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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1567	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Sturnira burtonlimi	Sturnira burtonlimi	Sturnira burtonlimi	Sturnira burtonlimi	Sturnira burtonlimi	Sturnira burtonlimi	Sturnira burtonlimi	Sturnira burtonlimi	Sturnira burtonlimi		[HMW] Sturnira burtonlimi Velazco & Patterson, 2014 , “Ojo de Agua, 2 km N of Santa Clara, Chiriqui , Panama , approximately 8°42’N , 82°45’W , 1500m .” Sturnira burtonlimi was previously confused with S. ludovici and S. hondurensis . Molecular and morphological analyses identified it as a distinct species. Monotypic.; [MDD2022] recently described; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														burtonlimi				burtonlimi	burtonlimi			burtonlimi Velazco & B. D. Patterson, 2014						N/A																																								NA																											03A687BCFF8EFF8E16B8FEB4F809F905	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	544	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF8DFF8D1388F432F6F2F46C.xml	Sturnira burtonlimi	Phyllostomidae	Sturnira	burtonlimi	Velazco & Patterson	2014	Sturnire de Burton Lim @fr | Burton-Gelbschulterfledermaus @de | Sturniro de Burton Lim @es | Burton Lim's Yellow-shouldered Bat @en	Sturnira burtonlimi Velazco & Patterson, 2014 , “Ojo de Agua, 2 km N of Santa Clara, Chiriqui , Panama , approximately 8°42’N , 82°45’W , 1500m .” Sturnira burtonlimi was previously confused with S. ludovici and S. hondurensis . Molecular and morphological analyses identified it as a distinct species. Monotypic.	Known only from two localities, one in C Costa Rica ( Cartago Province ) and the other in W Panama (type locality).	Head—body 70-72 mm (tailless), ear 15 mm , hindfoot 14 mm , forearm 44 mm ; weight 19 g (all measurements from two specimens). Burton's Yellow-shouldered Bat is medium-sized, similar to the Honduran Yellow-shouldered Bat ( S. hondurensis ). Dorsal fur is dark brown. Dorsal hairs have four bands, with short pale gray base (c.10% of the length of each hair), long dark gray band (c.40%), long pale gray band (c.40%), and short dark brown tip (c.10%). Ventral fur is dark brown and tricolored, with narrow pale gray base, long dark brown subterminal band (c.45% of each hair), and long gray terminal band (c.45%). Furis long and woolly: 7 mm between shoulders and 5 mm on chest. Proximal part of forearm is densely furred with short hairs. Wing membranes of are dark brown. Dorsal surfaces of femur and tibia are densely covered with long hairs; dorsal surfaces of feet are densely covered with short hairs. Tail membrane is very short and well-furred with long hairs. I' is procumbent and atleast twice the height of I, and I, and I, are bicuspidate and subequalin size. Lingual cuspids (metaconid and entoconid) of M, and M, are poorly defined and are not separated by deep notch.	Collected in pre-montane forest near coffee fields at elevations of 1290-1500 m. Based on E. Dinerstein’s 1986 ecological study on the Costa Rican Highland Yellow-shouldered Bat ( S. ludovici ) that could have actually been Burton’s Yellow-shouldered Bat, fruits of abundant second-growth shrubs are more frequently eaten than mature forest plants.	Dinerstein stated that Burton’s Yellow-shouldered Bat consumes fruits of 27 plant species, including Solanum spp. ( Solanaceae ), Piperspp., and P. umbellatum ( Piperaceae ).	Dinerstein stated that Burton’s Yellow-shouldered Bat has birth peaks in April-May and September—October.	Burton's Yellow-shouldered Bat is nocturnal.	No information.	Classified as Data Deficient on The [UCN Red List. Although Burton’s Yellow-shouldered Bat has a restricted distribution, there are no major conservation threats in this area.	