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authorityParentheses originalNameCombination authoritySpeciesCitation authoritySpeciesLink typeVoucher typeKind typeVoucherURIs typeLocality typeLocalityLatitude typeLocalityLongitude taxonomyNotes taxonomyNotesCitation distributionNotes distributionNotesCitation subregionDistribution countryDistribution continentDistribution biogeographicRealm iucnStatus extinct domestic flagged CMW_sciName diffSinceCMW MSW3_matchtype MSW3_sciName diffSinceMSW3 docOrigin_batnames2025 Family Genus Subgenus Species Author Date Parentheses (1=author & date in parentheses) Citation Pages Common Name Synonyms Type Locality Distribution CITES IUCN Comments column3781 column3791 subtribe CONCAT_ALTNAMES line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L159 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali Cloeotis percivali [HMW] Cloeotis pemivali Thomas, 1901 , “Takaungu, N. of Mombasa , British East Africa Kenya ]. Subspecies often noted, with nominate perrivalz are smaller in body size a than a subspecies australıs and conﬁned to Kenya (and perhaps Tanzania ) vs. southern AF rica. Validity of these two subspecies remains open to question, and no revision based on well representative material is available. Monotypic.; [MDD2022] moved from Hipposideridae to Rhinonycteridae; [IUCN] Meester et al. (1986) listed two subspecies, of which only Cloeotis p . australis occurs in the assessment region, while the other, C . p . percivali occurs in East Africa.; [MDD2023] moved from Hipposideridae to Rhinonycteridae; [MDD2025_2.0] moved from Hipposideridae to Rhinonycteridae; [MDD2025_2.2] moved from Hipposideridae to Rhinonycteridae australis. percivali, australis percivali , australis australis percivali, australis Meester et al. (1986) listed two subspecies, of which only Cloeotis p . australis occurs in the assessment region, while the other, C . p . percivali occurs in East Africa. australis, percivali percivali, australis percivali, australis australis, percivali percivali O. Thomas, 1901|australis A. Roberts, 1917 Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp. Short-eared trident bat Kenya – SE Botswana – Mozambique Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp. Cloeotis percivali Kenya, Coast Prov., Takaungu. Thomas 1901 Ann. Mag. Nat. Hist., ser. 7, 8:28. Distribution: Same as for genus. Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5 Short-eared trident bat (Percival's trident bat) Kenya – SE Botswana – Mozambique, Transvaal, Swaziland Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp. Thomas 1901 Ann. Mag. Nat. Hist., ser. 7, 8:28. Kenya, Tanzania, S Zaire, Mozambique, Zambia, Zimbabwe, SE Botswana, Swaziland, Transvaal (South Africa). Kenya, Coast Prov., Takaungu. THOMAS 1901 Size small (forearm length, 30-39 mm). Distribution: Same as for genus. Two subspecies: C. p. percivali (southeastern Kenya, northeastern Tan zania), C.p. australis (southern Zaire to Botswana and Swaziland). 68 species C. percivali THOMAS 1901 Cloeotis genus Cloeotis percivali Size small (forearm length, 30-39 mm). Two subspecies: 1. C. percivali THOMAS 1901. 1 _C. p. australis_ Roberts, 1917; _C. p. percivali_ Thomas, 1901 Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu). CHIROPTERA Hipposideridae Cloeotis percivali Cloeotis percivali Thomas 1901 Ann. Mag. Nat. Hist., ser. 7 8 28 Percival's Short-eared Trident Bat Kenya, Coast Prov., Takaungu. Kenya, Tanzania, S Dem. Rep. Congo, Mozambique, Zambia, Zimbabwe, SE Botswana, Swaziland, NE South Africa. IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt). australis Roberts, 1917. 