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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1582	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Sturnira perla	Sturnira perla	Sturnira perla	Sturnira perla	Sturnira perla	Sturnira perla	Sturnira perla	Sturnira perla	Sturnira perla		[HMW] Sturnira perla Jarrin-Valladares & Kunz, 2011 , “ Ecuador , Provincia Santo Domingo de los Tsachilas , Bosque Protector La Perla at 200 meters of altitude ( 0°08’N , 79°30°W ), 2 km south of La Concordia.” This species is monotypic.; [MDD2022] recently described; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														perla				perla	perla			perla JarrÃ­n-Valladares & Kunz, 2011						N/A																																								NA																											03A687BCFF8FFF8F1384FCE9FC77F85A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	546	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF8FFF8E1690F3D4FCE2F46C.xml	Sturnira perla	Phyllostomidae	Sturnira	perla	Jarrin-Valladares & Kunz	2011	Sturnire de La Perla @fr | Perla-Gelbschulterfledermaus @de | Sturniro de La Perla @es | La Perla Forest Yellow-shouldered Bat @en	Sturnira perla Jarrin-Valladares & Kunz, 2011 , “ Ecuador , Provincia Santo Domingo de los Tsachilas , Bosque Protector La Perla at 200 meters of altitude ( 0°08’N , 79°30°W ), 2 km south of La Concordia.” This species is monotypic.	Endemic to NW Ecuador ; its presence in SW Colombia is expected.	Head-body 56-66 mm (tailless), ear 14-17 mm, forearm 41-45 mm; weight 18 g . The Perla Yellowshouldered Bat is medium-sized, externally similar to Luis’s Yellow-shouldered Bat (S. wisi) and the Northern Yellowshouldered Bat ( S. parvidens ). Dorsal fur is grayish brown with four-banded hairs; sometimes shoulders and neck of adults (especially males) have yellow or reddish patch; ventral fur is paler, with tricolored hairs. The Perla Yellow-shouldered Bat is characterized by a spherical skull, noticeably round and robust, with pronounced curved zygomatic arches. Lower molars have poorly defined cusps toward tongue.	[.owlands and adjacent slopes in primary and secondary humid tropical forests on western side of the Andes at elevations of 35-700 m.	There is no specific information available for this species, but yellow-shouldered bats feed on fruits, being largely dependent on Solanum ( Solanaceae ) and Piper ( Piperaceae ) on the lowlands. The distinct shape of the skull of the Perla Yellow-shouldered Bat suggests that it might also be divergent in its feeding behavior.	No information.	The Perla Yellow-shouldered Bat is nocturnal.	No information.	Classified as Data Deficient on The IUCN Red List. The Perla Yellow-shouldered Bat is rare and poorly known, with a restricted distribution. There is no information available on population size, and therefore its conservation status is unknown. Deforestation and mining are probably its main threats. Records came from two protected areas: Cotacachi-Cayapas Ecological Reserve and La Perla Protected Forest. Nevertheless, La Perla is an isolated patch of Ecuadorean Choco forest consisting of only 226 ha of primary forest surrounded by vast expanses of monocultures consisting mostly of grazing lands, oil palms, and banana trees.	Jarrin-Valladares & Kunz (2011) | Tirira (2017)		131. Perla Yellow-shouldered Bat Sturnira perla French: Sturnire de La Perla / German: Perla-Gelbschulterfledermaus / Spanish: Sturniro de La Perla Other common names: La Perla Forest Yellow-shouldered Bat Taxonomy. Sturnira perla Jarrin-Valladares & Kunz, 2011 , “ Ecuador , Provincia Santo Domingo de los Tsachilas , Bosque Protector La Perla at 200 meters of altitude ( 0°08’N , 79°30°W ), 2 km south of La Concordia.” This species is monotypic. Distribution. Endemic to NW Ecuador ; its presence in SW Colombia is expected. Descriptive notes. Head-body 56-66 mm (tailless), ear 14-17 mm, forearm 