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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1638	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Triaenops parvus	Triaenops parvus	Triaenops parvus	Triaenops parvus	Triaenops parvus	Triaenops parvus	Triaenops parvus	Triaenops parvus	Triaenops parvus		[HMW] Tríamops panms Benda 8c Vallo, 2009 , “ Republic of Yemen , Province of Al Mahra , oasis of Hawf (easternmost edge of the country), 16 39 ' N , 53 03 ’ E , 410 l. ” m a. s. This species is monotypic.; [MDD2022] recently described; moved from Hipposideridae to Rhinonycteridae; [IUCN] This species has been split from Triaenops persicus by Benda and Vallo (2009).; [MDD2023] recently described; moved from Hipposideridae to Rhinonycteridae; [MDD2025_2.0] recently described; moved from Hipposideridae to Rhinonycteridae; [MDD2025_2.2] recently described; moved from Hipposideridae to Rhinonycteridae														parvus	This species has been split from Triaenops persicus by Benda and Vallo (2009).			parvus	parvus			parvus Benda & Vallo, 2009						N/A																																								NA																											0383245F222097788B1EF4BAFCBDF99E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinonycteridae_194.pdf.imf	hash://md5/ffba5c272223977c8e73f509ffb1ffd0	206	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/83/24/0383245F222797788EC5F396F852FB4D.xml	Triaenops parvus	Rhinonycteridae	Triaenops	parvus	Benda & Vallo	2009	Tr aen du Yémen German @fr | Yemen Tr dont Lent-nosed Bat @en	Tríamops panms Benda 8c Vallo, 2009 , “ Republic of Yemen , Province of Al Mahra , oasis of Hawf (easternmost edge of the country), 16 39 ' N , 53 03 ’ E , 410 l. ” m a. s. This species is monotypic.	Endemic to a belt of c 300 km along coast of S Arabia; known from three sites in EYemen and one site in SW Oman .	Head-body 52-61 mm, tail 30-34 mm, ear 11-4-14 mm, forearm 44-48 mm; weight 4-9 g. No specific data are available for hindfoot length. The Yerneni Trident Bat is a small rhinonycterid and the smallest species of Triamops. Pelage is similarly tinged dorsally and ventrallv, being beige, yellowish brown, or brownish gray but never reddish or rusty, hairs can be darker around eyes. Wing membranes are dark brown; ears and noseleaf are pale pinkish (unpigmented), pale gray, or grayish brown. Noseleaf is large (width 7 -4-8 -3 mm) and rounded or pentagonal, with suap-like longitudinal projection that resembles an hourglass in outline, lying across anterior part of anterior leaf, above deep anterior medial emargination. Three tall pointed projections are present on posterior part of posterior leaf, and its anterior part in medial position has long lanceolated projection, with sharply pointed tip. Total width of three posterior projections is c.50% the width of posterior leaf. Numerous cells, separated by fleshy septa, scallop lateral parts of posterior leaf, with only one small cell in central position; lateral margins of two of three tall posterior projections are emarginated at their bases. Two supplementary leaflets occur lateral to noseleaf. Ears are short, wide, and pointed, with prominent step-like emargination on anterior margin. Second phalanx of fourth wing finger has unique transverse bone projection into membrane c.2 -5-3 mm long. Skull has very prominent nasal swellings laterally and posteriorly, and braincase is almost as high as rostrum and has very low but long sagittal crest. Rostrum is relatively short compared with congeners. Wide plates are present on dorsal sides of zygomatic bones. Greatest skull lengths are 16-8-18 mm, condylo-canine lengths are 14-4-15 -3 mm. zygomatic widths are 7 -6-8 mm, and upper tooth row (C -M‘) lengths are 5 -8-6-2 mm. f’ is bilobed; C is slender, with slight cingulum and large posterior secondary cusp, extending one-halfc' crown height; P’ is small and extruded; C and P‘ are in or nearly in contact; and M is only little reduced, with metacone. I, is bicuspid, l is tricuspid and larger) than, C, is slender, P 2 has about two-thirds the crown area and one-half the height of P‘, and M is unreduced. Baculum is gracile and c. 1 -6 mm long, with broad basal epiphysis and bifurcated distal epiphysis; has relatively narrow diaphvsis (c.8 % of baculum length), relatively short arms at its distal epiphysis (arm length 0 -18-0-24 mm; c. 17-20% of baculum length), and relatively narrow proximal epiphysis (basis width 0-3-0-4 mm; c.20-30% of baculum length).	