http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1607	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Taphozous hildergardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae	Taphozous hildegardeae		[MSW2] Subgenus Taphozous.; [MSW3] Subgenus Taphozous. See Colket and Wilson (1998).; [HMW] Taphozous hildegardeaeThomas, 1909 , “ Rabai , 700 ’ [= 213 m ],” Mombasa District , Kenya . Taphozous hildegardeae is in the subgenus Taphozous . Some authorities question locality records attributable to central Kenya because. hildegardeae is otherwise known only from coastal distribution and habitats; however, these inland records coincide with coastal vegetation along the Tana River, possibly offering suitable habitat Monotypic.; [batnames2022] See Colket and Wilson (1998).; [batnames2023] See Colket and Wilson (1998).; [batnames2025_1.7] See Colket and Wilson (1998).														hildegardeae				hildegardeae	hildegardeae, hildergardeae			hildegardeae O. Thomas, 1909|hildergardeae Corbet & J. Edwards Hill, 1980 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Hildegarde's tomb bat	Kenya	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Taphozous hildegardeae	Kenya, Coast Province, Rabai (near Mombasa).	Thomas	1909	Ann. Mag. Nat. Hist., ser. 8, 4:98.	Dis tribution: Confined to a small area of southeastern Kenya and northeastern Tanzania including Zan zibar island.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Hildegarde's tomb bat	Kenya, NE Tanzania, Zanzibar	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1909	Ann. Mag. Nat. Hist., ser. 8, 4:98.	Subgenus Taphozous.	Kenya, NE Tanzania, Zanzibar.	Kenya, Coast Province, Rabai (near Mombasa).		THOMAS	1909	Gular sac ab sent. A black beard-like throat patch in males. Skull relatively broad. Fur relatively pale in color. Size medium (forearm length, 63-70 mm).	Dis tribution: Confined to a small area of southeastern Kenya and northeastern Tanzania including Zan zibar island.	No subspecies.		42	species	T. hildegardeae	THOMAS	1909	Taphozous	subgenus	Taphozous hildegardeae				Gular sac ab sent. A black beard-like throat patch in males. Skull relatively broad. Fur relatively pale in color. Size medium (forearm length, 63-70 mm).	No subspecies.		1. T. hildegardeae THOMAS 1909.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Taphozoinae		Taphozous hildegardeae	Taphozous	Taphozous	hildegardeae	Thomas		1909		Ann. Mag. Nat. Hist., ser. 8	4		98		Hildegarde's Tomb Bat	Kenya, Coast Province, Rabai (near Mombassa).	Kenya, NE Tanzania, Zanzibar.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Subgenus Taphozous. See Colket and Wilson (1998).	03D587F2FFC94C02F8FA36ADF71EF0C4	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	352	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFCC4C00F834333AFC4CFC29.xml	Taphozous hildegardeae	Emballonuridae	Taphozous	hildegardeae	Thomas	1909	Hildegarde’s Tomb Bat @en | Taphien de Hildegarde @fr | Hildegarde-Grabfledermaus @de | Tafozode Hildegarde @es | White-wingedTomb Bat @en	Taphozous hildegardeaeThomas, 1909 , “ Rabai , 700 ’ [= 213 m ],” Mombasa District , Kenya . Taphozous hildegardeae is in the subgenus Taphozous . Some authorities question locality records attributable to central Kenya because. hildegardeae is otherwise known only from coastal distribution and habitats; however, these inland records coincide with coastal vegetation along the Tana River, possibly offering suitable habitat Monotypic.	E Africa along coast of SE Kenya and NE Tanzania (from Lower Tana River S to Dar es Salaam), also on Pemba and Unguja Is (Zanzibar Archipelago); recorded in C Kenya, but verification of museum vouchers and additional fieldwork could clarify distribution there.	Head-body 79-82 mm, tail 25-29 mm, ear 13-22 mm, hindfoot 11— 18 mm, forearm 63-66 mm; weight 24-36 g (largest weights are for males with heavy fat deposits). Hairs on dorsum of Hildegarde’s Tomb Bat are tricolored, with pale brown above white bases and chocolate-brown tips; venter is white, with lightly frosted brown scattered tips. Exposed skin on limbs and other body parts is brown. Dorsal wing surfaces and uropatagium are brown, but distal part of wing membranes is white. Sparse fur on dorsal and ventral sides of flight membranes is white. Profile of head is concave. Both sexes have radio-metacarpal sacs. Although some sources state that glandular gular pouches are absent, others have observed these glands on mature males. Breeding age males have distinct black beards, covering undersides of their throats.	