Dinerstein (1986) | Reid (2009) | Velazco & Patterson (2013, 2014)	https://zenodo.org/record/6458855/files/figure.png	123. Burton's Yellow-shouldered Bat Sturnira burtonlimi French: Sturnire de Burton Lim / German: Burton-Gelbschulterfledermaus / Spanish: Sturniro de Burton Lim Other common names: Burton Lim's Yellow-shouldered Bat Taxonomy. Sturnira burtonlimi Velazco & Patterson, 2014 , “Ojo de Agua, 2 km N of Santa Clara, Chiriqui , Panama , approximately 8°42’N , 82°45’W , 1500m .” Sturnira burtonlimi was previously confused with S. ludovici and S. hondurensis . Molecular and morphological analyses identified it as a distinct species. Monotypic. Distribution. Known only from two localities, one in C Costa Rica ( Cartago Province ) and the other in W Panama (type locality). Descriptive notes. Head—body 70-72 mm (tailless), ear 15 mm , hindfoot 14 mm , forearm 44 mm ; weight 19 g (all measurements from two specimens). Burton's Yellow-shouldered Bat is medium-sized, similar to the Honduran Yellow-shouldered Bat ( S. hondurensis ). Dorsal fur is dark brown. Dorsal hairs have four bands, with short pale gray base (c.10% of the length of each hair), long dark gray band (c.40%), long pale gray band (c.40%), and short dark brown tip (c.10%). Ventral fur is dark brown and tricolored, with narrow pale gray base, long dark brown subterminal band (c.45% of each hair), and long gray terminal band (c.45%). Furis long and woolly: 7 mm between shoulders and 5 mm on chest. Proximal part of forearm is densely furred with short hairs. Wing membranes of are dark brown. Dorsal surfaces of femur and tibia are densely covered with long hairs; dorsal surfaces of feet are densely covered with short hairs. Tail membrane is very short and well-furred with long hairs. I' is procumbent and atleast twice the height of I, and I, and I, are bicuspidate and subequalin size. Lingual cuspids (metaconid and entoconid) of M, and M, are poorly defined and are not separated by deep notch. Habitat. Collected in pre-montane forest near coffee fields at elevations of 1290-1500 m. Based on E. Dinerstein’s 1986 ecological study on the Costa Rican Highland Yellow-shouldered Bat ( S. ludovici ) that could have actually been Burton’s Yellow-shouldered Bat, fruits of abundant second-growth shrubs are more frequently eaten than mature forest plants. Food and Feeding. Dinerstein stated that Burton’s Yellow-shouldered Bat consumes fruits of 27 plant species, including Solanum spp. ( Solanaceae ), Piperspp., and P. umbellatum ( Piperaceae ). Breeding. Dinerstein stated that Burton’s Yellow-shouldered Bat has birth peaks in April-May and September—October. Activity patterns. Burton's Yellow-shouldered Bat is nocturnal. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The [UCN Red List. Although Burton’s Yellow-shouldered Bat has a restricted distribution, there are no major conservation threats in this area. Bibliography. Dinerstein (1986), Reid (2009), Velazco & Patterson (2013, 2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Sturnira burtonlimi	Sturnira		burtonlimi	Velazco & Patterson	2014	0	ZooKeys	######	Burton Lim's Yellow-shouldered Bat	None.	Panama, Chiriqui, Ojo de Agua, 2 km N of Santa Clara	Costa Rica, Panama.	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Sturnira burtonlimi	23	Burton's Yellow-shouldered Bat	Burton Lim's Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STURNIRINI	Sturnira	Sturnira	burtonlimi	Velazco & B. D. Patterson	2014	0	Sturnira_burtonlimi	Velazco, P. M., & Patterson, B. D. (2014). Two new species of yellow-shouldered bats, genus Sturnira Gray, 1842 (Chiroptera, Phyllostomidae) from Costa Rica, Panama and western Ecuador. ZooKeys, (402), 54.	https://zookeys.pensoft.net/article/3760/	ROM 104294		"Ojo de Agua, 2 km N of Santa Clara, ChiriquÃ­, Panama, approximately 8Â°42'N, 82Â°45'W, 1500m."	8.7	-82.75	burtonlimi Velazco & B. D. Patterson, 2014	recently described	Velazco, P. M., & Patterson, B. D. (2014). Two new species of yellow-shouldered bats, genus Sturnira Gray, 1842 (Chiroptera, Phyllostomidae) from Costa Rica, Panama and western Ecuador. ZooKeys, (402), 43.	Panama|Costa Rica	North America	Neotropic	DD	0	0	0	Sturnira_burtonlimi	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Sturnira burtonlimi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Sturnira	burtonlimi	Velazco &; Patterson, 2014		90000000	Sturnira burtonlimi	Data Deficient		2017	2016-11-15 00:00:00 UTC	3.1	English	This species is listed as Data Deficient because the available information is inadequate to make a reliable assessment of its risk of extinction based on its distribution and/or population status. The three known records come from two localities far apart and not well characterized, meaning that the species could have a very specific combination of conditions with a narrower distribution. Both localities are nearby protected areas.	