41-45 mm; weight 18 g . The Perla Yellowshouldered Bat is medium-sized, externally similar to Luis’s Yellow-shouldered Bat (S. wisi) and the Northern Yellowshouldered Bat ( S. parvidens ). Dorsal fur is grayish brown with four-banded hairs; sometimes shoulders and neck of adults (especially males) have yellow or reddish patch; ventral fur is paler, with tricolored hairs. The Perla Yellow-shouldered Bat is characterized by a spherical skull, noticeably round and robust, with pronounced curved zygomatic arches. Lower molars have poorly defined cusps toward tongue. Habitat. [.owlands and adjacent slopes in primary and secondary humid tropical forests on western side of the Andes at elevations of 35-700 m. Food and Feeding. There is no specific information available for this species, but yellow-shouldered bats feed on fruits, being largely dependent on Solanum ( Solanaceae ) and Piper ( Piperaceae ) on the lowlands. The distinct shape of the skull of the Perla Yellow-shouldered Bat suggests that it might also be divergent in its feeding behavior. Breeding. No information. Activity patterns. The Perla Yellow-shouldered Bat is nocturnal. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Perla Yellow-shouldered Bat is rare and poorly known, with a restricted distribution. There is no information available on population size, and therefore its conservation status is unknown. Deforestation and mining are probably its main threats. Records came from two protected areas: Cotacachi-Cayapas Ecological Reserve and La Perla Protected Forest. Nevertheless, La Perla is an isolated patch of Ecuadorean Choco forest consisting of only 226 ha of primary forest surrounded by vast expanses of monocultures consisting mostly of grazing lands, oil palms, and banana trees. Bibliography. Jarrin-Valladares & Kunz (2011), Tirira (2017).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Sturnira perla	Sturnira		perla	Jarrin-V & Kunz	2011	0	Zootaxa	######	La Perla Forest Yellow-shouldered Bat	None.	Ecaudor, Provincia Santo Domingo de los Tsachilas, Bosque Protector La Perla	NW Ecuador; may extend into NW Colombia	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Sturnira perla	23	Perla Yellow-shouldered Bat	La Perla Forest Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STURNIRINI	Sturnira	Sturnira	perla	JarrÃ­n-Valladares & Kunz	2011	0	Sturnira_perla	JarrÃ­n-V. & Kunz, T. H. (2011). A new species of Sturnira (Chiroptera: Phyllostomidae) from the Choco forest of Ecuador. Zootaxa, 2755, 6.	https://www.mapress.com/zootaxa/2011/2/zt02755p035.pdf	QCAZ 120		"Ecuador, Provincia Santo Domingo de los Tsachilas, Bosque Protector La Perla at 200 meters of altitude (0Â°08'N, 79Â°30'W), 2 km south of La Concordia."	0.1333	-79.5	perla JarrÃ­n-Valladares & Kunz, 2011	recently described	Roca, Q. (2011). A new species of Sturnira (Chiroptera: Phyllostomidae) from the Choco forest of Ecuador. Zootaxa, 2755, 1-35.	Ecuador|Colombia?	South America	Neotropic	DD	0	0	0	Sturnira_perla	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Sturnira perla	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Sturnira	perla	Jarrin &; Kunz, 2011		90000000	Sturnira perla	Data Deficient		2016	2016-07-15 00:00:00 UTC	3.1	English	This species is listed as Data Deficient as it has only recently been described and its ecological requirements are poorly known. The current known distribution is limited, although given our current lack of major data on its ecology, the true distribution may be more restricted or may be broader. The dominant habitat has been extensively affected by deforestation and fragmentation within its presumed range. Indeed, more surveys are required.	