Tropical dry forests in very restricted coastal areas from sea level to elevations of C.214 m. Hildegarde’s Tomb Bat also might occur further inland in gallery forests along the Tana River.	Hildegarde’s Tomb Bat eats species of Orthoptera and Lepidoptera .	In coastal Kenya, reproductive cycle ofHildegarde’s Tomb Bat closely corresponds to regional seasonality in rainfall. Males have pronounced bimodal cycles of fat deposition, corresponding to relative insect abundance during dual rainy seasons in Kenya. During active courtship and copulation in the first dry season of the year when moth abundance is lowest, males deplete body fat; fat accumulation in males is greatest in the long rainy season in Kenya. Females do not put on appreciable fat deposits, and even near-term pregnant females do not reach large seasonal body weights of males. A secondary peak in body weight and development of male sexual glands are associated with “short rains” in October-December. Second rainy season apparently does not result in reproduction but appears to allow maintenance of year-round harems that might reflect a vestigial polyestrous reproductive cycle.	Hildegard’s Tomb Bat is nocturnal and uses coral sea caves as roosts along the coast. It emerges from day roosts well after dark.	Hildegard’s Tomb Bat is polygynous, and males maintain roost territories to defend their female harems. In some larger harems, multiple males defend a group’s roosting site. Distance between harems is slightly greater than striking distance of the forearm. Adult males defend harems from intruders by wing flicking, vocalizations, scent marking, postural changes, and aggressive attacks. Males identify females as harem members by anal-genital sniffing. Males also mark harem females by pressing throat glands over females’ dorsum and mark themselves by rubbing their folded wings and forearms over throat glands. Bachelor males andjuveniles roost separately from harems. Colonies of hundreds to c.1000 individuals occur in caves, suggesting that local foraging movements occur at considerable distances from roosting sites. Hildegard’s Tomb Bats often share lit parts of caves with African Sheath-tailed Bats ( Coleura afra ). The cimicid bug Loxaspis miranda and the bat fly Brachytarsina aUuaudi have been identified as common ectoparasites in the larger colonies of Hildegard’s Tomb Bats.	Classified as Vulnerable on The IUCNRed List. Extent of occurrence of Hildegard’s Tomb Bat is less than 20,000 km2, its distribution is severely fragmented, and there is continuing decline in extent and quality of its forest habitat. It has been recorded from fewer than ten coastal localities. No distribution-wide population estimates are available, but surveys in Kenya and Tanzania in 1988 documented more than 130, 300, and 1000 individuals in three caves. Hildegard’s Tomb Bat likely is declining due to loss of habitat in coastal dry forests and disturbance of cave sites. Updated surveys of cave populations are needed to reassess it conservation status.	ACR (2017) | Colket & Wilson (1998) | Harrison (1962) | Kock (1974) | McWilliam (1988b) | Mickleburgh , Hutson & Bergmans (2008b) | Usinger (1966)	https://zenodo.org/record/3747912/files/figure.png	5 . Hildegarde’s Tomb Bat Taphozous hildegardeae French: Taphien de Hildegarde / German: Hildegarde-Grabfledermaus I Spanish: Tafozo de Hildegarde Other common names: White-wingedTomb Bat Taxonomy . Taphozous hildegardeaeThomas, 1909 , “ Rabai , 700 ’ [= 213 m ],” Mombasa District , Kenya . Taphozous hildegardeae is in the subgenus Taphozous . Some authorities question locality records attributable to central Kenya because. hildegardeae is otherwise known only from coastal distribution and habitats; however, these inland records coincide with coastal vegetation along the Tana River, possibly offering suitable habitat Monotypic. Distribution. E Africa along coast of SE Kenya and NE Tanzania (from Lower Tana River S to Dar es Salaam), also on Pemba and Unguja Is (Zanzibar Archipelago); recorded in C Kenya, but verification of museum vouchers and additional fieldwork could clarify distribution there. Descriptive notes. Head-body 79-82 mm, tail 25-29 mm, ear 13-22 mm, hindfoot 11— 18 mm, forearm 63-66 mm; weight 24-36 g (largest weights are