The species has been taken in premontane forest near coffee fields (Velazco and Patterson 2014). Like other species in the genus, ;Sturnira burtonlimi ; probably forages in the understory and prefers the fruit of plants in the family Solanaceae (Fleming 1986).	Threats to this species are unknown at present.	The available information on this species is based on three specimens, two from the type locality in Panama (Velazco and Patterson 2014). The southern part of Central America is well-studied in regard to its bat fauna, so the species seems to be extremely rare.	Unknown	Sturnira burtonlimi is known from only two localities, one in Costa Rica and the other in Panama (Velazco and Patterson 2014).		Terrestrial	Although there are some protected areas nearby the localities where S. burtonlimi has been collected, there is no confirmation of its presence at any of these. A lot of research is necessary to assess ;the actual situation of its populations, and to establish its distribution.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Sturnira		burtonlimi	Velazco & Patterson	2014	0	ZooKeys	402:54:00	Burton Lim's Yellow-shouldered Bat	None.	Panama, Chiriqui, Ojo de Agua, 2 km N of Santa Clara	Costa Rica, Panama.	Not listed.	Data Deficient		Sturnira burtonlimi	1005076	23	Burton's Yellow-shouldered Bat	Burton Lim's Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STURNIRINI	Sturnira	Sturnira	burtonlimi	Velazco & B. D. Patterson	2014	0	Sturnira_burtonlimi	Velazco, P. M., & Patterson, B. D. (2014). Two new species of yellow-shouldered bats, genus Sturnira Gray, 1842 (Chiroptera, Phyllostomidae) from Costa Rica, Panama and western Ecuador. ZooKeys, (402), 54.	https://zookeys.pensoft.net/article/3760/	ROM 104294		"Ojo de Agua, 2 km N of Santa Clara, ChiriquÃ­, Panama, approximately 8Â°42'N, 82Â°45'W, 1500m."	8.7	-82.75	burtonlimi Velazco & B. D. Patterson, 2014	recently described	Velazco, P. M., & Patterson, B. D. (2014). Two new species of yellow-shouldered bats, genus Sturnira Gray, 1842 (Chiroptera, Phyllostomidae) from Costa Rica, Panama and western Ecuador. ZooKeys, (402), 43.				Panama|Costa Rica	North America	Neotropic	DD	0	0	0	Sturnira_burtonlimi	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Sturnira_burtonlimi	1005076	23	Burton's Yellow-shouldered Bat	Burton Lim's Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Sturnirini	Sturnira	Sturnira	burtonlimi	Velazco & B. D. Patterson	0	Sturnira burtonlimi	Velazco, P.M. and Patterson, B.D. 2014-04-16. Two new species of yellow-shouldered bats, genus _Sturnira_ Gray, 1842 (Chiroptera, Phyllostomidae) from Costa Rica, Panama and western Ecuador. ZooKeys 402:43-66.	https://doi.org/10.3897/zookeys.402.7228	ROM:MAM:104294	holotype	http://portal.vertnet.org/o/rom/mammals?id=uri-catalog-rom-mammals-104294	"Ojo de Agua, 2 km N of Santa Clara, ChiriquÃ­, Panama, approximately 8Â°42'N, 82Â°45'W, 1500m."	8.7	-82.75	recently described	Velazco, P. M., & Patterson, B. D. (2014). Two new species of yellow-shouldered bats, genus Sturnira Gray, 1842 (Chiroptera, Phyllostomidae) from Costa Rica, Panama and western Ecuador. ZooKeys, (402), 43.				Panama|Costa Rica	North America	Neotropic	DD	0	0	0	Sturnira_burtonlimi	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Sturnira		burtonlimi	Velazco & Patterson	2014	0	ZooKeys	402:54:00	Burton Lim's Yellow-shouldered Bat	None.	Panama, Chiriqui, Ojo de Agua, 2 km N of Santa Clara	Costa Rica, Panama.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/88152206/88152209/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Sturnira burtonlimi; Sturnira burtonlimi; Sturnira burtonlimi; Sturnira burtonlimi; Sturnira burtonlimi; burtonlimi; Sturnire de Burton Lim; Burton-Gelbschulterfledermaus; Sturniro de Burton Lim; Burton Lim's Yellow-shouldered Bat; Burton's Yellow-shouldered Bat; Burton Lim's Yellow-shouldered Bat; Burton Lim's Yellow-shouldered Bat; S. burtonlimi
