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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1578	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana	Sturnira nana		[MSW2] Subgenus Corvira.; [MSW3] Subgenus Corvira.; [HMW] Sturnira nana Gardner & O’Neill, 1971 , “Huanhuachayo ( 12°44’S , 73°47'W ), ca. 1,660 meters, Departamento de Ayacucho , Pera.” Sturnira nana was originally associated with S. bidens in the subgenus Corvira, but phylogenetic analyses shown they are not close relatives. Monotypic.; [batnames2022] Subgenus Corvira .; [batnames2023] Subgenus Corvira .; [batnames2025_1.7] Subgenus Corvira.														nana				nana	nana			nana A. L. Gardner & J. P. O'Neill, 1971		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		C Peru	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Sturnira nana	Peru, Ayacucho, Huanhuachayo, 1660 m.	Gardner and O'Neill	1971	Occas. Pap. Mus. Zool. La. St. Univ., 42:1.	Distribution: Known only from a small area in the mountains of southern Peru.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		S Peru	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Gardner and O'Neill	1971	Occas. Pap. Mus. Zool. La. St. Univ., 42:1.	Subgenus Corvira.	S Peru.	Peru, Ayacucho, Huanhuachayo, 1,660 m.		GARDNER & O'NEILL	1971	Size very small (forearm length, 34-36 mm; condylobasal length, 16-18 mm). Vestigial outer lower incisors frequently present.	Distribution: Known only from a small area in the mountains of southern Peru.	No subspecies.		86	species	S. nana	GARDNER & O'NEILL	1971	Corvira	subgenus	Sturnira nana				Size very small (forearm length, 34-36 mm; condylobasal length, 16-18 mm). Vestigial outer lower incisors frequently present.	No subspecies.		11. S. nana GARDNER & O'NEILL 1971.	11	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Stenodermatinae	Sturnirini	Sturnira nana	Sturnira	Corvira	nana	Gardner and O'Neill		1971		Occas. Pap. Mus. Zool. La. St. Univ.	42		1		Lesser Yellow-shouldered Bat	Peru, Ayacucho, Huanhuachayo, 1,660 m.	S Peru.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Subgenus Corvira.	03A687BCFF8CFF8C1640FDA2F710FB71	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	543	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF8AFF8A1342F527F85DF432.xml	Sturnira nana	Phyllostomidae	Sturnira	nana	Gardner & O'Neill	1971	Sturnire naine @fr | Kleine Gelbschulterfledermaus @de | Sturniroenano @es	Sturnira nana Gardner & O’Neill, 1971 , “Huanhuachayo ( 12°44’S , 73°47'W ), ca. 1,660 meters, Departamento de Ayacucho , Pera.” Sturnira nana was originally associated with S. bidens in the subgenus Corvira, but phylogenetic analyses shown they are not close relatives. Monotypic.	Known only from SE Ecuador (Cordillera del Condor, Zamora Chinchipe Province ) and type locality in SC Peru .	Head—body 47-50 mm (tailless), ear 13-15 mm, hindfoot 9-15 mm, forearm 32-6-35-7 mm; weight 10-12 g. The Lesser Yellow-shouldered Batis the smallest species of Sturnira . Upperparts are dark grayish brown. Shoulder glands are absent. Dorsal hairs are long (7-8 mm), with 3—4 bands: almost white basal band, brown medial band, then light silvery brown band, and brownish narrow tips. Underparts are slightly lighter because hairs lack brownish tips. Noseleaf is long and narrow. Forearm, legs, feet, adjacent part of the wings, and short tail membrane are sparsely haired. Skull is relatively long, narrow, and anteriorly sloping, with weak or incomplete zygomatic arches. There are two pairs of lower incisors; I, are broad and trilobed; I, are minute (or absent) and hardly distinct in live individuals; I' have broader tips compared with the Bidentate Yellow-shouldered Bat ( S. bidens ), and they are in contactat tips; and small molars and premolars are separated by narrow gaps. Dental formula for all species of Sturnira except the Bidentate Yellow-shouldered Batis12/1-2,C1/1,P 2/2, M 3/3 ( x2 ) = 30-32. Chromosomal complement has 2n = 30 and FN = 46, with 14 biarmed autosomal chromosomes. X-chromosome is subtelocentric, and Y-chromosome is acrocentric.	Only known from three Andean localities in Peru and Ecuador within a narrow elevational band of 1250-1670 m. Lesser Yellow-shouldered Bats were netted in low montane forests and at edges of clearings on eastern slope of the Andes.	