for males with heavy fat deposits). Hairs on dorsum of Hildegarde’s Tomb Bat are tricolored, with pale brown above white bases and chocolate-brown tips; venter is white, with lightly frosted brown scattered tips. Exposed skin on limbs and other body parts is brown. Dorsal wing surfaces and uropatagium are brown, but distal part of wing membranes is white. Sparse fur on dorsal and ventral sides of flight membranes is white. Profile of head is concave. Both sexes have radio-metacarpal sacs. Although some sources state that glandular gular pouches are absent, others have observed these glands on mature males. Breeding age males have distinct black beards, covering undersides of their throats. Habitat . Tropical dry forests in very restricted coastal areas from sea level to elevations of C.214 m. Hildegarde’s Tomb Bat also might occur further inland in gallery forests along the Tana River. Food and Feeding . Hildegarde’s Tomb Bat eats species of Orthoptera and Lepidoptera . Breeding . In coastal Kenya, reproductive cycle ofHildegarde’s Tomb Bat closely corresponds to regional seasonality in rainfall. Males have pronounced bimodal cycles of fat deposition, corresponding to relative insect abundance during dual rainy seasons in Kenya. During active courtship and copulation in the first dry season of the year when moth abundance is lowest, males deplete body fat; fat accumulation in males is greatest in the long rainy season in Kenya. Females do not put on appreciable fat deposits, and even near-term pregnant females do not reach large seasonal body weights of males. A secondary peak in body weight and development of male sexual glands are associated with “short rains” in October-December. Second rainy season apparently does not result in reproduction but appears to allow maintenance of year-round harems that might reflect a vestigial polyestrous reproductive cycle. Activity patterns. Hildegard’s Tomb Bat is nocturnal and uses coral sea caves as roosts along the coast. It emerges from day roosts well after dark. Movements, Home range and Social organization. Hildegard’s Tomb Bat is polygynous, and males maintain roost territories to defend their female harems. In some larger harems, multiple males defend a group’s roosting site. Distance between harems is slightly greater than striking distance of the forearm. Adult males defend harems from intruders by wing flicking, vocalizations, scent marking, postural changes, and aggressive attacks. Males identify females as harem members by anal-genital sniffing. Males also mark harem females by pressing throat glands over females’ dorsum and mark themselves by rubbing their folded wings and forearms over throat glands. Bachelor males andjuveniles roost separately from harems. Colonies of hundreds to c.1000 individuals occur in caves, suggesting that local foraging movements occur at considerable distances from roosting sites. Hildegard’s Tomb Bats often share lit parts of caves with African Sheath-tailed Bats ( Coleura afra ). The cimicid bug Loxaspis miranda and the bat fly Brachytarsina aUuaudi have been identified as common ectoparasites in the larger colonies of Hildegard’s Tomb Bats. Status and Conservation . Classified as Vulnerable on The IUCNRed List. Extent of occurrence of Hildegard’s Tomb Bat is less than 20,000 km2, its distribution is severely fragmented, and there is continuing decline in extent and quality of its forest habitat. It has been recorded from fewer than ten coastal localities. No distribution-wide population estimates are available, but surveys in Kenya and Tanzania in 1988 documented more than 130, 300, and 1000 individuals in three caves. Hildegard’s Tomb Bat likely is declining due to loss of habitat in coastal dry forests and disturbance of cave sites. Updated surveys of cave populations are needed to reassess it conservation status. Bibliography. ACR (2017), Colket & Wilson (1998), Harrison (1962), Kock (1974), McWilliam (1988b), Mickleburgh , Hutson & Bergmans (2008b), Usinger (1966).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Taphozous hildegardeae	Taphozous		hildegardeae	Thomas	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 98	Hildegarde's Tomb Bat	None.	Kenya, Coast Province, Rabai (near Mombassa).	Kenya, NE Tanzania, Zanzibar.	Not listed.	