0383245F2226977A8B0EFE1EF738FDF3 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Rhinonycteridae_194.pdf.imf hash://md5/ffba5c272223977c8e73f509ffb1ffd0 207 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/83/24/0383245F2226977A8B0EFE1EF738FDF3.xml Cloeotis percivali Rhinonycteridae Cloeotis percivali Thomas 1901 é he de Per va German @fr | AtrcanTidenr Bat AfrcanTrdent nosed Bat asrAtrcan Trdent Bat Por iva s Short-cared @en Cloeotis pemivali Thomas, 1901 , “Takaungu, N. of Mombasa , British East Africa Kenya ]. Subspecies often noted, with nominate perrivalz are smaller in body size a than a subspecies australıs and conﬁned to Kenya (and perhaps Tanzania ) vs. southern AF rica. Validity of these two subspecies remains open to question, and no revision based on well representative material is available. Monotypic. SE Kenya , Maﬁa I ( Tanzania ), SE DR Congo , Zambia , Mozambique , Zimbabue , SE Botswana , NE South Africa , and Swaziland . Head-body 30-43 mm, tail 18-36 mm, ear 7-10 mm, hindfoot 5-8 mm, forearm 31-39 mm; weight 3-6 g. Percival's Trident Bat is the smallest species in the family. Pelage usually has slightly darker tinge dorsally than ventrally, 0r venter is sometimes very pale or whitish; color very variable, including bright brownish orange, pale yellowish, pale brown, pale or dark grayish brown, and dull gray; hairs are dark brown, brownish gray, or gray around eyes, noseleaf, and mouth. Wing membranes arc dark grayish brown, and ears and noseleaf are pale pinkish (unpigrnented) to pale grayish brown. Noseleaf is small (width 3 -4-3-9 mm) and rounded, with straplike longitudinal projection that is diamond-shaped in outline and anteriorly forked and lying across anterior part of anterior leaf, above deep anterior medial emargination. Three tall pointed posterior projections are present on posterior pan of posterior leaf, and anterior part in medial position has short and narrow process, with sharply pointed tip. Total width of three posterior projections is c. 50% the width of posterior noseleaf; lateral projections have no emargination at their bases. lateral pans of posterior noseleaf are scalloped by six small cells on each side, plus pair of large cells in central position, separated from each other by ﬂeshy septum. Two supplementary leaflets occur lateral to noseleaf. Ears are very small, rounded, and only bluntly pointed; pinna membrane is thickened. Skull has posteriorly but not laterally prominent nasal swellings. Braincase is much higher than narrow rostrum, and minute sagittal crest is developed only in frontal region. Zygomatic bones are anteriorly strongly convergent and thin, with low dorsal triangular projection on posterior margin. Greatest skull lengths are 13-136 mm, condvlo-canine lengths are 10-9-11 -3 mm, zygomatic widths are 7-7-7 mm, and upper tooth row (C —M‘) lengths are 3 -8-4-4 mm. f is bilobed; C' is slender, with slight cingulum and large posterior secondary cusp, extending about one-third the canine crown height; P‘ is small and extruded; C‘ and P‘ are in or nearly in contact; and M‘ is only little reduced, with metacone. Lower incisors are tricuspid, C, is slender, P, is about one-third to threequarters the crown area and one-half to twothirds the height of P4, and M, is unreduced. Baculum has not been described. Diploid number is 2n 40, but number of autosome arms is unknown. Dry woodland savannas with mopane (Colophospennum mopane, Fabaceae ) and various miombo woodlands, bushvelds, riverine woodlands, coastal forest mosaic, and others habitats from sea level to elevations of c. l 540 m . Percival`s Trident Bats are aerial hawkers, preying on small ﬂying insects at various heights aboveground and mostly among vegetation. Individuals were documented foraging at various water bodies, in riparian vegetation, and among woodland vegetation. Stomach contents were almost entirely small-sized adult moths (97-1 % by overall volume). All stomachs contained moths, and 86% of them contained only moths. Termites (1 %), beetles 1 -1 %), ﬂies (0-6 %), bugs (04 %). and other items were also consumed. There was little variation in prey type consumed during the night or during annual cycles (moths were 94-7% in summer diets and 983% in winter); small-sized moths were clearly preferred over other similarly sized insects that were equally available. Pregnant Percival's Trident Bats, each containing one fetus, were found in October in Zimbabwe , and pregnant and lactating females with attached young were found in late November in north-eastem South Africa and early December in southern Zambia . This indicated that births occur in November and early December. Examinations of females in other seasons did not reveal any signs of ongoing reproductive activity. Hence, Percival's Trident Bat is, like other species of the family, probably seasonallv monoestrous, with females