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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1562	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi	Sturnira aratathomasi		[MSW2] Subgenus Sturnira. See Soriano and Molinari (1987, Mammalian Species, 284).; [MSW3] Subgenus Sturnira. See Soriano and Molinari (1987).; [HMW] Sturnira aratathomas: Peterson & Tamsit, 1968 , “ 2 km south of Pance (approximately 20 km southwest of Cali), Dept. of Valle, Colombia , at 1650 metres elevation.” Phylogenetic analyses of molecular data indicate that S. aratathomasi represents a unique lineage of Sturnira , not closely related to any other species. Monotypic.; [batnames2022] Subgenus Sturnira .  See Soriano and Molinari (1987).; [batnames2023] Subgenus Sturnira .  See Soriano and Molinari (1987).; [batnames2025_1.7] Subgenus Sturnira. See Soriano and Molinari (1987).														aratathomasi				aratathomasi	aratathomasi			aratathomasi R. L. Peterson & Tamsitt, 1968		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		SW Colombia, W Ecuador	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Sturnira aratathomasi	Colombia, Valle, 2 km S. Pance (ca. 20 km S.W. Cali), 1650 m.	Peterson and Tamsitt	1968	R. Ont. Mus. Life Sci. Occ. Pap., 12:1.	Distribution: Restricted to the northern Andean region from western Venezuela to northern Ecuador.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Venezuela, W Ecuador	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peterson and Tamsitt	1968	R. Ontario Mus. Life Sci. Occas. Pap., 12:1.	Subgenus Sturnira. See Soriano and Molinari (1987, Mammalian Species, 284).	Colombia, Ecuador, NW Venezuela, Peru.	Colombia, Valle, 2 km S Pance (ca. 20 km SW Cali), 1,650 m.		PETERSON & TAMSITT	1968	Lingual cusps of anterior and middle lower molars (metaconid and entoconid) well defined by vertical notches. Size relatively large (forearm length, 56-61 mm; condylobasal length, 25-27 mm).	Distribution: Restricted to the northern Andean region from western Venezuela to northern Ecuador.	No subspecies.		85	species	S. aratathomasi	PETERSON & TAMSITT	1968	Sturnira	subgenus	Sturnira aratathomasi				Lingual cusps of anterior and middle lower molars (metaconid and entoconid) well defined by vertical notches. Size relatively large (forearm length, 56-61 mm; condylobasal length, 25-27 mm).	No subspecies.		5. S. aratathomasi PETERSON & TAMSITT 1968.	5	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Stenodermatinae	Sturnirini	Sturnira aratathomasi	Sturnira	Sturnira	aratathomasi	Peterson and Tamsitt		1968		R. Ontario Mus. Life Sci. Occas. Pap.	12		1		Aratathomas's Yellow-shouldered Bat	Colombia, Valle, 2 km S Pance (ca. 20 km SW Cali), 1,650 m.	Colombia, Ecuador, NW Venezuela, Peru.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Subgenus Sturnira. See Soriano and Molinari (1987).	03A687BCFFF0FFF013B6FCA6FADCF890	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	543	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF8AFF8D1647F905FC8AF660.xml	Sturnira aratathomasi	Phyllostomidae	Sturnira	aratathomasi	Peterson & Tamsitt	1968	Sturnire Arata-Thomas @fr | Arata-Thomas-Gelbschulterfledermaus @de | Sturniro de Arata-Thomas @es | Arata Yellow-shouldered Bat @en | Giant Yellow-shouldered Bat @en | Thomas's Yellow-shouldered Bat @en	Sturnira aratathomas: Peterson & Tamsit, 1968 , “ 2 km south of Pance (approximately 20 km southwest of Cali), Dept. of Valle, Colombia , at 1650 metres elevation.” Phylogenetic analyses of molecular data indicate that S. aratathomasi represents a unique lineage of Sturnira , not closely related to any other species. Monotypic.	W Colombia , W Venezuela , Ecuador , and Peru . Records from Ecuador do not have precise localities.	Head-body 88-101 mm (tailless), ear 19-22 mm, hindfoot 17-21 mm, forearm 55-61 mm; weight 47-54 g. The Arata-Thomas Yellow-shouldered Bat is one of the largest species of Sturnira . Upperparts range from grayish brown to brownish tones; head is pale grayish brown, with some darker spots on rostrum and frontal area. Fur is long (7-8 mm), soft, and woolly. Hairs on shoulder area have four distinguishable bands: short white basal band, light gray epibasal band, white subterminal band, and grayish brown terminal band. Basal band is missing on back. Shoulder patches (sebaceous glands) are not evident. Underparts are paler, usually with three-banded hairs. Fur is longer on posterior one-half of back. Elbow area and adjacent areas of wings are sparsely covered with short hairs. Wing membranes are mostly blackish. Tail membraneis very short and moderately furred; there is no calcar. Feet are hairy. Skull is robust and elongated. Upper incisors protrude, and cusps of I! are pointed and have no contact at tips.	