Endangered	See Colket and Wilson (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Taphozous hildegardeae	23	Hildegarde's Tomb Bat	White-winged Tomb Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	TAPHOZOINAE	NA	Taphozous	Taphozous	hildegardeae	O. Thomas	1909	0	Taphozous_hildegardeae	Thomas, O. (1909). New African small mammals in the British Museum Collection. Annals and Magazine of Natural History, ser. 8, 4, 98.	https://www.biodiversitylibrary.org/item/71907#page/124/mode/1up	BM 1909.6.12.7		"Rabai, 700 ' [= 213 m]," Mombasa District, Kenya.			hildegardeae O. Thomas, 1909	NA	NA	Tanzania|Kenya	Africa	Afrotropic	EN	0	0	0	Taphozous_hildegardeae	0	sciname match	Taphozous_hildegardeae	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	21456	Taphozous hildegardeae	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Taphozous	hildegardeae	Thomas, 1909		20000000	Taphozous hildegardeae	Endangered	B2ab(i,ii,iii)	2020	2019-06-22 00:00:00 UTC	3.1	English	This bat is listed as Vulnerable under criterion B2ab(i,ii,iii) because its area of occupancy (AOO) is estimated at 48 kmÂ² (based on 12 known roost caves across its distribution), a severely fragmented population, and continuing decline in the extent and quality of its cave roost and forest habitat, leading to the reduction of its EOO and area of occupancy (AOO).	Hildegarde's Tomb Bat is a cave-dependent species that appears to be restricted to tropical dry coastal forests (coastal forest mosaic), with a restricted range along the coast of Kenya and Tanzania, including on Pemba and Zanzibar Islands (McWilliam and Happold 2013). Along the coast, it roosts colonially in coral reef caves in remnant coastal forests but likely to forage in adjacent scrublands and plantations (McWilliam and Happold 2013). The species usually roosts adjacent to or together with Coleura afra , ;Miniopterus minor , Rousettus aegyptiacus , Triaenops afer , Macronycteris gigas , M. vittata , Hipposideros caffer and Cloeotis percivali (McWilliam and Happold 2013, Makori 2015, Musila et al. 2019). Taphozous hildegardeae is insectivorous, feeding on Orthoptera and Lepidoptera and seasonal variation in insect prey has been observed (McWilliam and Happold 2013). Copulation and ovulation occur when rainfall is lowest (late July to early August), and females give birth during the second wet-season (mid- to late-December). There have been reports of pronounced delay in giving birth, with some females still pregnant in late January or early February, following a severe dry season when body condition in females likely to be very poor and linked to delayed implantation (McWilliam and Happold 2013).	This species is endemic to Kenya and Tanzania, with a small geographic range. Disturbance, including tourism and recreational activities of the coral caves on which it depends, and loss of forest habitat along the coastal strip, could result in population declines. Because the roost sites in coral caves are on unprotected community lands, at least in Kenya, this in itself is a threat because the future of these caves is uncertain. The land on which these caves are located are at risk of being converted to farmlands and the caves being closed off or destroyed.	The overall global population is thought to be declining due to cave roost and forest degradation and loss. Large colonies roosting at Makuruhu and Kaboga Caves near Watamu town in Kilifi County has been estimated to include over 21,240 and 3,800 individuals, respectively (Makori 2015, Musila et al. , 2019, P. Webala, unpublished data). Another coral cave, called Kuruwitu, just a few kilometres north of Mombasa has about 2,000 individuals (P. Webala, unpublished data). The Fikirini/Mdenyenye Caves in Kwale County, south coast of Kenya are estimated to have over 20,000 (P. Webala, unpublished data). A few individuals in the hundreds are also resident at the historical Shimoni Slave Cave on Kenyaâ€™s south coast. Therefore, Kenyan populations at northern and southern coastal Kenya are probably stable at the moment. However, as these caves are unprotected on private and community lands they are at great risk of development and loss. Two large populations in coastal Kenya were estimated at about 1,000 to 2,000 individuals (McWilliam and Happold 2013). During a survey in Kenya and Tanzania in 1988, population estimates from three caves surveyed yielded more than 1,500 individuals (1,000+, 300+, 130+) (Susan M. Pont, personal observation).	