This small species related to Myotinae was considered as a subspecies of more widespread taxa such as Myotis muricola (e.g. Simmons 2005) or M. mystacinus (e.g. Tate 1941). However, global molecular studies (Stadelmann et al. 2007, Lack et al . 2010, Ruedi et al. 2013) showed that this species actually differs from any other known Myotis and appears in a basal position in the radiation of Myotinae (Ruedi et al . 2013). In addition, unique dental and cranial characters indicate that this species actually belongs to a distinct genus that is morphologically closely related to Submyotodon, a fossil Myotinae of Miocene age (Ziegler 2003). To reflect these large differences compared to all other species of Myotis, it is currently classified as S. latirostris (Ruedi et al . 2015) and is endemic to Taiwan. Other recent species belonging to the genus Submyotodon are found in the Himalayan region.
; [batnames2023] Does not include caliginosus , blanfordi , or moupinensis ; see Benda, (2010), Benda and Gaisler (2015), Ruedi et al. (2015) and Ruedi et al. (2021). Distinct from Myotis muricola and Myotis mystacinus ; see Stadelmann et al. (2007) and Ruedi et al. (2015).; [MDD2023] split from Myotis muricola; moved from Myotis to Submyotodon; [MDD2025_2.0] split from Myotis muricola; moved from Myotis to Submyotodon; [batnames2025_1.7] Does not include caliginosus, blanfordi, or moupinensis; see Benda, (2010), Benda and Gaisler (2015), Ruedi et al. (2015) and Ruedi et al. (2021). Distinct from Myotis muricola and Myotis mystacinus; see Stadelmann et al. (2007) and Ruedi et al. (2015).; [MDD2025_2.2] split from Myotis muricola; moved from Myotis to Submyotodon latirostris, oriiThis small species related to Myotinae was considered as a subspecies of more widespread taxa such as Myotis muricola (e.g. Simmons 2005) or M. mystacinus (e.g. Tate 1941). However, global molecular studies (Stadelmann et al. 2007, Lack et al . 2010, Ruedi et al. 2013) showed that this species actually differs from any other known Myotis and appears in a basal position in the radiation of Myotinae (Ruedi et al . 2013). In addition, unique dental and cranial characters indicate that this species actually belongs to a distinct genus that is morphologically closely related to Submyotodon, a fossil Myotinae of Miocene age (Ziegler 2003). To reflect these large differences compared to all other species of Myotis, it is currently classified as S. latirostris (Ruedi et al . 2015) and is endemic to Taiwan. Other recent species belonging to the genus Submyotodon are found in the Himalayan region.
latirostris latirostris - orii latirostris, orii latirostris, orii latirostris latirostris - orii latirostris (Kishida, 1932)|orii (Kuroda, 1935) N/A NA 4C3D87E8FF596AE6FF4F9ED71A3BB6B1 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Vespertilionidae_716.pdf.imf hash://md5/b004ff90fffb6a44fffc96591e00bb32 923 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF5A6AE6FA8B92451A0CBA4E.xml Submyotodon latirostris Vespertilionidae Submyotodon latirostris Kishida 1932 Murin de Kishida @fr | Taiwan-Breitmaulfledermaus @de | Ratonero de Taiwan @es | Broad-muzzled Bat @en | Taiwan Broad-muzzled Myotis @en Myotis latirostris Kishida, 1932 , central Taiwan . Submyotodon was originally described based on fossil material. After comparing molars of the extinct type species, S. petersbuchensis, and S. latirostris (then included under Myotis muricola as a subspecies), S. latirostris and two taxa previously considered subspecies of M. muricola ( caliginosus and moupinensis ) were moved to Submyotodon . The genus issister to the rest of Myotinae , except for Fudiscopus (somewhat tentatively in Myotinae ). Relationships among S. latirostris , S. caliginosus , and S. moupinensis have not yet been thoroughly examined. Monotypic. Taiwan I. Head-body 35-40- 5 mm , tail 31-3-40- 2 mm , ear 9-5-14- 2 mm , hindfoot 5-7- 4 mm , forearm 31- 7-35 mm . The Taiwan Broad-muzzled Bat is similar to the Nepalese Whiskered Myotis ( M. muricola ). Its pelage is long and shaggy. Dorsal hairs are dark