Dry inter-Andean lower, pre-montane, and montane forests, including cloudy, evergreen, and secondary forests, at elevations of 1650-3165 m (usually above 2000 m ). In Colombia , the Arata-Thomas Yellow-shouldered Bat was recorded in all three Cordilleras, but in Peru , it is reported only in lower montane forests on eastern slope of the Andes. Based on this distribution, it is expected to occur in northern Bolivia . On following pages: 122. Honduran Yellow-shouldered Bat ( Sturnira hondurensis ); 123. Burton's Yellow-shouldered Bat ( Sturnira burtonlimi ); 124. Highland Yellow-shouldered Bat ( Sturnira ludovici ); 125. Adriana’s Yellow-shouldered Bat ( Sturnira adrianae ); 126. Tschudi’s Yellow-shouldered Bat ( Sturnira oporaphilum ); 127. Mistratoan Yellow-shouldered Bat ( Sturnira mistratensis ); 128. Soriano's Yellow-shouldered Bat ( Sturnira sorianoi ); 129. Choco Yellow-shouldered Bat ( Sturnira koopmanhill)); 130. Talamancan Yellow-shouldered Bat ( Sturnira mordax ); 131. Perla Yellow-shouldered Bat ( Sturnira perla ); 132. Tilda's Yellow-shouldered Bat ( Sturnira tildae ); 133. Hairy Yellow-shouldered Bat ( Sturnira erythromos ); 134. Bogota Yellow-shouldered Bat ( Sturnira bogotensis ); 135. Greater Yellow-shouldered Bat ( Sturnira magna ); 136. Northern Yellow-shouldered Bat ( Sturnira parvidens ); 137 Baker's Yellow-shouldered Bat ( Sturnira baker ); 138. Little Yellow-shouldered Bat ( Sturnira lilium ); 139. Gianna's Yellow-shouldered Bat ( Sturnira giannae ); 140. Dominica Yellow-shouldered Bat ( Sturnira angell); 141. Paulson's Yellow-shouldered Bat ( Sturnira paulsoni ); 142. Luis's Yellow-shouldered Bat ( Sturnira luisi ).	In one of the few ecological studies in Colombia , the Arata-Thomas Yellow-shouldered Bat preferred fruits of Solanaceae (44% frequency), with a particular preference for species of Solanum ( Solanaceae ). It is expected to have a predominantly frugivorous diet but including some pollen, nectar, and insects.	Pregnant Arata-Thomas Yellow-shouldered Bats have been found in February and August, suggesting bimodal polyestry. Its reproductive cycle probably follows the two peaks in food resources associated with wet seasons in Andean forests.	The Arata-Thomas Yellow-shouldered Bat is nocturnal. It has been captured in understories and subcanopies. A cave is the only record of a roost.	The Arata-Thomas Yellow-shouldered Bat was found sharing a cave with the Tailed Tailless Bat ( Anoura caudifer ) and the Bidentate Yellow-shouldered Bat ( Sturnira bidens ) at Cueva de los Guacharos, Cordillera Oriental of Colombia .	Classified as Least Concern on The IUCN Red List. The Arata-Thomas Yellow-shouldered Bat might be locally rare, but there is no information on specific requirements for its occurrence at a local scale. Its total distribution is rather extensive.	Estrada-Villegas et al. (2007) | Gardner ( 2008g) | McCarthy et al. (1991) | Pacheco & Hocking (2006) | Pacheco & Patterson (1991) | Peterson & Tamsitt (1968) | Soriano & Molinari (1984, 1987) | Tamsitt et al. (1986) | Thomas & McMurray (1974) | Velazco & Patterson (2013)	https://zenodo.org/record/6458851/files/figure.png	121. Arata-Thomas Yellow-shouldered Bat Sturnira aratathomasi French: Sturnire Arata-Thomas / German: Arata-Thomas-Gelbschulterfledermaus / Spanish: Sturniro de Arata-Thomas Other common names: Arata Yellow-shouldered Bat , Giant Yellow-shouldered Bat , Thomas's Yellow-shouldered Bat Taxonomy. Sturnira aratathomas: Peterson & Tamsit, 1968 , “ 2 km south of Pance (approximately 20 km southwest of Cali), Dept. of Valle, Colombia , at 1650 metres elevation.” Phylogenetic analyses of molecular data indicate that S. aratathomasi represents a unique lineage of Sturnira , not closely related to any other species. Monotypic. Distribution. W Colombia , W Venezuela , Ecuador , and Peru . Records from Ecuador do not have precise localities. Descriptive notes. Head-body 88-101 mm (tailless), ear 19-22 mm, hindfoot 17-21 mm, forearm 55-61 mm; weight 47-54 g. The Arata-Thomas Yellow-shouldered Bat is one of the largest species of Sturnira . Upperparts range from grayish brown to brownish tones; head is pale grayish brown, with some darker spots on rostrum and frontal area. Fur is long (7-8 mm), soft, and woolly. Hairs on shoulder area have four distinguishable bands: short white basal band, light gray epibasal band, white subterminal band, and grayish brown terminal band. Basal