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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1586	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei	Styloctenium wallacei		[MSW3] See Bergmans and Rozendaal (1988).; [HMW] Pteropus wallacei J. E. Gray, 1866 , “ Celebes [= Sulawesi , Indonesia ], Macassar.” Styloctenium is sister to other living flying foxes in genera Acerodon , Neopteryx , and Pteropus . Monotypic.; [batnames2022] See Bergmans and Rozendaal (1988).; [batnames2023] See Bergmans and Rozendaal (1988).; [batnames2025_1.7] See Bergmans and Rozendaal (1988).														wallacei				wallacei	wallacei			wallacei (J. E. Gray, 1866)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Stripe-faced fruit bat	Celebes	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Styloctenium wallacei	Indonesia, Sulawesi, Macassar.	Gray	1866	Proc. Zool. Soc. Lond., 1866:65.	Distribution: Same as genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Striped-faced fruit bat	Sulawesi	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Gray	1866	Proc. Zool. Soc. Lond., 1866:65.		Sulawesi.	Indonesia, Sulawesi, Macassar.		GRAY	1866	Size relatively small (forearm length, 90-96 mm).	Distribution: Same as genus.	No subspecies.		28	species	S. wallacei	GRAY	1866	Styloctenium	genus	Styloctenium wallacei				Size relatively small (forearm length, 90-96 mm).	No subspecies.		1. S. wallacei (GRAY 1866).	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Styloctenium wallacei	Styloctenium		wallacei	Gray	y	1866		Proc. Zool. Soc. Lond.	1866		65		Stripe-faced Fruit Bat	Indonesia, Sulawesi, Macassar.	Sulawesi, Tongian Isls.	IUCN/SSC Action Plan (1992) – No Data. IUCN 2003 – Lower Risk (nt).		See Bergmans and Rozendaal (1988).	4C3D87E8FF596AE6FF4A979C1C83B377	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	133	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF8FF661896A35BDFCB8F701.xml	Styloctenium wallacei	Pteropodidae	Styloctenium	wallacei		1866	Roussette de Wallace @fr | Sulawesi-Streifengesichtflughund @de | Styloctenio de Wallace @es | Sulawesi Stripe-faced Fruit Bat @en	Pteropus wallacei J. E. Gray, 1866 , “ Celebes [= Sulawesi , Indonesia ], Macassar.” Styloctenium is sister to other living flying foxes in genera Acerodon , Neopteryx , and Pteropus . Monotypic.	Sulawesi , including nearby Togian, Buton, and Selayar Is.	Head-body 145- 2— 200 mm (umilless), ear 17-24- 8 mm , hindfoot 27- 2-32 mm , forearm 90-103- 2 mm ; weight 174-218 g . Greatest lengths of skulls are 50-54- 3 mm . Hair of Wallace's Stripe-faced Fruit Bat is soft and silky. Face and crown are reddish brown to ocherous buff, with 11-105 mm nasal white stripe, one spot above cach eye, and white mustache. Eyes are brown. Ears are light brown, naked except at bases, and attenuated at tips. Throat, chest, and belly have light reddish-brown fur. Occiput, nape, shoulders, and back are usually gray, often with hint of reddish brown. Dorsal forearm is grayish white, with white humeral spot. Wings have patterns of translucent bands and dots, with bands generally parallel to digits. Wing membranesattach to sides of back and insert on second toe. Index claw is present, and all claws are black. Rostrum is more gracile, and teeth are smaller than in the Mindoro Stripe-faced Fruit Bat (S. mindorensis ). Posterior edge of palate is more deeply cut, exposing much of sphenoid crest. Postorbital processes are strong and elongated. Dental formula for both species of Styloctenium 1s12/1, C 1/1, P 3/3, M 2/2 (x2) = 30. Incisors are cylindrical, slender, and pointed at tops. Canines are short and thick. In contrast to the Mindoro Stripefaced Fruit Bat, canines of Wallace's Stripe-faced Fruit Bat are not that complex. Cheekteeth have rounded cusps, I, and M? are absent, and P,is only rudimentary.	