Dry habitats of Sudanian savanna zone in southern Arabia and only in coastal areas with xeromorphic shrublands, surrounded by Acacia ( Fabaceae ) woodland savannas, from sea level to elevations of c. 650 m .	Diet of the Yemeni Trident Bat was studied only marginally; one sample of 20 fecal pellets from one individual contained only moths. Considering diet composition and knowledge of other species of Triamops, the Yemeni Trident Bat probably an aerial hawker. Foraging individuals were documented at various water bodies (e.g. livestock watering places, reservoir, and well) in oases and gardens. Yemeni Trident Bats are most commonly netted over and near open water.	No information.	Yemeni Trident Bats are probably unable to enter torpor and do not hibernate. Roosting was documented only once: subadult male was found in a natural cave in Yemen . They presumably leave clay roosts at sunset and forage for several hours; foraging bats were netted c. 2-3 hours after sunset. Echolocation consists of multiharmonic QCF calls where pulse is composed of long CF component, combined with FM component at end (CF-FM); terminating FM component is steep sweep of 3-10 kHz. Male calls last 6-6-11 -7 milliseconds (mean 8 milliseconds), interpulse intervals are 13-150-1 milliseconds (mean 49 -9 milliseconds); start frequencies are 93 -8-94 -9 kHz (mean 94 -3 kHz), peak frequencies are 93 -1-94 -2 kHz (mean 93-4 kHz), and end frequencies are 90 -3-92 -6 kHz (mean 9] -8 kHz).	No information.	Classified as Data Deficient on The IUCN Red List.	Benda & Va IO (2009) | Zdárská (2013)	https://zenodo.org/record/6611826/files/figure.png	3. Yemeni Trident Bat Triaenops parvus French: Tr aen du Yémen German: amen-Dre zahnb attnase Spanish: Rnonıcteno del Yemen Other common names: Yemen Tr dont Lent-nosed Bat Taxonomy. Tríamops panms Benda 8c Vallo, 2009 , “ Republic of Yemen , Province of Al Mahra , oasis of Hawf (easternmost edge of the country), 16 39 ' N , 53 03 ’ E , 410 l. ” m a. s. This species is monotypic. Distribution. Endemic to a belt of c 300 km along coast of S Arabia; known from three sites in EYemen and one site in SW Oman . Descriptive notes. Head-body 52-61 mm, tail 30-34 mm, ear 11-4-14 mm, forearm 44-48 mm; weight 4-9 g. No specific data are available for hindfoot length. The Yerneni Trident Bat is a small rhinonycterid and the smallest species of Triamops. Pelage is similarly tinged dorsally and ventrallv, being beige, yellowish brown, or brownish gray but never reddish or rusty, hairs can be darker around eyes. Wing membranes are dark brown; ears and noseleaf are pale pinkish (unpigmented), pale gray, or grayish brown. Noseleaf is large (width 7 -4-8 -3 mm) and rounded or pentagonal, with suap-like longitudinal projection that resembles an hourglass in outline, lying across anterior part of anterior leaf, above deep anterior medial emargination. Three tall pointed projections are present on posterior part of posterior leaf, and its anterior part in medial position has long lanceolated projection, with sharply pointed tip. Total width of three posterior projections is c.50% the width of posterior leaf. Numerous cells, separated by fleshy septa, scallop lateral parts of posterior leaf, with only one small cell in central position; lateral margins of two of three tall posterior projections are