giving birth to one young each year. Percival’s Trident Bat probably does not enter torpor or hibernate. probably an acrobatic ﬂier. leaves day roosts shortly after sunset and forages for several hours or the entire night. Percival's Trident Bat roosts in underground spaces in natural caves and human-made structures (e.g. mine tunnels and dam body corridors). lt roosts in small and narrow crevices and were once found in a cavity on the ground that was thought to be a porcupine den. Echolocation consists of multiharmonic QCF calls where pulse is composed of short CF component, combined with short FM component at end (CF-FM); tenninating FM component is steep sweep up to 29 kHz. Call is unique due to its unusually high peak frequency of ﬁrst harmonic, well above 200 kHz; however, fundamental frequency often used during foraging. Call has mean durations of 1 -9 milliseconds (fundamental frequency) and 4 -6 milliseconds (ﬁrst harmonic). Peak frequencies (fundamental frequencies) are 101-6-105 -3 kHz (mean 103 -4 kHz), ﬁrst hannonics are 204-212 kHz (mean 207 -8 kHz). highest frequency is 212 kHz, and end (lowest) frequency is 183 kHz. Slight sexual dimorphism was detected in CF component of fundamental harmonic frequency: females ha\e, on average, lower frequency (102-9 kHz) than males (103-7 kHz). Extremely high frequency of echolocation calls is speculated to be an adaptation to more efficiently catch moths. Remains of Percivafs Trident Bats were found in small amounts of the diet of the bat hawk (Madmmmphus alnnus). Roosting Percival’s Trident Bats hang on cavity ceilings in loose groups, with indiriduals separated from each other by 10-20 cm. Roosting groups have 10-300 individuals, but there are no data on their sexual composition. ln Zambia , a colony of Percival's Trident Bat shared its roost (large karst cave) with colonies of Sundevalfs Leaf-nosed Bats (Hıpposidems mfƒrr), Su-iped Leaf-nosed Bats (Mammycleris vittalus), and Natal bong-ﬁngered Bats (Mmaopterus natalmsis). https://zenodo.org/record/6611836/files/figure.png 6. Percival`s Trident Bat Cloeotis percivali French: é he de Per va German: Pei va Kenohr-Drezahnbartna e Spanish: Rn meter dePercrva Other common names: AtrcanTidenr Bat AfrcanTrdent nosed Bat asrAtrcanTrdent Bat Por iva s Short-cared Bat Per va Sh ﬂ-GBIOGTIIÖGM Bat Sh n-earedTııdent Bat Taxonomy. Cloeotis pemivali Thomas, 1901 , “Takaungu, N. of Mombasa , British East Africa Kenya ]. Subspecies often noted, with nominate perrivalz are smaller in body size a than a subspecies australıs and conﬁned to Kenya (and perhaps Tanzania ) vs. southern AF rica. Validity of these two subspecies remains open to question, and no revision based on well representative material is available. Monotypic. Distribution. SE Kenya , Maﬁa I ( Tanzania ), SE DR Congo , Zambia , Mozambique , Zimbabue , SE Botswana , NE South Africa , and Swaziland . Descriptive notes. Head-body 30-43 mm, tail 18-36 mm, ear 7-10 mm, hindfoot 5-8 mm, forearm 31-39 mm; weight 3-6 g. Percival's Trident Bat is the smallest species in the family. Pelage usually has slightly darker tinge dorsally than ventrally, 0r venter is sometimes very pale or whitish; color very variable, including bright brownish orange, pale yellowish, pale brown, pale or dark grayish brown, and dull gray; hairs are dark brown, brownish gray, or gray around eyes, noseleaf, and mouth. Wing membranes arc dark grayish brown, and ears and noseleaf are pale pinkish (unpigrnented) to pale grayish brown. Noseleaf is small (width 3 -4-3-9 mm) and rounded, with straplike longitudinal projection that is diamond-shaped in outline and anteriorly forked and lying across anterior part of anterior leaf, above deep anterior medial emargination. Three tall pointed posterior projections are present on posterior pan of posterior leaf, and anterior part in medial position has short and narrow process, with sharply pointed tip. Total width of three posterior projections is c. 50% the width of posterior noseleaf; lateral projections have no emargination at their bases. lateral pans of posterior noseleaf are scalloped by six small cells on each side, plus pair of large cells in central position, separated from each other by ﬂeshy septum. Two supplementary leaflets occur