slate-brown, with lighter brown tips; venter is also dark brown but with lighter, golden-tipped hairs (particularly long in posterior region). Face is dark and comparatively hairy, with bare flesh-colored skin around eyes. Ears are long and pointed, with distinctive sharp notch near rear edge of concha (unique to Submyotodon relative to Myotis ); tragus is relatively short and bent forward, with spatulated tip and parallel-sided base. Membranes are dark and sparsely covered in hair ventrally and close to body but with short whitish hairs near tibia on uropatagium. Wings are attached to bases of outer toes; uropatagium attaches to calcar that has keeled lobe and extends along one-half the hind edge of uropatagium. Penisis short and slightly club-shaped. Skull is delicate and smooth, with no visible crests; rostrum is elongated, relegating foramen lacrimal to level offirst molars rather than premolars as in Myotis ; zygomatic arches are thin; tooth row has clear diastema between second incisor and canine; premolars are not crowded and all visible in lateral view; there are three upper and lower premolars and well-developed paraconule to P* C! is relatively large and conspicuously taller than P* C, is weaker and onlyslightly taller than P,; lower molars are highly distinctive compared to Myotis (in which nearly all have all myotodont molars with one group of species having all nyctalodont molars) because M, and M, are nyctalodont and M, is uniquely submyotodont. Commonly broadleaf and mixed coniferous forests and urban areas at elevations of 1000-3000 m (marginally down to 200 m possibly only in winter). Taiwan Broad-muzzled Bats are insectivorous. Births are mostly observed in May-June, depending on elevation. Lactating females were caught in April-June and males with enlarged testis in August— March. Taiwan Broad-muzzled Bats are nocturnal and active throughout the year. Maternity colonies occur in buildings, but natural day roosts are still uncertain. No information. Classified as Least Concern on The [UCN Red List. The Taiwan Broad-muzzled Bat is often the most common species of bat caught in forests in its distribution. It does not seem to face any major threats. On following pages: 368. Himalayan Broad-muzzled Bat ( Submyotodon caliginosus ); 369. Moupin Broad-muzzled Bat ( Submyotodon moupinensis ); 370. Northern Myotis ( Myotis septentrionalis ); 371. South-western Myotis ( Myotis auriculus ); 372. Western Small-footed Myotis ( Myotis ciliolabrum ); 373. Dark-nosed Small-footed Myotis ( Myotis melanorhinus ); 374. Eastern Small-footed Myotis ( Myotis leibil); 375. California Myotis ( Myotis californicus ); 376. Little Brown Myotis ( Myotis lucifugus ); 377. Keen's Myotis ( Myotis keenii ); 378. Long-eared Myotis ( Myotis evotis ); 379. Fringed Myotis ( Myotis thysanodes ); 380. Arizona Myotis ( Myotis occultus ); 381. Longlegged Myotis ( Myotis volans ); 382. Flat-headed Myotis ( Myotis planiceps ); 383. Indiana Myotis ( Myotis sodalis ); 384. Cinnamon Myotis ( Myotis fortidens ); 385. Findley's Myotis ( Myotis findley)); 386. Northern Hairy-legged Myotis ( Myotis pilosatibialis); 387. Southern Hairy-legged Myotis ( Myotis keaysi ); 388. Red Myotis ( Myotis ruber ); 389. Riparian Myotis ( Myotis riparius ); 390. Velvety Myotis ( Myotis simus ); 391. Golden Myotis ( Myotis midastactus ); 392. Elegant Myotis ( Myotis elegans ); 393. Fish-eating Myotis ( Myotis vives). Benda (2010a) | Cheng Hsichi et al. (2017) | Francis et al. (2010) | Kruskop & Borisenko (2013) | Lack et al. (2010) | Ruedi & Mayer (2001) | Ruedi, Csorba et al. (2015, 2017¢) | Ruedi, Stadelmann et al. (2013) | Stadelmann et al. (2007) | Wiantoro et al. (2012) | Zhang Zhenzhen et al. (2009) https://zenodo.org/record/6398638/files/figure.png 367. Taiwan Broad-muzzled Bat Submyotodon latirostris French: Murin de Kishida / German: Taiwan-Breitmaulfledermaus / Spanish: Ratonero de Taiwan Other common names: Broad-muzzled Bat , Taiwan Broad-muzzled Myotis Taxonomy. Myotis latirostris Kishida, 1932 , central Taiwan . Submyotodon was originally described based on