emarginated at their bases. Two supplementary leaflets occur lateral to noseleaf. Ears are short, wide, and pointed, with prominent step-like emargination on anterior margin. Second phalanx of fourth wing finger has unique transverse bone projection into membrane c.2 -5-3 mm long. Skull has very prominent nasal swellings laterally and posteriorly, and braincase is almost as high as rostrum and has very low but long sagittal crest. Rostrum is relatively short compared with congeners. Wide plates are present on dorsal sides of zygomatic bones. Greatest skull lengths are 16-8-18 mm, condylo-canine lengths are 14-4-15 -3 mm. zygomatic widths are 7 -6-8 mm, and upper tooth row (C -M‘) lengths are 5 -8-6-2 mm. f’ is bilobed; C is slender, with slight cingulum and large posterior secondary cusp, extending one-halfc' crown height; P’ is small and extruded; C and P‘ are in or nearly in contact; and M is only little reduced, with metacone. I, is bicuspid, l is tricuspid and larger) than, C, is slender, P 2 has about two-thirds the crown area and one-half the height of P‘, and M is unreduced. Baculum is gracile and c. 1 -6 mm long, with broad basal epiphysis and bifurcated distal epiphysis; has relatively narrow diaphvsis (c.8 % of baculum length), relatively short arms at its distal epiphysis (arm length 0 -18-0-24 mm; c. 17-20% of baculum length), and relatively narrow proximal epiphysis (basis width 0-3-0-4 mm; c.20-30% of baculum length). Habitat. Dry habitats of Sudanian savanna zone in southern Arabia and only in coastal areas with xeromorphic shrublands, surrounded by Acacia ( Fabaceae ) woodland savannas, from sea level to elevations of c. 650 m . Food and Feeding. Diet of the Yemeni Trident Bat was studied only marginally; one sample of 20 fecal pellets from one individual contained only moths. Considering diet composition and knowledge of other species of Triamops, the Yemeni Trident Bat probably an aerial hawker. Foraging individuals were documented at various water bodies (e.g. livestock watering places, reservoir, and well) in oases and gardens. Yemeni Trident Bats are most commonly netted over and near open water. Breeding. No information. Activity patterns. Yemeni Trident Bats are probably unable to enter torpor and do not hibernate. Roosting was documented only once: subadult male was found in a natural cave in Yemen . They presumably leave clay roosts at sunset and forage for several hours; foraging bats were netted c. 2-3 hours after sunset. Echolocation consists of multiharmonic QCF calls where pulse is composed of long CF component, combined with FM component at end (CF-FM); terminating FM component is steep sweep of 3-10 kHz. Male calls last 6-6-11 -7 milliseconds (mean 8 milliseconds), interpulse intervals are 13-150-1 milliseconds (mean 49 -9 milliseconds); start frequencies are 93 -8-94 -9 kHz (mean 94 -3 kHz), peak frequencies are 93 -1-94 -2 kHz (mean 93-4 kHz), and end frequencies are 90 -3-92 -6 kHz (mean 9] -8 kHz). Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Bibliography . Benda & Va IO (2009) Zdárská (2013)	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinonycteridae	Triaenops parvus	Triaenops		parvus	Benda & Vallo	2009	0	Folia Zoologica	58(1): 29	Little Trident Bat	None.	Republic of Yemen, Province of al Mahra, oasis of Hawf (easternmost edge of the country)	SE Yemen	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Triaenops parvus	23	Yemeni Trident Bat	Yemeni Trident Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINONYCTERIDAE	NA	NA	Triaenops	NA	parvus	Benda & Vallo	2009	0	Triaenops_parvus	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 29.	https://search.proquest.com/openview/7b0af8ba46bdcd2c9dbe53a842e506b2/1?pq-origsite=gscholar&cbl=27999	NMP 92270		"Republic of Yemen, Province of Al Mahra, oasis of Hawf (easternmost edge of the country), 16° 39' N, 53° 03' E, 410 m a. s. l."	16.65	53.05	parvus Benda & Vallo, 2009	recently described; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.	