Primary forests and slightly disturbed forests with clearings in understories for coffee and cocoa plantations from sea level up to elevations of ¢. 1100 m .	Wallace’s Stripe-faced Fruit Bats probably eat fruits and flower products.	No information.	Wallace's Stripe-faced Fruit Bat is probably crepuscular or nocturnal.	Wallace's Stripe-faced Fruit Bats have been captured crossing gaps between fragmented forests.	Classified as Near Threatened on The IUCN Red List. Populations of Wallace's Stripe-faced Fruit Bat are decreasing due to decline of primary forests, and it is hunted in some areas. It is generally rare, but there are recent records from Lore Lindu National Park and Lambusango Forest Reserve on Buton Island.	Almeida et al. (2014) | Andersen (1912b) | Bergmans (2001) | Bergmans & Rozendaal (1988) | Esselstyn (2007) | Gray (1866a) | Heinsohn (2002) | Jentink (1883) | Matschie (1899) | Nowak (1999) | Patterson etal. (2017) | Ruedas et al. (2010) | Simmons (2005) | Yusti et al. (2015)		127. Wallace's Stripe-faced Fruit Bat Styloctenium wallacei French: Roussette de Wallace / German: Sulawesi-Streifengesichtflughund / Spanish: Styloctenio de Wallace Other common names: Sulawesi Stripe-faced Fruit Bat Taxonomy. Pteropus wallacei J. E. Gray, 1866 , “ Celebes [= Sulawesi , Indonesia ], Macassar.” Styloctenium is sister to other living flying foxes in genera Acerodon , Neopteryx , and Pteropus . Monotypic. Distribution. Sulawesi , including nearby Togian, Buton, and Selayar Is. Descriptive notes. Head-body 145- 2— 200 mm (umilless), ear 17-24- 8 mm , hindfoot 27- 2-32 mm , forearm 90-103- 2 mm ; weight 174-218 g . Greatest lengths of skulls are 50-54- 3 mm . Hair of Wallace's Stripe-faced Fruit Bat is soft and silky. Face and crown are reddish brown to ocherous buff, with 11-105 mm nasal white stripe, one spot above cach eye, and white mustache. Eyes are brown. Ears are light brown, naked except at bases, and attenuated at tips. Throat, chest, and belly have light reddish-brown fur. Occiput, nape, shoulders, and back are usually gray, often with hint of reddish brown. Dorsal forearm is grayish white, with white humeral spot. Wings have patterns of translucent bands and dots, with bands generally parallel to digits. Wing membranesattach to sides of back and insert on second toe. Index claw is present, and all claws are black. Rostrum is more gracile, and teeth are smaller than in the Mindoro Stripe-faced Fruit Bat (S. mindorensis ). Posterior edge of palate is more deeply cut, exposing much of sphenoid crest. Postorbital processes are strong and elongated. Dental formula for both species of Styloctenium 1s12/1, C 1/1, P 3/3, M 2/2 (x2) = 30. Incisors are cylindrical, slender, and pointed at tops. Canines are short and thick. In contrast to the Mindoro Stripefaced Fruit Bat, canines of Wallace's Stripe-faced Fruit Bat are not that complex. Cheekteeth have rounded cusps, I, and M? are absent, and P,is only rudimentary. Habitat. Primary forests and slightly disturbed forests with clearings in understories for coffee and cocoa plantations from sea level up to elevations of ¢. 