lateral to noseleaf. Ears are very small, rounded, and only bluntly pointed; pinna membrane is thickened. Skull has posteriorly but not laterally prominent nasal swellings. Braincase is much higher than narrow rostrum, and minute sagittal crest is developed only in frontal region. Zygomatic bones are anteriorly strongly convergent and thin, with low dorsal triangular projection on posterior margin. Greatest skull lengths are 13-136 mm, condvlo-canine lengths are 10-9-11 -3 mm, zygomatic widths are 7-7-7 mm, and upper tooth row (C —M‘) lengths are 3 -8-4-4 mm. f is bilobed; C' is slender, with slight cingulum and large posterior secondary cusp, extending about one-third the canine crown height; P‘ is small and extruded; C‘ and P‘ are in or nearly in contact; and M‘ is only little reduced, with metacone. Lower incisors are tricuspid, C, is slender, P, is about one-third to threequarters the crown area and one-half to twothirds the height of P4, and M, is unreduced. Baculum has not been described. Diploid number is 2n 40, but number of autosome arms is unknown. Habitat. Dry woodland savannas with mopane (Colophospennum mopane, Fabaceae ) and various miombo woodlands, bushvelds, riverine woodlands, coastal forest mosaic, and others habitats from sea level to elevations of c. l 540 m . Food and Feeding. Percival`s Trident Bats are aerial hawkers, preying on small ﬂying insects at various heights aboveground and mostly among vegetation. Individuals were documented foraging at various water bodies, in riparian vegetation, and among woodland vegetation. Stomach contents were almost entirely small-sized adult moths (97-1 % by overall volume). All stomachs contained moths, and 86% of them contained only moths. Termites (1 %), beetles 1 -1 %), ﬂies (0-6 %), bugs (04 %). and other items were also consumed. There was little variation in prey type consumed during the night or during annual cycles (moths were 94-7% in summer diets and 983% in winter); small-sized moths were clearly preferred over other similarly sized insects that were equally available. Breeding. Pregnant Percival's Trident Bats, each containing one fetus, were found in October in Zimbabwe , and pregnant and lactating females with attached young were found in late November in north-eastem South Africa and early December in southern Zambia . This indicated that births occur in November and early December. Examinations of females in other seasons did not reveal any signs of ongoing reproductive activity. Hence, Percival's Trident Bat is, like other species of the family, probably seasonallv monoestrous, with females giving birth to one young each year. Activity patterns. Percival’s Trident Bat probably does not enter torpor or hibernate. probably an acrobatic ﬂier. leaves day roosts shortly after sunset and forages for several hours or the entire night. Percival's Trident Bat roosts in underground spaces in natural caves and human-made structures (e.g. mine tunnels and dam body corridors). lt roosts in small and narrow crevices and were once found in a cavity on the ground that was thought to be a porcupine den. Echolocation consists of multiharmonic QCF calls where pulse is composed of short CF component, combined with short FM component at end (CF-FM); tenninating FM component is steep sweep up to 29 kHz. Call is unique due to its unusually high peak frequency of ﬁrst harmonic, well above 200 kHz; however, fundamental frequency often used during foraging. Call has mean durations of 1 -9 milliseconds (fundamental frequency) and 4 -6 milliseconds (ﬁrst harmonic). Peak frequencies (fundamental frequencies) are 101-6-105 -3 kHz (mean 103 -4 kHz), ﬁrst hannonics are 204-212 kHz (mean 207 -8 kHz). highest frequency is 212 kHz, and end (lowest) frequency is 183 kHz. Slight sexual dimorphism was detected in CF component of fundamental harmonic frequency: females ha\e, on average, lower frequency (102-9 kHz) than males (103-7 kHz). Extremely high frequency of echolocation calls is speculated to be an adaptation to more efficiently catch moths. Remains of Percivafs Trident Bats were found in small amounts of the diet of the bat hawk (Madmmmphus alnnus). Movements, Home range and Social organization. Roosting Percival’s Trident Bats hang on cavity ceilings