Yemen|Oman	Asia	Afrotropic	DD	0	0	0	Triaenops_parvus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Triaenops parvus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Triaenops	parvus	Benda &; Vallo, 2009	This species has been split from Triaenops persicus by Benda and Vallo (2009).	90000000	Triaenops parvus	Data Deficient		2017	2016-06-24 00:00:00 UTC	3.1	English	This is a newly established species without sufficient data on distribution and ecology, therefore listed as ;Data Deficient (DD). Given its small distribution, it could result in being listed as threatened once more information are gathered.	This bat inhabits savannah vegetation zone where suitable roosts and food are available. It roosts in caves. Similarly as in other species of the genus, this species is ;perhaps ;sedentary.	Main threats to this species are human disturbance in roost sites and pesticide use against insects.	This species is known from four sites and no other informations are available regarding its population dimensions and trend.	Unknown	This bat occurs in the semi-arid and savannah areas of south-eastern Yemen and most south-western Oman.		Terrestrial	No specific measures are known or are in place.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinonycteridae	Triaenops		parvus	Benda & Vallo	2009	0	Folia Zoologica	58(1): 29	Little Trident Bat	None.	Republic of Yemen, Province of al Mahra, oasis of Hawf (easternmost edge of the country)	SE Yemen	Not listed.	Data Deficient		Triaenops parvus	1004768	23	Yemeni Trident Bat	Yemeni Trident Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinonycteridae	NA	NA	Triaenops	NA	parvus	Benda & Vallo	2009	0	Triaenops_parvus	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 29.	https://search.proquest.com/openview/7b0af8ba46bdcd2c9dbe53a842e506b2/1?pq-origsite=gscholar&cbl=27999	NMP 92270		"Republic of Yemen, Province of Al Mahra, oasis of Hawf (easternmost edge of the country), 16° 39' N, 53° 03' E, 410 m a. s. l."	16.65	53.05	parvus Benda & Vallo, 2009	recently described; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.				Yemen|Oman	Asia	Afrotropic	DD	0	0	0	Triaenops_parvus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Triaenops_parvus	1004768	23	Yemeni Trident Bat	Yemeni Trident Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinonycteridae	NA	NA	Triaenops	NA	parvus	Benda & Vallo	0	Triaenops parvus	Benda, P. and Vallo, P. 2009. Taxonomic revision of the genus _Triaenops_ (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica 58(Monograph 1):1-45.	https://www.researchgate.net/publication/47057233_Taxonomic_revision_of_the_genus_Triaenops_Chiroptera_Hipposideridae_with_description_of_a_new_species_from_southern_Arabia_and_definitions_of_a_new_genus_and_tribe	NMPr P6V-092270	holotype		"Republic of Yemen, Province of Al Mahra, oasis of Hawf (easternmost edge of the country), 16° 39' N, 53° 03' E, 410 m a. s. l."	16.65	53.05	recently described; moved from Hipposideridae to Rhinonycteridae	Benda, P., & Vallo, P. (2009). Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica, 58, 1-45.|Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333.				Yemen|Oman	Asia	Afrotropic	DD	0	0	0	Triaenops_parvus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinonycteridae	Triaenops		parvus	Benda & Vallo	2009	0	Folia Zoologica	58(1): 29	Little Trident Bat	None.	Republic of Yemen, Province of al Mahra, oasis of Hawf (easternmost edge of the country)	SE Yemen	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/81082829/89457381/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Triaenops parvus; Triaenops parvus; Triaenops parvus; Triaenops parvus; Triaenops parvus; parvus; Tr aen du Yémen German; Yemen Tr dont Lent-nosed Bat; Yemeni Trident Bat; Yemeni Trident Leaf-nosed Bat; Little Trident Bat; T. parvus