fossil material. After comparing molars of the extinct type species, S. petersbuchensis, and S. latirostris (then included under Myotis muricola as a subspecies), S. latirostris and two taxa previously considered subspecies of M. muricola ( caliginosus and moupinensis ) were moved to Submyotodon . The genus issister to the rest of Myotinae , except for Fudiscopus (somewhat tentatively in Myotinae ). Relationships among S. latirostris , S. caliginosus , and S. moupinensis have not yet been thoroughly examined. Monotypic. Distribution. Taiwan I. Descriptive notes. Head-body 35-40- 5 mm , tail 31-3-40- 2 mm , ear 9-5-14- 2 mm , hindfoot 5-7- 4 mm , forearm 31- 7-35 mm . The Taiwan Broad-muzzled Bat is similar to the Nepalese Whiskered Myotis ( M. muricola ). Its pelage is long and shaggy. Dorsal hairs are dark slate-brown, with lighter brown tips; venter is also dark brown but with lighter, golden-tipped hairs (particularly long in posterior region). Face is dark and comparatively hairy, with bare flesh-colored skin around eyes. Ears are long and pointed, with distinctive sharp notch near rear edge of concha (unique to Submyotodon relative to Myotis ); tragus is relatively short and bent forward, with spatulated tip and parallel-sided base. Membranes are dark and sparsely covered in hair ventrally and close to body but with short whitish hairs near tibia on uropatagium. Wings are attached to bases of outer toes; uropatagium attaches to calcar that has keeled lobe and extends along one-half the hind edge of uropatagium. Penisis short and slightly club-shaped. Skull is delicate and smooth, with no visible crests; rostrum is elongated, relegating foramen lacrimal to level offirst molars rather than premolars as in Myotis ; zygomatic arches are thin; tooth row has clear diastema between second incisor and canine; premolars are not crowded and all visible in lateral view; there are three upper and lower premolars and well-developed paraconule to P* C! is relatively large and conspicuously taller than P* C, is weaker and onlyslightly taller than P,; lower molars are highly distinctive compared to Myotis (in which nearly all have all myotodont molars with one group of species having all nyctalodont molars) because M, and M, are nyctalodont and M, is uniquely submyotodont. Dental formula for all species of Submyotodon is12/3,C1/1,P 3/3, M 3/3 (x2) = 38 Habitat. Commonly broadleaf and mixed coniferous forests and urban areas at elevations of 1000-3000 m (marginally down to 200 m possibly only in winter). Food and Feeding. Taiwan Broad-muzzled Bats are insectivorous. Breeding. Births are mostly observed in May-June, depending on elevation. Lactating females were caught in April-June and males with enlarged testis in August— March. Activity patterns. Taiwan Broad-muzzled Bats are nocturnal and active throughout the year. Maternity colonies occur in buildings, but natural day roosts are still uncertain. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The [UCN Red List. The Taiwan Broad-muzzled Bat is often the most common species of bat caught in forests in its distribution. It does not seem to face any major threats. On following pages: 368. Himalayan Broad-muzzled Bat ( Submyotodon caliginosus ); 369. Moupin Broad-muzzled Bat ( Submyotodon moupinensis ); 370. Northern Myotis ( Myotis septentrionalis ); 371. South-western Myotis ( Myotis auriculus ); 372. Western Small-footed Myotis ( Myotis ciliolabrum ); 373. Dark-nosed Small-footed Myotis ( Myotis melanorhinus ); 374. Eastern Small-footed Myotis ( Myotis leibil); 375. California Myotis ( Myotis californicus ); 376. Little Brown Myotis ( Myotis lucifugus ); 377. Keen's Myotis ( Myotis keenii ); 378. Long-eared Myotis ( Myotis evotis ); 379. Fringed Myotis ( Myotis thysanodes ); 380. Arizona Myotis ( Myotis occultus ); 381. Longlegged Myotis ( Myotis volans ); 382. Flat-headed Myotis ( Myotis planiceps ); 383. Indiana Myotis ( Myotis sodalis ); 384. Cinnamon Myotis ( Myotis fortidens ); 385. Findley's Myotis ( Myotis findley)); 386. Northern Hairy-legged Myotis ( Myotis pilosatibialis); 387. Southern Hairy-legged Myotis ( Myotis keaysi ); 388. Red Myotis ( Myotis ruber ); 389. Riparian Myotis ( Myotis riparius ); 390. Velvety Myotis ( Myotis simus ); 391. Golden Myotis ( Myotis midastactus ); 392. Elegant Myotis ( Myotis elegans ); 393. Fish-eating Myotis ( Myotis vives). Bibliography. Benda (2010a), Cheng Hsichi et al. (2017), Francis et al. (2010), Kruskop & Borisenko (2013), Lack et al. (2010), Ruedi & Mayer (2001), Ruedi, Csorba et al. (2015, 2017¢), Ruedi, Stadelmann et al. (2013), Stadelmann et al. (2007), Wiantoro et al. (2012), Zhang Zhenzhen et al. (2009). Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Vespertilionidae Submyotodon latirostris Submyotodon latirostris Kishida 1932 1 Lansania 0.2729 Broad-nosed Bat Yes. Taiwan Not listed. Least Concern Does not include caliginosus , blanfordi , or moupinensis ; see Benda, (2010), Benda and Gaisler (2015), Ruedi et al. (2015) and Ruedi et al. (2021). Distinct from Myotis muricola and Myotis mystacinus ; see Stadelmann et al. (2007) and Ruedi et al. (2015). Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Submyotodon latirostris 23 Taiwan Broad-muzzled Bat Broad-muzzled Bat|Taiwan Broad-muzzled Myotis Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA VESPERTILIONIFORMES NA NA VESPERTILIONOIDEA VESPERTILIONIDAE MYOTINAE NA Submyotodon NA latirostris Kishida 1932 1 central Taiwan. latirostris (Kishida, 1932)|orii (Kuroda, 1935) split from Myotis muricola; moved from Myotis to Submyotodon Ruedi, M., Csorba, G., Lin, L. K., & Chou, C. H. (2015). Molecular phylogeny and morphological revision of Myotis bats (Chiroptera: Vespertilionidae) from Taiwan and adjacent China. Zootaxa, 3920(1), 301-342.|Csorba, G., Lin, L.-K., Chou, C.-H., Ruedi, M. 2015. Molecular phylogeny and morphological revision of Myotis bats (Chiroptera: Vespertilionidae) from Taiwan and adjacent China. Zootaxa 3920(2). doi: 10.11646/zootaxa.3920.2.6.|Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology, 101(4), 465-480. Taiwan Asia Palearctic LC 0 0 0 Submyotodon_latirostris 0 unmatched NA 1 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 90000000 Submyotodon latirostris ANIMALIA CHORDATA MAMMALIA CHIROPTERA VESPERTILIONIDAE Submyotodon latirostris Kishida, 1932This small species related to Myotinae was considered as a subspecies of more widespread taxa such as Myotis muricola (e.g. Simmons 2005) or M. mystacinus (e.g. Tate 1941). However, global molecular studies (Stadelmann et al. 2007, Lack et al . 2010, Ruedi et al. 2013) showed that this species actually differs from any other known Myotis and appears in a basal position in the radiation of Myotinae (Ruedi et al . 2013). In addition, unique dental and cranial characters indicate that this species actually belongs to a distinct genus that is morphologically closely related to Submyotodon, a fossil Myotinae of Miocene age (Ziegler 2003). To reflect these large differences compared to all other species of Myotis, it is currently classified as S. latirostris (Ruedi et al . 2015) and is endemic to Taiwan. Other recent species belonging to the genus Submyotodon are found in the Himalayan region.
90000000 Submyotodon latirostris Least Concern 2017 2016-06-13 00:00:00 UTC 3.1 EnglishAlthough this species occurs only in a single island and therefore has a restricted range, it is believed to be stable, widespread and not currently facing any specific threat. ;
This forest bat is associated to a number of different habitats found in mountain areas of Taiwan, where it feeds on small insects (Cheng, Fang and Chou 2015). It has been found in broadleaf as well as in mixed coniferous forests. Breeding colonies have been found in buildings, but natural roosts are still unknown. It is often the most common species of bat caught in forests. Parturition occurs mostly in May and June, depending on elevation.This widespread species is apparently adapted to a variety of forested habitats in Taiwan and may not be particularly under threat. ;
Commonly recorded in forest habitats, but no population estimate has been made.