Diet of the Lesser Yellow-shouldered Bat is presumably similar to other small species of Sturnira , especially those from montane forests. It might depend on fruits of Solanum ( Solanaceae ) but also include Vismia ( Hypericaceae ), Ficus ( Moraceae ), Piper ( Piperaceae ), and Anthurium (Araceae) species.	No information.	Lesser Yellow-shouldered Bats are nocturnal.	No information.	Classified as Endangered on The IUCN Red List. Known area of occupancy of the Lesser Yellow-shouldered Bat is less than 500 km ? at two locations (south-eastern Ecuador and central Peru ). This small area is severely fragmented and threatened by habitat conversion (small-scale agriculture and wood logging).	Boada (2011) | Gardner (1977a, 2008g) | Gardner & O'Neill (1971) | Pacheco & Patterson (1991) | Regalado & Albuja (2012) | Velazco & Patterson (2013)	https://zenodo.org/record/6458849/files/figure.png	120. Lesser Yellow-shouldered Bat Sturnira nana French: Sturnire naine / German: Kleine Gelbschulterfledermaus / Spanish: Sturniro enano Taxonomy. Sturnira nana Gardner & O’Neill, 1971 , “Huanhuachayo ( 12°44’S , 73°47'W ), ca. 1,660 meters, Departamento de Ayacucho , Pera.” Sturnira nana was originally associated with S. bidens in the subgenus Corvira, but phylogenetic analyses shown they are not close relatives. Monotypic. Distribution. Known only from SE Ecuador (Cordillera del Condor, Zamora Chinchipe Province ) and type locality in SC Peru . Descriptive notes. Head—body 47-50 mm (tailless), ear 13-15 mm, hindfoot 9-15 mm, forearm 32-6-35-7 mm; weight 10-12 g. The Lesser Yellow-shouldered Batis the smallest species of Sturnira . Upperparts are dark grayish brown. Shoulder glands are absent. Dorsal hairs are long (7-8 mm), with 3—4 bands: almost white basal band, brown medial band, then light silvery brown band, and brownish narrow tips. Underparts are slightly lighter because hairs lack brownish tips. Noseleaf is long and narrow. Forearm, legs, feet, adjacent part of the wings, and short tail membrane are sparsely haired. Skull is relatively long, narrow, and anteriorly sloping, with weak or incomplete zygomatic arches. There are two pairs of lower incisors; I, are broad and trilobed; I, are minute (or absent) and hardly distinct in live individuals; I' have broader tips compared with the Bidentate Yellow-shouldered Bat ( S. bidens ), and they are in contactat tips; and small molars and premolars are separated by narrow gaps. Dental formula for all species of Sturnira except the Bidentate Yellow-shouldered Batis12/1-2,C1/1,P 2/2, M 3/3 ( x2 ) = 30-32. Chromosomal complement has 2n = 30 and FN = 46, with 14 biarmed autosomal chromosomes. X-chromosome is subtelocentric, and Y-chromosome is acrocentric. Habitat. Only known from three Andean localities in Peru and Ecuador within a narrow elevational band of 1250-1670 m. Lesser Yellow-shouldered Bats were netted in low montane forests and at edges of clearings on eastern slope of the Andes. Food and Feeding. Diet of the Lesser Yellow-shouldered Bat is presumably similar to other small species of Sturnira , especially those from montane forests. It might depend on fruits of Solanum ( Solanaceae ) but also include Vismia ( Hypericaceae ), Ficus ( Moraceae ), Piper ( Piperaceae ), and Anthurium (Araceae) species. Breeding. No information. Activity patterns. Lesser Yellow-shouldered Bats are nocturnal. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Endangered on The IUCN Red List. Known area of occupancy of the Lesser Yellow-shouldered Bat is less than 500 km ? at two locations (south-eastern Ecuador and central Peru ). This small area is severely fragmented and threatened by habitat conversion (small-scale agriculture and wood logging). Bibliography. Boada (2011), Gardner (1977a, 2008g ), Gardner & O'Neill (1971), Pacheco & Patterson (1991), Regalado & Albuja (2012), Velazco & Patterson (2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Sturnira nana	Sturnira	Corvira	nana	Gardner & O'Neill	1971	0	Occas. Pap. Mus. Zool. La. St. Univ.	######	Lesser Yellow-shouldered Bat	None.	Peru, Ayacucho, Huanhuachayo, 1,660 m.	S Peru.	Not listed.	Endangered	Subgenus Corvira .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Sturnira nana	23	Lesser Yellow-shouldered Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STURNIRINI	Sturnira	Corvira	nana	A. L. Gardner & O'Neill	1971	0	Sturnira_nana	Gardner, A. L. & O'Neill, J. P. (1971). A new species of Sturnira (Chiroptera: Phyllostomidae) from Peru. Occasional Papers of the Museum of Zoology Louisiana State University, 42, 1.	