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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1622	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Thoopterus suhaniahae	Thoopterus suhaniahae	Thoopterus suhaniahae	Thoopterus suhaniahae	Thoopterus suhaniahae	Thoopterus suhaniahae	Thoopterus suhaniahae	Thoopterus suhaniahae	Thoopterus suhaniahae		[HMW] Thoopterus suhaniahae Maryanto et al., 2012 , “ Indonesia , Central Sulawesi ... Lore Lindu National Park, Watumeta (23°57'S , 120° 18'07"E , 1125 m ).” This species is monotypic.; [MDD2022] recently described; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														suhaniahae				suhaniahae	suhaniahae			suhaniahae Maryanto, Yani, Prijono, & Wiantoro, 2012						N/A																																								NA																											67120269A438E04FFF4CF87DFAD9F24B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Thyropteridae_418.pdf.imf	hash://md5/9b2b7a11a43ae04dffe6ffc7ff89ffd6	75	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFD5F63B8996310CF87CFE41.xml	Thoopterus suhaniahae	Pteropodidae	Thoopterus	suhaniahae	Maryanto et al.	2012	Cynoptere de Suhaniah @fr | Suhaniah-Schwarzfligelflughund @de | Toptérido de Suhaniah @es	Thoopterus suhaniahae Maryanto et al., 2012 , “ Indonesia , Central Sulawesi ... Lore Lindu National Park, Watumeta (23°57'S , 120° 18'07"E , 1125 m ).” This species is monotypic.		Head-body 89- 109 mm (tail minute or absent), ear 16-19 mm , forearm 74-82 mm ; weight 52-100 g . Males are slightly larger than females in most dimensions. Head of Suhaniah’s Swift Fruit Bat has relatively long muzzle, nostrils are short and barely divergent, philtrum reaches upperlip, and lower lip has two large and flat triangular pads. Eyes are moderately large. Ears are short and rounded. Head pelage is darker than body, moderately long, dense, and dark grayish brown; nape has short guard hairs; dorsal pelage is long (c. 10 mm ), dark brown, and darker in rump. Uropatagium 1s narrow at center, with dorsal hairs extending also over tibia; underside of uropatagium is furred to its proximal one-half; and calcar is moderately long ( 10-11 mm ). Chest and belly are brownish gray, and flanks are washed with warmer brown. Wing membranes are black from sides of body and attach to second toe; index claw is present. Skull is robust, without basicranial deflection; rostrum is short and deep; orbit is large with obvious rim; zygomatic root is from above alveolar line; zygomais moderately thick and strongly arched throughout; and braincase is round. Dorsally, rostrum is broad; paranasal recesses are moderately visible; postorbital processes are directed posterolaterally; there is no postorbital foramen; postorbital constriction is long and obvious; braincase is oval; sagittal crest is marked; and nuchal crestis large, projecting posteriorly. Ventrally, palate is flat, with diverging tooth rows; postdental is moderately short, and sides are slightly convergent; end of post-dental palate is straight; and ectotympanic is small and wide anteriorly. Mandible is moderately thick; coronoid is tall, almost vertical with wide tip; condyle is very wide; and angle is distinct. There are 9-10 arched interdental palatal ridges, and posterior one-half is divided at middle, followed by three post-dental, divided ridges, the last one denticulate, possibly a last ill-defined ridge, denticulate, near end of post-dental palate. Upper incisors are conical, with I' slightly longer than I; C' is large, long, and almost straight, with anterointernal groove and strong inner cingulum, longer and bulkier in males; P' is small; posterior cheekteeth are large, very long, and of very similar length, with squarish outline and lingual cusps reduced; and M' is more rectangular. Lower incisors are small and crowded; C, is rather small and straight, with cingulum; P is well developed; posterior cheekteeth are tall; large premolars are very wide and squarish, with reduced lingual cusp and strong surface cusp; and M,is very low, small, and peg-like. Chromosomal complement has 2n = 38, FN = 64, and FNa = 60, with two pairs of metacentric, six pairs of submetacentric, four pairs of subtelocentric, and six pairs of telocentric autosomes. X-chromosome is metacentric.	Lowland, hill, and montane rainforest at elevations of 60-2100 m . Suhaniah’s Swift Fruit Bats are more common in primary forests at low and middle elevations.	Suhaniah’s Swift Fruit Bat is primarily frugivorous. Strong dentition suggests that some hard fruits are included in the diet.	