1100 m . Food and Feeding. Wallace’s Stripe-faced Fruit Bats probably eat fruits and flower products. Breeding. No information. Activity patterns. Wallace's Stripe-faced Fruit Bat is probably crepuscular or nocturnal. Movements, Home range and Social organization. Wallace's Stripe-faced Fruit Bats have been captured crossing gaps between fragmented forests. Status and Conservation. Classified as Near Threatened on The IUCN Red List. Populations of Wallace's Stripe-faced Fruit Bat are decreasing due to decline of primary forests, and it is hunted in some areas. It is generally rare, but there are recent records from Lore Lindu National Park and Lambusango Forest Reserve on Buton Island. Bibliography. Almeida et al. (2014), Andersen (1912b), Bergmans (2001), Bergmans & Rozendaal (1988), Esselstyn (2007), Gray (1866a), Heinsohn (2002), Jentink (1883), Matschie (1899), Nowak (1999), Patterson etal. (2017), Ruedas et al. (2010), Simmons (2005), Yusti et al. (2015).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Styloctenium wallacei	Styloctenium		wallacei	Gray	1866	1	Proc. Zool. Soc. Lond.	1867:05:00	Stripe-faced Fruit Bat	None.	Indonesia, Sulawesi, Macassar.	Sulawesi, Togian Isls.	Not listed.	Near Threatened	See Bergmans and Rozendaal (1988).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Styloctenium wallacei	23	Wallace's Stripe-faced Fruit Bat	Sulawesi Stripe-faced Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTEROPODINI	Styloctenium	NA	wallacei	J. E. Gray	1866	1	Pteropus_wallacei	Gray, J. E. (1866). A revision of the genera of pteropine bats (Pteropidae), and the descriptions of some apparently undescribed species. Proceedings of the Zoological Society of London, 1866, 65.	https://www.biodiversitylibrary.org/item/90954#page/99/mode/1up	BM 1866.2.13.11		"Celebes [= Sulawesi, Indonesia], Macassar."			wallacei (J. E. Gray, 1866)	NA	NA	Indonesia	Asia	Australasia/Oceania	NT	0	0	0	Styloctenium_wallacei	0	sciname match	Styloctenium_wallacei	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	21100	Styloctenium wallacei	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Styloctenium	wallacei	(Gray, 1866)		200000000	Styloctenium wallacei	Near Threatened	A2cd	2021	2021-06-13 00:00:00 UTC	3.1	English	Styloctenium wallacei is assessed as Near Threatened (nearly meets the threshold for criterion A). This species is suspected to have declined over the past 15 years (three generations, GL = 5 years, Pacifici et al. 2013) at a rate of 25–29% due to widespread forest loss and hunting throughout much of its range on Sulawesi and adjacent islands.	Styloctenium wallacei is most closely associated with tropical rainforests up to 1,800 m asl and has also been encountered in mixed-gardens (e.g. cocoa, rambutan, and durian plantation) (Maryanto et al. 2011, Yusti 2015, Widayati and Nurjana 2018, Widayati et al. 2020). This species is a solitary, nectarivorous species that also feeds on fruits (Maryanto et al. 2011).	Degradation and the loss of forest habitats due to logging and small-scale agriculture are the primary drivers of the species decline. Further, it is at risk of bushmeat hunting for local consumption in North Sulawesi close to Bogani Nani Wartabone National Park (Mickleburgh et al. 2009, Ransaleleh et al. 2020).	<span id="docs-internal-guid-0b8e208b-7fff-cb41-5ed7-be88b45b407d"><p>Population data on Styloctenium wallacei are not available as the species is usually encountered in very small numbers (1–2 individuals), which is often much lower compared to other Pteropodid bats in Sulawesi. The species population is suspected to be declining by an estimated 25–29% due to forest loss and hunting (Mickleburgh et al. 2009, Ransaleleh et al. 2020). It is unknown if the species' distribution reflects a single location or if it comprises a single population.