https://sites01.lsu.edu/wp/mnspapers/files/2014/09/42.pdf	LSUMZ 15683		"Huanhuachayo (12Â°44'S, 73Â°47'W), ca. 1,660 meters, Departamento de Ayacucho, PerÃº."	-12.73	-73.78	nana A. L. Gardner & O'Neill, 1971	NA	NA	Ecuador|Peru	South America	Neotropic	EN	0	0	0	Sturnira_nana	0	sciname match	Sturnira_nana	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	20958	Sturnira nana	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Sturnira	nana	Gardner &; O'Neill, 1971		20000000	Sturnira nana	Endangered	B2ab(iii)	2016	2015-10-22 00:00:00 UTC	3.1	English	Listed as Endangered because its area of occupancy is &lt;500 kmÂ², known only from 2 locations. This area of occupancy is threatened by severe habitat conversion (small scale agriculture, wood logging). Even with new records in southern Ecuador, its distribution is still severely fragmented, and there is continuing decline in the extent and quality of its major habitat (moist montane forests).	Little is known of the behavior of this species, but it is probably similar to other members of the genus. Specimens were netted in lower montane forest and at the edges of clearings on the eastern slope of the Andes (Gardner 2008).	Areas in southern Peru are threatened by illicit crops and other agriculture.	It is rare and its distribution restricted (Gardner 2008, Regalado and Albuja 2012). Nine specimens were caught at a single locality in southern Ecuador (Boada 2011).	Unknown	This species occurs in southern Peru (Simmons 2005), but there are two additional localities from southern Ecuador (Boada 2011, Regalado and Albuja 2012).		Terrestrial	Drug enforcement may help to reduce drastic changes in forest cover. A binational protected area has been proposed for the Cordillera del Condor, the border between Ecuador and Peru.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Sturnira	Corvira	nana	Gardner & O'Neill	1971	0	Occas. Pap. Mus. Zool. La. St. Univ.	42:01:00	Lesser Yellow-shouldered Bat	None.	Peru, Ayacucho, Huanhuachayo, 1,660 m.	S Peru.	Not listed.	Endangered	Subgenus Corvira .	Sturnira nana	1005087	23	Lesser Yellow-shouldered Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STURNIRINI	Sturnira	Corvira	nana	A. L. Gardner & O'Neill	1971	0	Sturnira_nana	Gardner, A. L. & O'Neill, J. P. (1971). A new species of Sturnira (Chiroptera: Phyllostomidae) from Peru. Occasional Papers of the Museum of Zoology Louisiana State University, 42, 1.	https://sites01.lsu.edu/wp/mnspapers/files/2014/09/42.pdf	LSUMZ 15683		"Huanhuachayo (12Â°44'S, 73Â°47'W), ca. 1,660 meters, Departamento de Ayacucho, PerÃº."	-12.7333	-73.7833	nana A. L. Gardner & O'Neill, 1971	NA	NA				Ecuador|Peru	South America	Neotropic	EN	0	0	0	Sturnira_nana	0	sciname match	Sturnira_nana	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Sturnira_nana	1005087	23	Lesser Yellow-shouldered Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Sturnirini	Sturnira	Corvira	nana	A. L. Gardner & J. P. O'Neill	0	Sturnira nana	Gardner, A.L. and O'Neill, J.P. 1971-06-30. A new species of _Sturnira_ (Chiroptera: Phyllostomidae) from Peru. Occasional Papers of the Museum of Zoology, Louisiana State University 42:1-7.	https://doi.org/10.31390/opmns.042	LSUMZ:Mamm:15683	holotype	http://portal.vertnet.org/o/lsumz/mammals?id=urn-catalog-lsumz-mammals-15683	"Huanhuachayo (12Â°44'S, 73Â°47'W), ca. 1,660 meters, Departamento de Ayacucho, PerÃº."	-12.7333	-73.7833	NA	NA				Peru	South America	Neotropic	EN	0	0	0	Sturnira_nana	0	sciname match	Sturnira_nana	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Sturnira	Corvira	nana	Gardner & O'Neill	1971	0	Occas. Pap. Mus. Zool. La. St. Univ.	42:01:00	Lesser Yellow-shouldered Bat	None.	Peru, Ayacucho, Huanhuachayo, 1,660 m.	S Peru.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/20958/22050195/' target='_blank'>Endangered</a>	Subgenus Corvira.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Sturnira nana; Sturnira nana; Sturnira nana; Sturnira nana; Sturnira nana; Sturnira nana; nana; Sturnire naine; Kleine Gelbschulterfledermaus; Sturniroenano; Lesser Yellow-shouldered Bat; Lesser Yellow-shouldered Bat; Lesser Yellow-shouldered Bat; S. nana