No information.	No information.	No information.	Not assessed on The [UCN Red List. Suhaniah’s Swift Fruit Bat is reported to be one of the most abundant bat species in Sulawesi . It is likely affected by hunting in North and Central Sulawesi and deforestation throughoutits distribution.	Maryanto (2004) | Mubarok et al. (2018)		24. Suhaniah’s Swift Fruit Bat Thoopterus suhaniahae French: Cynoptere de Suhaniah / German: Suhaniah-Schwarzfligelflughund / Spanish: Toptérido de Suhaniah Taxonomy. Thoopterus suhaniahae Maryanto et al., 2012 , “ Indonesia , Central Sulawesi ... Lore Lindu National Park, Watumeta (23°57'S , 120° 18'07"E , 1125 m ).” This species is monotypic. Distribution. Talaud Is and Sulawesi , including nearby Wowoni L Descriptive notes. Head-body 89- 109 mm (tail minute or absent), ear 16-19 mm , forearm 74-82 mm ; weight 52-100 g . Males are slightly larger than females in most dimensions. Head of Suhaniah’s Swift Fruit Bat has relatively long muzzle, nostrils are short and barely divergent, philtrum reaches upperlip, and lower lip has two large and flat triangular pads. Eyes are moderately large. Ears are short and rounded. Head pelage is darker than body, moderately long, dense, and dark grayish brown; nape has short guard hairs; dorsal pelage is long (c. 10 mm ), dark brown, and darker in rump. Uropatagium 1s narrow at center, with dorsal hairs extending also over tibia; underside of uropatagium is furred to its proximal one-half; and calcar is moderately long ( 10-11 mm ). Chest and belly are brownish gray, and flanks are washed with warmer brown. Wing membranes are black from sides of body and attach to second toe; index claw is present. Skull is robust, without basicranial deflection; rostrum is short and deep; orbit is large with obvious rim; zygomatic root is from above alveolar line; zygomais moderately thick and strongly arched throughout; and braincase is round. Dorsally, rostrum is broad; paranasal recesses are moderately visible; postorbital processes are directed posterolaterally; there is no postorbital foramen; postorbital constriction is long and obvious; braincase is oval; sagittal crest is marked; and nuchal crestis large, projecting posteriorly. Ventrally, palate is flat, with diverging tooth rows; postdental is moderately short, and sides are slightly convergent; end of post-dental palate is straight; and ectotympanic is small and wide anteriorly. Mandible is moderately thick; coronoid is tall, almost vertical with wide tip; condyle is very wide; and angle is distinct. There are 9-10 arched interdental palatal ridges, and posterior one-half is divided at middle, followed by three post-dental, divided ridges, the last one denticulate, possibly a last ill-defined ridge, denticulate, near end of post-dental palate. Upper incisors are conical, with I' slightly longer than I; C' is large, long, and almost straight, with anterointernal groove and strong inner cingulum, longer and bulkier in males; P' is small; posterior cheekteeth are large, very long, and of very similar length, with squarish outline and lingual cusps reduced; and M' is more rectangular. Lower incisors are small and crowded; C, is rather small and straight, with cingulum; P is well developed; posterior cheekteeth are tall; large premolars are very wide and squarish, with reduced lingual cusp and strong surface cusp; and M,is very low, small, and peg-like. Chromosomal complement has 2n = 38, FN = 64, and FNa = 60, with two pairs of metacentric, six pairs of submetacentric, four pairs of subtelocentric, and six pairs of telocentric autosomes. X-chromosome is metacentric. Habitat. Lowland, hill, and montane rainforest at elevations of 60-2100 m . Suhaniah’s Swift Fruit Bats are more common in primary forests at low and middle elevations. Food and Feeding. Suhaniah’s Swift Fruit Bat is primarily frugivorous. Strong dentition suggests that some hard fruits are included in the diet. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The [UCN Red List. Suhaniah’s Swift Fruit Bat is reported to be one of the most abundant bat species in Sulawesi . It is likely affected by hunting in North and Central Sulawesi and deforestation throughoutits distribution. Bibliography. Maryanto (2004), Mubarok et al. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Thoopterus suhaniahae	Thoopterus		suhaniahae	Maryanto, Yani, Prijono & Wiantoro	2012	0	Rec. West. Aust. Mus.	28:16:00	Suhaniah&apos;s Fruit Bat	None.	