Decreasing	Hildegarde's Tomb Bat is endemic to Kenya and Tanzania, occurring in the coastal zone of Kenya and north-east Tanzania (from Malindi south to Dar es Salaam), as well as on Zanzibar Island (Kityo et al . 2009). It has been recorded from more than 15 coastal localities. Recently, the species was documented in the hundreds to thousands in a number of coral caves in Kilifi County, northern coastal Kenya (e.g., Kaboga, Kuruwitu Makururu Caves, Makori 2015, Musila et al . 2019, P Webala, unpublished data). It is present at suitable sites throughout the year, with some local movement (McWilliam and Happold 2013). However, a population of migrating individuals has been observed inhabiting a coastal cave in Kenya, appearing in November and disappearing in May (McWilliam and Happold 2013). There are two central Kenya records collected by Harrison (1962) that have been questioned (Kock (1974), since these are the only localities not along the East African coast. Although the questioned locations do coincide with coastal vegetation along the Tana River, which could afford this species suitable habitat (Colket and Wilson 1998), they are excluded from the species distribution for this assessment.		Terrestrial	It may occur in some protected areas in Tanzania. In Kenya it occurs on unprotected community lands (Makori 2015, Musila et al. 2019, P. Webala, personal observation) and therefore stricter conservation of roosting and breeding sites is needed, and positive community engagement. Potential to protect these species through gating important maternity roosts and restoring degraded habitat within close proximity to these important roosts. Other research needs include: better understanding of the species distribution, including confirmation of records on Zanzibar and Pemba Islands and current survey to update historical records, baseline and monitoring data to understand population size and trends, including roost use throughout the year, and more information on significant threats and potential mitigation strategies.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Taphozous		hildegardeae	Thomas	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 98	Hildegarde's Tomb Bat	None.	Kenya, Coast Province, Rabai (near Mombassa).	Kenya, NE Tanzania, Zanzibar.	Not listed.	Endangered	See Colket and Wilson (1998).	Taphozous hildegardeae	1004821	23	Hildegarde's Tomb Bat	White-winged Tomb Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	TAPHOZOINAE	NA	Taphozous	Taphozous	hildegardeae	O. Thomas	1909	0	Taphozous_hildegardeae	Thomas, O. (1909). New African small mammals in the British Museum Collection. Annals and Magazine of Natural History, ser. 8, 4, 98.	https://www.biodiversitylibrary.org/item/71907#page/124/mode/1up	BM 1909.6.12.7		"Rabai, 700 ' [= 213 m]," Mombasa District, Kenya.			hildegardeae O. Thomas, 1909	NA	NA				Tanzania|Kenya	Africa	Afrotropic	EN	0	0	0	Taphozous_hildegardeae	0	sciname match	Taphozous_hildegardeae	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Taphozous_hildegardeae	1004821	23	Hildegarde's Tomb Bat	White-winged Tomb Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Taphozoinae	NA	Taphozous	NA	hildegardeae	O. Thomas	0	Taphozous hildegardeÃ¦	Thomas, O. 1909-08-01. New African small mammals in the British Museum. Annals and Magazine of Natural History (8)4(20):98-112.	https://www.biodiversitylibrary.org/page/22097473	BMNH:Mamm:1909.6.12.7	holotype	https://data.nhm.ac.uk/object/314c8922-282c-4bb2-943c-97dddaed1325	"Rabai, 700 ' [= 213 m]," Mombasa District, Kenya.			NA	NA				Tanzania|Kenya	Africa	Afrotropic	EN	0	0	0	Taphozous_hildegardeae	0	sciname match	Taphozous_hildegardeae	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Taphozous		hildegardeae	Thomas	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 98	Hildegarde's Tomb Bat	None.	Kenya, Coast Province, Rabai (near Mombassa).	Kenya, NE Tanzania, Zanzibar.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/21456/22111960/' target='_blank'>Endangered</a>	See Colket and Wilson (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Taphozous hildegardeae; Taphozous hildegardeae; Taphozous hildegardeae; Taphozous hildegardeae; Taphozous hildegardeae; Taphozous hildegardeae; hildegardeae; Hildegarde’s Tomb Bat; Taphien de Hildegarde; Hildegarde-Grabfledermaus; Tafozode Hildegarde; White-wingedTomb Bat; Hildegarde's Tomb Bat; White-winged Tomb Bat; Hildegarde's Tomb Bat; Hildegarde's Tomb Bat; T. hildegardeae
