeps of short duration (1:4-1-6 milliseconds) and high mean frequency (75-3 kHz).	On Trinidad Island , groups of up to twelve Niceforo’s Big-eared Bats were found in hollow trees, roosting with Seba’s Shorttailed Bats ( Carollia perspicillata ). In Colombia , individuals were captured roosting with Common Sword-nosed Bats ( Lonchorhina aurita ) and Tiny Big-eared Bats ( Micronycteris minuta ).	Classified as Least Concern on The IUCN Red List. Niceforo’s Big-eared Bat has a wide distribution and occurs in several protected areas throughout its distribution.	Baker (1979) | Baker, Genoways & Seyfarth (1981) | Delciellos et al. (2018.) | Giannini & Kalko (2004) | Goodwin & Greenhall (1961) | Handley (1966b, 1976) | Pérez-Consuegra et al. (2012) | Pio et al. (2010) | Rocha , Garbino & Aires (2013) | Sanborn (1949) | Simmons & Voss (1998) | Williams & Genoways (2008)	https://zenodo.org/record/6458837/files/figure.png	114. Niceforo’s Big-eared Bat Trinycteris nicefori French: Trinyctere de Niceforo / German: Niceforo-GroRohrblattnase / Spanish: Trinicterio de Niceforo Other common names: Niceforo's Bat Taxonomy. Micronycteris (Trinycteris) nicefori Sanborn, 1949 , “Cucuta, Colombia .” This species is monotypic. Distribution. S Mexico ( Chiapas ), Belize , Guatemala , S Nicaragua , Costa Rica , Panama , Colombia , Venezuela , the Guianas, W, N & SE Brazil , E Ecuador , E Peru , and NW Bolivia ; also on Trinidad I. Descriptive notes. Head—body 55-68 mm, tail 9-13 mm, ear 16-20 mm, hindfoot 11-14 mm, forearm 36—41 mm ; weight 7-8-9 g. Dorsal fur of Niceforo’s Big-eared Bat can be grayish or orange-brown. Underparts are dark. Dorsal hairs have four bands: narrow and inconspicuous pale basal band followed by dark band, pale middle band, and dark tip. Pale median dorsalstripe can be present. Ears are pointed, whitish at bases, and blackish on remaining length. Noseleatis blackish, with base of horseshoe fused to upperlip. Chin has pair of longitudinal fleshy pads, converging ventrally. Wing membranes and uropatagium are blackish. Uropatagium is relatively long, naked, and V-shaped posteriorly. Calcar is smaller than foot. Tail perforates uropatagium dorsally. Rostrum is small, less than one-half the length of braincase. Zygomatic arches are slender. Basisphenoid pits are deep. Palate is relatively broad and long. Mandible condyle is above tooth row. Dental formula is 12,2. C1/1.,P 2/3. M 3/3 ( x2 ) = 34. I! isabout the same size as C'. P? touches C' and is separated from P* by a small gap. M' and M* are dilambdodont (W-shaped). P, is much smaller than and P,. Chromosomal complement has 2n = 28 and FN = 52, with ten pairs of metacentric or submetacentric and three pairs of subtelocentric autosomes. X-chromosome is submetacentric, and Y-chromosome is acrocentric. Habitat. Mature and secondary moist tropical forests, captured at elevations of 24— 460 m in Venezuela . In Mexico and Central America, Niceforo’s Big-eared Bat occurs along the humid Caribbean coast. Food and Feeding. In Panama , 87-5% of diets of Niceforo’s Big-eared Bats was arthropods, and the remaining was fruit from species of Piper ( Piperaceae ). Breeding. No information. Activity patterns. Niceforo’s Big-eared Bat roosts in tree-cavities. It also uses artificial shelters; specimens were captured in a concrete building in Panama and an artificial tunnel. Echolocation calls recorded on Trinidad Island are characterized by multiharmonic FM sweeps of short duration (1:4-1-6 milliseconds) and high mean frequency (75-3 kHz). Movements, Home range and Social organization. On Trinidad Island , groups of up to twelve Niceforo’s Big-eared Bats were found in hollow trees, roosting with Seba’s Shorttailed Bats ( Carollia perspicillata ). In Colombia , individuals were captured roosting with Common Sword-nosed Bats ( Lonchorhina aurita ) and Tiny Big-eared Bats ( Micronycteris minuta ). Status and Conservation. Classified as Least Concern on The IUCN Red List. Niceforo’s Big-eared Bat has a wide distribution and occurs in several protected areas throughout its distribution. Bibliography. Baker (1979), Baker, Genoways & Seyfarth (1981), Delciellos et al. (2018.), Giannini & Kalko (2004), Goodwin & Greenhall (1961), Handley (1966b, 1976), Pérez-Consuegra et al. (2012), Pio et al. (2010), Rocha , Garbino & Aires (2013), Sanborn (1949), Simmons & Voss (1998), Williams & Genoways (2008).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Trinycteris nicefori	Trinycteris		nicefori	Sanborn	1949	1	Fieldiana Zool.	34:50:00	Niceforo's Bat	None.	Colombia, Norte de Santander, Cucuta.	Belize to Colombia, Venezuela, Guianas, Amazonian Brazil, Ecuador, and Peru; Bolivia; Trinidad.	Not listed.	Least Concern	See Simmons and Voss (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Trinycteris nicefori	23	Niceforo's Big-eared Bat	Niceforo's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	GLYPHONYCTERINAE	NA	Trinycteris	NA	nicefori	Sanborn	1949	1	Micronycteris_(Trinycteris)_nicefori	Sanborn, C. C. (1949). Bats of the genus Micronycteris and its subgenera. Fieldiana Zoology, 31(27), 230.	