Unknown Widespread and common mostly at mid- to higher elevations throughout Taiwan. Most records were made above 1000 m Asl, but it can occur marginally at lower elevation (down to about 200 m Asl). Terrestrial Present in many well protected areas such as national parks or national forests. Indomalayan FALSE FALSE Global Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Vespertilionidae Submyotodon latirostris Kishida 1932 1 Lansania 4(40): 153 Broad-nosed Bat Yes. Taiwan Not listed. Least Concern Does not include caliginosus , blanfordi , or moupinensis ; see Benda, (2010), Benda and Gaisler (2015), Ruedi et al. (2015) and Ruedi et al. (2021). Distinct from Myotis muricola and Myotis mystacinus ; see Stadelmann et al. (2007) and Ruedi et al. (2015). Submyotodon latirostris 1005490 23 Taiwan Broad-muzzled Bat Broad-muzzled Bat|Taiwan Broad-muzzled Myotis Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA VESPERTILIONIFORMES NA NA VESPERTILIONOIDEA Vespertilionidae MYOTINAE NA Submyotodon NA latirostris Kishida 1932 1 central Taiwan. latirostris (Kishida, 1932)|orii (Kuroda, 1935) split from Myotis muricola; moved from Myotis to Submyotodon Ruedi, M., Csorba, G., Lin, L. K., & Chou, C. H. (2015). Molecular phylogeny and morphological revision of Myotis bats (Chiroptera: Vespertilionidae) from Taiwan and adjacent China. Zootaxa, 3920(1), 301-342.|Csorba, G., Lin, L.-K., Chou, C.-H., Ruedi, M. 2015. Molecular phylogeny and morphological revision of Myotis bats (Chiroptera: Vespertilionidae) from Taiwan and adjacent China. Zootaxa 3920(2). doi: 10.11646/zootaxa.3920.2.6.|Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology, 101(4), 465-480. Taiwan Asia Palearctic LC 0 0 0 Submyotodon_latirostris 0 unmatched NA 1 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Submyotodon_latirostris 1005490 23 Taiwan Broad-muzzled Bat Broad-muzzled Bat|Taiwan Broad-muzzled Myotis Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yangochiroptera NA NA Vespertilionoidea Vespertilionidae Myotinae NA Submyotodon NA latirostris Kishida 1 Myotis latirostris Kishida, K. 1932-11-11. Notes on a new Formosan whiskered bat. Lansania 4(40):153-160. untraced (number not known) holotype central Taiwan. split from Myotis muricola; moved from Myotis to Submyotodon Ruedi, M., Csorba, G., Lin, L. K., & Chou, C. H. (2015). Molecular phylogeny and morphological revision of Myotis bats (Chiroptera: Vespertilionidae) from Taiwan and adjacent China. Zootaxa, 3920(1), 301-342.|Csorba, G., Lin, L.-K., Chou, C.-H., Ruedi, M. 2015. Molecular phylogeny and morphological revision of Myotis bats (Chiroptera: Vespertilionidae) from Taiwan and adjacent China. Zootaxa 3920(2). doi: 10.11646/zootaxa.3920.2.6.|Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology, 101(4), 465-480. Taiwan Asia Palearctic LC 0 0 0 Submyotodon_latirostris 0 unmatched NA 1 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Vespertilionidae Submyotodon latirostris Kishida 1932 1 Lansania 4(40): 153 Broad-nosed Bat Yes. Taiwan Not Listed Least Concern Does not include caliginosus, blanfordi, or moupinensis; see Benda, (2010), Benda and Gaisler (2015), Ruedi et al. (2015) and Ruedi et al. (2021). Distinct from Myotis muricola and Myotis mystacinus; see Stadelmann et al. (2007) and Ruedi et al. (2015). Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Submyotodon latirostris; Submyotodon latirostris; Submyotodon latirostris; Submyotodon latirostris; Submyotodon latirostris; latirostris; orii; Murin de Kishida; Taiwan-Breitmaulfledermaus; Ratonero de Taiwan; Broad-muzzled Bat; Taiwan Broad-muzzled Myotis; Taiwan Broad-muzzled Bat; Broad-muzzled Bat; Taiwan Broad-muzzled Myotis; Broad-nosed Bat; S. latirostris