Indonesia, C. Sulawesi, Lore Lindu National Park, Watumeta	Sulawesi, Talaud, and Wowoni Isls.	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Thoopterus suhaniahae	23	Suhaniah's Swift Fruit Bat	Suhaniah's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	CYNOPTERINAE	BALIONYCTERINI	Thoopterus	NA	suhaniahae	Maryanto, Yani, Prijono, & Wiantoro	2012	0						"Indonesia, Central Sulawesi â€¦ Lore Lindu National Park, Watumeta (1Â°23'57"S, 120Â°18'07"E, 1125 m)."	-1.399	120.3	suhaniahae Maryanto, Yani, Prijono, & Wiantoro, 2012	recently described	Maryanto, I., Yani, M., Prijono, S. N., & Wiantoro, S. (2012). A new species of fruit bat (Megachiroptera: Pteropodidae: Thoopterus) from Sulawesi and adjacent islands, Indonesia. Records of the Western Australian Museum, 27(1), 68-84.	Indonesia	Asia	Indomalaya|Australasia/Oceania	LC	0	0	0	Thoopterus_suhaniahae	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Thoopterus suhaniahae	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Thoopterus	suhaniahae	Maryanto, Yani, Prijono &; Wiantoro, 2012		80000000	Thoopterus suhaniahae	Least Concern		2020	2019-05-14 00:00:00 UTC	3.1	English	Listed as Least Concern because its extent of occurrence is greater than 20,000 kmÂ², it is very common, and its populations are not declining fast enough for listing in a more threatened category.	This species is generally found in the primary forest at elevations up to 2,100 m asl (Maryanto et al. 2012)	Deforestation is occurring throughout the species' range.	This species is common and widely distributed in Sulawesi (Maryanto et al. 2012).	Stable	This species known from greater Sulawesi, Talaud, and Wowoni Islands, Indonesia (Maryanto et al. 2012).		Terrestrial		Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Thoopterus		suhaniahae	Maryanto, Yani, Prijono & Wiantoro	2012	0	Rec. West. Aust. Mus.	28:16:00	Suhaniah&apos;s Fruit Bat	None.	Indonesia, C. Sulawesi, Lore Lindu National Park, Watumeta	Sulawesi, Talaud, and Wowoni Isls.	Not listed.	Least Concern		Thoopterus suhaniahae	1004373	23	Suhaniah's Swift Fruit Bat	Suhaniah's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	CYNOPTERINAE	BALIONYCTERINI	Thoopterus	NA	suhaniahae	Maryanto, Yani, Prijono, & Wiantoro	2012	0						"Indonesia, Central Sulawesi â€¦ Lore Lindu National Park, Watumeta (1Â°23'57"S, 120Â°18'07"E, 1125 m)."	-1.39917	120.3019	suhaniahae Maryanto, Yani, Prijono, & Wiantoro, 2012	recently described	Maryanto, I., Yani, M., Prijono, S. N., & Wiantoro, S. (2012). A new species of fruit bat (Megachiroptera: Pteropodidae: Thoopterus) from Sulawesi and adjacent islands, Indonesia. Records of the Western Australian Museum, 27(1), 68-84.				Indonesia	Asia	Indomalaya|Australasia/Oceania	LC	0	0	0	Thoopterus_suhaniahae	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Thoopterus_suhaniahae	1004373	23	Suhaniah's Swift Fruit Bat	Suhaniah's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Cynopterinae	Balionycterini	Thoopterus	NA	suhaniahae	Maryanto, Yani, Prijono, & Wiantoro	0	Thoopterus suhaniahae	Maryanto, I., Yani, M., Prijono, S.N. and Wiantoro, S. 2012. A new species of fruit bat (Megachiroptera: Pteropodidae: _Thoopterus_) from Sulawesi and adjacent islands, Indonesia. Records of the Western Australian Museum 27:68-84.	https://www.biodiversitylibrary.org/page/59832697	MZB 23390	holotype		"Indonesia, Central Sulawesi â€¦ Lore Lindu National Park, Watumeta (1Â°23'57"S, 120Â°18'07"E, 1125 m)."	-1.39917	120.3019	recently described	Maryanto, I., Yani, M., Prijono, S. N., & Wiantoro, S. (2012). A new species of fruit bat (Megachiroptera: Pteropodidae: Thoopterus) from Sulawesi and adjacent islands, Indonesia. Records of the Western Australian Museum, 27(1), 68-84.				Indonesia	Asia	Australasia	LC	0	0	0	Thoopterus_suhaniahae	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Thoopterus		suhaniahae	Maryanto, Yani, Prijono & Wiantoro	2012	0	Rec. West. Aust. Mus.	28:16:00	Suhaniah&apos;s Fruit Bat	None.	Indonesia, C. Sulawesi, Lore Lindu National Park, Watumeta	Sulawesi, Talaud, and Wowoni Isls.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84463939/84463943/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Thoopterus suhaniahae; Thoopterus suhaniahae; Thoopterus suhaniahae; Thoopterus suhaniahae; Thoopterus suhaniahae; suhaniahae; Cynoptere de Suhaniah; Suhaniah-Schwarzfligelflughund; Toptérido de Suhaniah; Suhaniah's Swift Fruit Bat; Suhaniah's Fruit Bat; Suhaniah&apos;s Fruit Bat; T. suhaniahae