https://www.biodiversitylibrary.org/item/21663#page/21/mode/1up	FMNH 64266		"CÃºcuta, Colombia."			nicefori (Sanborn, 1949)	NA	NA	Mexico|Belize|Guatemala|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Trinycteris_nicefori	0	sciname match	Trinycteris_nicefori	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	40026	Triaenops rufus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Triaenops	rufus	Milne-Edwards, 1881	This taxon was recognized as a full species by Peterson et al. (1995).	20000000	Triaenops rufus	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution across Madagascar. It can tolerate some degree of habitat transformation, is relatively common, and is not thought to be declining fast enough to place it in a category of higher threat.	It is generally associated with tropical forests, but it may not need large areas of intact habitat and is not considered to be forest-dependent (Goodman et al. 2005). It is known from the spiny forest of the south-west and also in dry deciduous forest, and in the rainforest of the north-east of Madagascar (Goodman et al . 2005; Russ et al . 2003). It roosts in caves which it frequently cohabits with Triaenops furculus (Goodman et al . 2005; Olsson et al . 2006). It feeds mainly on lepidopterans (Rakotoarivelo et al . 2007) and frequently forages at the edge of relatively intact forest rather than inside (Kofoky et al . 2007; Rakotoarivelo et al . 2007).	There are no significant threats to this species even though its is occasionally hunted for food (Goodman 2006) and is often associated with forests that are subject to ongoing degradation (Russell et al. 2007).	The species is thought to be locally common in many parts of western Madagascar. A single roost in deciduous forest near the Onilahy River in southern Madagascar contained over 40,000 individuals (Olsson et al. 2006).	Unknown	This species is endemic to the island of Madagascar (Peterson et al. 1995), where it is widespread (Ranivo and Goodman 2006). It is not thought to be common on the central plateau of the eastern humid forest (Russ et al . 2003), but is common along the west coast (Goodman et al . 2005) and occurs between sea-level and 1,300 m Asl.		Terrestrial	This species is known to occur in many protected areas: RÃ©serve SpÃ©ciale dâ€™Ankarana, RÃ©serve SpÃ©ciale dâ€™Analamerana, Parc National du Tsingy de Bemaraha, Parc National de Namoroka, Parc National de Tsimanampetsotsa, Parc National dâ€™Ankarafantsika, Parc National dâ€™Isalo, RÃ©serve SpÃ©ciale dâ€™Ambohitantely, Parc National de Masoala and RÃ©serve Naturelle IntÃ©grale de Tsaratanana (Ranivo and Goodman 2006; Russ et al. 2003; Russell et al . 2007).	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Trinycteris		nicefori	Sanborn	1949	1	Fieldiana Zool.	34:50:00	Niceforo's Bat	None.	Colombia, Norte de Santander, Cucuta.	Belize to Colombia, Venezuela, Guianas, Amazonian Brazil, Ecuador, and Peru; Bolivia; Trinidad.	Not listed.	Least Concern	See Simmons and Voss (1998).	Trinycteris nicefori	1004923	23	Niceforo's Big-eared Bat	Niceforo's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	GLYPHONYCTERINAE	NA	Trinycteris	NA	nicefori	Sanborn	1949	1	Micronycteris_(Trinycteris)_nicefori	Sanborn, C. C. (1949). Bats of the genus Micronycteris and its subgenera. Fieldiana Zoology, 31(27), 230.	https://www.biodiversitylibrary.org/item/21663#page/21/mode/1up	FMNH 64266		"CÃºcuta, Colombia."			nicefori (Sanborn, 1949)	NA	NA				Mexico|Belize|Guatemala|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Trinycteris_nicefori	0	sciname match	Trinycteris_nicefori	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Trinycteris_nicefori	1004923	23	Niceforo's Big-eared Bat	Niceforo's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Glyphonycterinae	NA	Trinycteris	NA	nicefori	Sanborn	1	Micronycteris (Trinycteris) nicefori	Sanborn, C.C. 1949-04-29. Bats of the genus _Micronycteris_ and its subgenera. Fieldiana Zoology 31(27):215-233.	https://www.biodiversitylibrary.org/page/2871649	FMNH:Mamm:64266	holotype	http://portal.vertnet.org/o/fmnh/mammals?id=378cd3e9-1555-47e6-85e5-26b9e998cf42	"CÃºcuta, Colombia."			NA	NA				Mexico|Belize|Guatemala|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad and Tobago|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Trinycteris_nicefori	0	sciname match	Trinycteris_nicefori	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Trinycteris		nicefori	Sanborn	1949	1	Fieldiana Zool.	34:50:00	Niceforo's Bat	None.	Colombia, Norte de Santander, Cucuta.	Belize to Colombia, Venezuela, Guianas, Amazonian Brazil, Ecuador, and Peru; Bolivia; Trinidad.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/13381/22123365/' target='_blank'>Least Concern</a>	See Simmons and Voss (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Triaenops rufus; Trinycteris nicefori; Trinycteris nicefori; Trinycteris nicefori; Triaenops rufus; Trinycteris nicefori; humbolti; nicefori; Trinyctere de Niceforo; Niceforo-Gro Rohrblattnase; Trinicterio de Niceforo; Niceforo's Bat; Niceforo's Big-eared Bat; Niceforo's Bat; Rufous Trident Bat; Niceforo's Bat; Trinicteris nicefori; Micronycteris nicefori; T. nicefori