</p>	Decreasing	This species has been recorded from throughout Sulawesi, including the nearby Buton Island and the Togian Islands (Bergmans and Rozendaal 1988, Maryanto et al. 2011, Yusti 2015, Patterson et al. 2017, Mubarok et al. 2018). The species is suspected to occur on Selayar Island (Heinsohn 2002), but that has yet to be confirmed. Styloctenium wallacei has been found from sea level up to 1,800 m asl (Maryanto et al. 2011, Widayati and Nurjana 2018, Widayati et al. 2020). The type locality for this species is Macassar, Sulawesi, Indonesia.	The species is threatened by hunting for bushmeat for local consumption in North Sulawesi close to Bogani Nani Wartabone National Park (Mickleburgh et al. 2009, Ransaleleh et al. 2020).	Terrestrial	While S. wallacei occurs in Lore Lindu National Park (Maryanto et al. 2011), Lambusango Forest Reserve (Patterson et al. 2017), and Bogani Nani Wartabone National Park (Ransaleleh et al. 2020), there are no known species-specific conservation initiatives in place for it. Forests where the species is known from should be protected, and degraded forests should be restored. Additional research is needed to inform the species distribution, its population status and trends, and the extent of the threats, including the impacts of hunting on the species. Insights into the species ecology are needed to better inform the species conservation and management.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Styloctenium		wallacei	Gray	1866	1	Proc. Zool. Soc. Lond.	1867:05:00	Stripe-faced Fruit Bat	None.	Indonesia, Sulawesi, Macassar.	Sulawesi, Togian Isls.	Not listed.	Near Threatened	See Bergmans and Rozendaal (1988).	Styloctenium wallacei	1004512	23	Wallace's Stripe-faced Fruit Bat	Sulawesi Stripe-faced Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTEROPODINI	Styloctenium	NA	wallacei	J. E. Gray	1866	1	Pteropus_wallacei	Gray, J. E. (1866). A revision of the genera of pteropine bats (Pteropidae), and the descriptions of some apparently undescribed species. Proceedings of the Zoological Society of London, 1866, 65.	https://www.biodiversitylibrary.org/item/90954#page/99/mode/1up	BM 1866.2.13.11		"Celebes [= Sulawesi, Indonesia], Macassar."			wallacei (J. E. Gray, 1866)	NA	NA				Indonesia	Asia	Australasia/Oceania	NT	0	0	0	Styloctenium_wallacei	0	sciname match	Styloctenium_wallacei	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Styloctenium_wallacei	1004512	23	Wallace's Stripe-faced Fruit Bat	Sulawesi Stripe-faced Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteropodini	Styloctenium	NA	wallacei	J. E. Gray	1	Pteropus wallacei	Gray, J.E. 1866-05. A revision of the genera of pteropine bats (Pteropidee), and the descriptions of some apparently undescribed species. Proceedings of the Zoological Society of London 1866(1):62-67.	https://www.biodiversitylibrary.org/page/28627501	BMNH:Mamm:1866.2.13.11	holotype	https://data.nhm.ac.uk/object/f057a1d2-cb1a-4176-9846-c18545fea8cc	"Celebes [= Sulawesi, Indonesia], Macassar."			NA	NA				Indonesia	Asia	Australasia	NT	0	0	0	Styloctenium_wallacei	0	sciname match	Styloctenium_wallacei	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Styloctenium		wallacei	Gray	1866	1	Proc. Zool. Soc. Lond.	1867:05:00	Stripe-faced Fruit Bat	None.	Indonesia, Sulawesi, Macassar.	Sulawesi, Togian Isls.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/21100/203829571/' target='_blank'>Near Threatened</a>	See Bergmans and Rozendaal (1988).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Styloctenium wallacei; Styloctenium wallacei; Styloctenium wallacei; Styloctenium wallacei; Styloctenium wallacei; Styloctenium wallacei; wallacei; Roussette de Wallace; Sulawesi-Streifengesichtflughund; Styloctenio de Wallace; Sulawesi Stripe-faced Fruit Bat; Wallace's Stripe-faced Fruit Bat; Sulawesi Stripe-faced Fruit Bat; Stripe-faced Fruit Bat; Stripe-faced Fruit Bat; S. wallacei