in loose groups, with indiriduals separated from each other by 10-20 cm. Roosting groups have 10-300 individuals, but there are no data on their sexual composition. ln Zambia , a colony of Percival's Trident Bat shared its roost (large karst cave) with colonies of Sundevalfs Leaf-nosed Bats (Hıpposidems mfƒrr), Su-iped Leaf-nosed Bats (Mammycleris vittalus), and Natal bong-ﬁngered Bats (Mmaopterus natalmsis). Status and Conservation. Classiﬁed as beast Ooncem on The IUCN Rad List. Percivafs Trident Bat reportedly highly sensitive to roost disturbance. In South Africa , religious ceremonies by local people are carried out in caves, and these rituals and other forms of disturbance can impact local populations and perhaps cause abandonment of roosts. ln Zambia , Percival's Trident Bat shared a cave with Striped Leaf-nosed Bats that were harvested by locals for food, which seriously disturb roosting Percival's Trident Bats. Due to observed ﬂuctuations in numbers of roosting bats at particular roosts, such activities could cause local extirpations. ln South Africa , a national law protects Percival's Trident Bat. Bibliogaphy. Baona (2016) Back (1979), Bat* et a (1979) HI (1982a). Jacobs (2013) Monadiam. Res de 8r Lumsden (2007) Monad |em, Tay or era (2010) Reutenbach eta (1993). Sdwoeman G« Jacobs (2008). Seamark (2005), Taylor (1999), Thomas (1901 b), Whitaker 8r B ack (1976) Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Rhinonycteridae Cloeotis percivali Cloeotis percivali Thomas 1901 0 Ann. Mag. Nat. Hist. ser. 7, 8: 28 Percival's Short-eared Trident Bat australis Roberts, 1917. Kenya, Coast Prov., Takaungu. Kenya, Tanzania, S Dem. Rep. Congo, Mozambique, Zambia, Zimbabwe, SE Botswana, Swaziland, NE South Africa. Not listed. Least Concern Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Cloeotis percivali 23 Percival's Trident Bat African Trident-nosed Bat|East African Trident Bat|Percival's Short-eared Bat|Percival's Short-eared Trident Bat|Short-eared Trident Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA RHINONYCTERIDAE NA NA Cloeotis NA percivali O. Thomas 1901 0 Cloeotis_Percivali Thomas, O. (1901). Some new African bats (including one from the Azores) and a new Galago. Annals and Magazine of Natural History, ser. 7, 8, 28. https://www.biodiversitylibrary.org/item/94923#page/48/mode/1up BM 1901.5.1.11 "Takaungu, N. of Mombasa, British East Africa [= Kenya]." percivali O. Thomas, 1901|australis Roberts, 1917 moved from Hipposideridae to Rhinonycteridae Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333. Kenya|Tanzania|Democratic Republic of the Congo|Zambia|Mozambique|Zimbabwe|Botswana|South Africa|Eswatini Africa Afrotropic LC 0 0 0 Cloeotis_percivali 0 sciname match Cloeotis_percivali 0 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 4983 Cloeotis percivali ANIMALIA CHORDATA MAMMALIA CHIROPTERA HIPPOSIDERIDAE Cloeotis percivali Thomas, 1901 Meester et al. (1986) listed two subspecies, of which only Cloeotis p . australis occurs in the assessment region, while the other, C . p . percivali occurs in East Africa. 20000000 Cloeotis percivali Least Concern 2017 2016-08-31 00:00:00 UTC 3.1 English Although poorly known and with some recorded population declines, it is listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category. It presumably occurs in savanna areas where there is sufficient cover in the form of caves and mine tunnels for day roosting (Taylor 2000). It feeds exclusively on moths, and appears to be very sensitive to disturbance. There have been seemingly stochastic extinctions of colonies, though it is not known if the animals simply move to new locations. Roost disturbance seems to be an important factors, but there might be others, and there has been speculation that DDT might be a cause of population disappearances. Large fluctuations in population numbers are known, and it is prone to local extinctions. It is never found in very large colonies, with the largest known colony (of about 300 individuals) being reported in KwaZulu-Natal, South Africa, on the southern section of its range. However, the current estimated population size of the KwaZulu-Natal colony is currently only 50 animals. There are no population estimates from elsewhere in its