Seven of the fourteen specimens known for S. perla were collected from an isolated patch of Ecuadorean ChocÃ³ forest consisting of 226 ha of primary forest, known as â€œBosque Protector la Perlaâ€, and surrounded by vast expanses of monocultures consisting mostly of grazing lands, oil palms and banana trees (Jarrin and Kunz 2011). These authors hypothesize that the distinct shape of the skull in this species, relative to the highly overlapped distribution that other lowland species have in morphometric space, suggests it ;may also be remarkably divergent in its ecological habits, especially those immediately related to feeding behaviour.	All 14 specimens known for this species come from a highly devastated area, suffering from an accelerated rate of deforestation, amidst a human population affected from increasingly levels of social unrest and poverty (Jarrin and Kunz 2011).	The most recent collection records for Sturnira perla are from over a decade ago (Jarrin and Kunz 2011). This species seems to be uncommon through its limited distribution area.	Unknown	This species is known only from the tropical rainforest lowlands of the ChocÃ³ forests in Ecuador. Its presence may also extend to the Colombian ChocÃ³ (Jarrin and Kunz 2011).		Terrestrial	Half of the known specimens come from a private protected reserve isolated from larger extensions of natural ChocÃ³ forests (Jarrin and Kunz 2011).	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Sturnira		perla	Jarrin-V & Kunz	2011	0	Zootaxa	########	La Perla Forest Yellow-shouldered Bat	None.	Ecaudor, Provincia Santo Domingo de los Tsachilas, Bosque Protector La Perla	NW Ecuador; may extend into NW Colombia	Not listed.	Data Deficient		Sturnira perla	1005091	23	Perla Yellow-shouldered Bat	La Perla Forest Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STURNIRINI	Sturnira	Sturnira	perla	JarrÃ­n-Valladares & Kunz	2011	0	Sturnira_perla	JarrÃ­n-V. & Kunz, T. H. (2011). A new species of Sturnira (Chiroptera: Phyllostomidae) from the Choco forest of Ecuador. Zootaxa, 2755, 6.	https://www.mapress.com/zootaxa/2011/2/zt02755p035.pdf	QCAZ 120		"Ecuador, Provincia Santo Domingo de los Tsachilas, Bosque Protector La Perla at 200 meters of altitude (0Â°08'N, 79Â°30'W), 2 km south of La Concordia."	0.133333	-79.5	perla JarrÃ­n-Valladares & Kunz, 2011	recently described	Roca, Q. (2011). A new species of Sturnira (Chiroptera: Phyllostomidae) from the Choco forest of Ecuador. Zootaxa, 2755, 1-35.				Ecuador|Colombia?	South America	Neotropic	DD	0	0	0	Sturnira_perla	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Sturnira_perla	1005091	23	Perla Yellow-shouldered Bat	La Perla Forest Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Sturnirini	Sturnira	Sturnira	perla	JarrÃ­n-Valladares & Kunz	0	Sturnira perla	JarrÃ­n-Valladares, P. and Kunz, T.H. 2011-02-02. A new species of _Sturnira_ (Chiroptera: Phyllostomidae) from the Choco forest of Ecuador. Zootaxa 2755:1-35.	https://doi.org/10.11646/zootaxa.2755.1.1	QCAZ 120	holotype		"Ecuador, Provincia Santo Domingo de los Tsachilas, Bosque Protector La Perla at 200 meters of altitude (0Â°08'N, 79Â°30'W), 2 km south of La Concordia."	0.133333	-79.5	recently described	Roca, Q. (2011). A new species of Sturnira (Chiroptera: Phyllostomidae) from the Choco forest of Ecuador. Zootaxa, 2755, 1-35.				Ecuador|Colombia?	South America	Neotropic	DD	0	0	0	Sturnira_perla	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Sturnira		perla	Jarrin-V & Kunz	2011	0	Zootaxa	########	La Perla Forest Yellow-shouldered Bat	None.	Ecaudor, Provincia Santo Domingo de los Tsachilas, Bosque Protector La Perla	NW Ecuador; may extend into NW Colombia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/88159664/88159667/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Sturnira perla; Sturnira perla; Sturnira perla; Sturnira perla; Sturnira perla; perla; Sturnire de La Perla; Perla-Gelbschulterfledermaus; Sturniro de La Perla; La Perla Forest Yellow-shouldered Bat; Perla Yellow-shouldered Bat; La Perla Forest Yellow-shouldered Bat; La Perla Forest Yellow-shouldered Bat; S. perla