range, although the largest population might be in Zimbabwe. Unknown Percival's trident bat is largely confined to southern Africa, with records from South Africa (KwaZulu-Natal), Swaziland, south-east Botswana, southern Zambia, Zimbabwe (the core of the distribution), and extralimital records from southern Democratic Republic of Congo, Malawi, northwestern Mozambique, and coastal Kenya (Taylor 2000). It has also been found on Mafia Island in Tanzania (unpublished record). It elevational range is from sea level to 1,000 m. Terrestrial In South Africa, KwaZulu-Natal province they are protected from human disturbance. It might occur in some protected areas. Protection of nesting and roosting sites from disturbance is important. Afrotropical FALSE FALSE Global Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Rhinonycteridae Cloeotis percivali Thomas 1901 0 Ann. Mag. Nat. Hist. ser. 7, 8: 28 Percival's Short-eared Trident Bat australis Roberts, 1917. Kenya, Coast Prov., Takaungu. Kenya, Tanzania, S Dem. Rep. Congo, Mozambique, Zambia, Zimbabwe, SE Botswana, Swaziland, NE South Africa. Not listed. Least Concern Cloeotis percivali 1004761 23 Percival's Trident Bat African Trident-nosed Bat|East African Trident Bat|Percival's Short-eared Bat|Percival's Short-eared Trident Bat|Short-eared Trident Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA Rhinonycteridae NA NA Cloeotis NA percivali O. Thomas 1901 0 Cloeotis_Percivali Thomas, O. (1901). Some new African bats (including one from the Azores) and a new Galago. Annals and Magazine of Natural History, ser. 7, 8, 28. https://www.biodiversitylibrary.org/item/94923#page/48/mode/1up BM 1901.5.1.11 "Takaungu, N. of Mombasa, British East Africa [= Kenya]." percivali O. Thomas, 1901|australis Roberts, 1917 moved from Hipposideridae to Rhinonycteridae Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333. Kenya|Tanzania|Democratic Republic of the Congo|Zambia|Mozambique|Zimbabwe|Botswana|South Africa|Eswatini Africa Afrotropic LC 0 0 0 Cloeotis_percivali 0 sciname match Cloeotis_percivali 0 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Cloeotis_percivali 1004761 23 Percival's Trident Bat African Trident-nosed Bat|East African Trident Bat|Percival's Short-eared Bat|Percival's Short-eared Trident Bat|Short-eared Trident Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yinpterochiroptera NA NA Rhinolophoidea Rhinonycteridae NA NA Cloeotis NA percivali O. Thomas 0 Cloeotis Percivali Thomas, O. 1901-07-01. Some new African bats (including one from the Azores) and a new _Galago_. Annals and Magazine of Natural History (7)8(43):27-34. https://www.biodiversitylibrary.org/page/29980030 BMNH:Mamm:1901.5.1.11 holotype https://data.nhm.ac.uk/object/5b5b7039-c837-4b1f-9955-efe3e93481dd "Takaungu, N. of Mombasa, British East Africa [= Kenya]." moved from Hipposideridae to Rhinonycteridae Foley, N. M., Thong, V. D., Soisook, P., Goodman, S. M., Armstrong, K. N., Jacobs, D. S., ... & Teeling, E. C. (2015). How and why overcome the impediments to resolution: lessons from rhinolophid and hipposiderid bats. Molecular Biology and Evolution, 32(2), 313-333. Kenya|Tanzania|Democratic Republic of the Congo|Zambia|Mozambique|Zimbabwe|Botswana|South Africa|Eswatini Africa Afrotropic LC 0 0 0 Cloeotis_percivali 0 sciname match Cloeotis_percivali 0 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Rhinonycteridae Cloeotis percivali Thomas 1901 0 Ann. Mag. Nat. Hist. ser. 7, 8: 28 Percival's Short-eared Trident Bat australis Roberts, 1917. Kenya, Coast Prov., Takaungu. Kenya, Tanzania, S Dem. Rep. Congo, Mozambique, Zambia, Zimbabwe, SE Botswana, Swaziland, NE South Africa. Not Listed Least Concern Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Cloeotis percivali; Cloeotis percivali; Cloeotis percivali; Cloeotis percivali; Cloeotis percivali; Cloeotis percivali; percivali ; australis; australis; percivali; australis; é he de Per va German; AtrcanTidenr Bat AfrcanTrdent nosed Bat asrAtrcan Trdent Bat Por iva s Short-cared; Percival's Trident Bat; African Trident-nosed Bat; East African Trident Bat; Percival's Short-eared Bat; Percival's Short-eared Trident Bat; Short-eared Trident Bat; Percival's Short-eared Trident Bat; Percival's Short-eared Trident Bat; C. percivali