band is missing on back. Shoulder patches (sebaceous glands) are not evident. Underparts are paler, usually with three-banded hairs. Fur is longer on posterior one-half of back. Elbow area and adjacent areas of wings are sparsely covered with short hairs. Wing membranes are mostly blackish. Tail membraneis very short and moderately furred; there is no calcar. Feet are hairy. Skull is robust and elongated. Upper incisors protrude, and cusps of I! are pointed and have no contact at tips. I are trilobed, I, are bilobed, and molars have deep lingualclefts on M| and M,, Habitat. Dry inter-Andean lower, pre-montane, and montane forests, including cloudy, evergreen, and secondary forests, at elevations of 1650-3165 m (usually above 2000 m ). In Colombia , the Arata-Thomas Yellow-shouldered Bat was recorded in all three Cordilleras, but in Peru , it is reported only in lower montane forests on eastern slope of the Andes. Based on this distribution, it is expected to occur in northern Bolivia . On following pages: 122. Honduran Yellow-shouldered Bat ( Sturnira hondurensis ); 123. Burton's Yellow-shouldered Bat ( Sturnira burtonlimi ); 124. Highland Yellow-shouldered Bat ( Sturnira ludovici ); 125. Adriana’s Yellow-shouldered Bat ( Sturnira adrianae ); 126. Tschudi’s Yellow-shouldered Bat ( Sturnira oporaphilum ); 127. Mistratoan Yellow-shouldered Bat ( Sturnira mistratensis ); 128. Soriano's Yellow-shouldered Bat ( Sturnira sorianoi ); 129. Choco Yellow-shouldered Bat ( Sturnira koopmanhill)); 130. Talamancan Yellow-shouldered Bat ( Sturnira mordax ); 131. Perla Yellow-shouldered Bat ( Sturnira perla ); 132. Tilda's Yellow-shouldered Bat ( Sturnira tildae ); 133. Hairy Yellow-shouldered Bat ( Sturnira erythromos ); 134. Bogota Yellow-shouldered Bat ( Sturnira bogotensis ); 135. Greater Yellow-shouldered Bat ( Sturnira magna ); 136. Northern Yellow-shouldered Bat ( Sturnira parvidens ); 137 Baker's Yellow-shouldered Bat ( Sturnira baker ); 138. Little Yellow-shouldered Bat ( Sturnira lilium ); 139. Gianna's Yellow-shouldered Bat ( Sturnira giannae ); 140. Dominica Yellow-shouldered Bat ( Sturnira angell); 141. Paulson's Yellow-shouldered Bat ( Sturnira paulsoni ); 142. Luis's Yellow-shouldered Bat ( Sturnira luisi ). Food and Feeding. In one of the few ecological studies in Colombia , the Arata-Thomas Yellow-shouldered Bat preferred fruits of Solanaceae (44% frequency), with a particular preference for species of Solanum ( Solanaceae ). It is expected to have a predominantly frugivorous diet but including some pollen, nectar, and insects. Breeding. Pregnant Arata-Thomas Yellow-shouldered Bats have been found in February and August, suggesting bimodal polyestry. Its reproductive cycle probably follows the two peaks in food resources associated with wet seasons in Andean forests. Activity patterns. The Arata-Thomas Yellow-shouldered Bat is nocturnal. It has been captured in understories and subcanopies. A cave is the only record of a roost. Movements, Home range and Social organization. The Arata-Thomas Yellow-shouldered Bat was found sharing a cave with the Tailed Tailless Bat ( Anoura caudifer ) and the Bidentate Yellow-shouldered Bat ( Sturnira bidens ) at Cueva de los Guacharos, Cordillera Oriental of Colombia . Status and Conservation. Classified as Least Concern on The IUCN Red List. The Arata-Thomas Yellow-shouldered Bat might be locally rare, but there is no information on specific requirements for its occurrence at a local scale. Its total distribution is rather extensive. Bibliography. Estrada-Villegas et al. (2007), Gardner ( 2008g ), McCarthy et al. (1991), Pacheco & Hocking (2006), Pacheco & Patterson (1991), Peterson & Tamsitt (1968), Soriano & Molinari (1984, 1987), Tamsitt et al. (1986), Thomas & McMurray (1974), Velazco & Patterson (2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Sturnira aratathomasi	Sturnira	Sturnira	aratathomasi	Peterson & Tamsitt	1968	0	R. Ontario Mus. Life Sci. Occas. Pap.	12:01	Aratathomas' Yellow-shouldered Bat	None.	Colombia, Valle, 2 km S Pance (ca. 20 km SW Cali), 1,650 m.	Colombia, Ecuador, NW Venezuela, Peru.	Not listed.	Least Concern	Subgenus Sturnira .  See Soriano and Molinari (1987).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Sturnira aratathomasi	23	Arata-Thomas Yellow-shouldered Bat	Arata Yellow-shouldered Bat|Giant Yellow-shouldered Bat|Thomas's Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STURNIRINI	Sturnira	Sturnira	aratathomasi	R. L. Peterson & Tamsitt	1968	0	Sturnira_aratathomasi	Peterson, R. L., & Tamsitt, J. R. (1968). A New Species of Bat of the Genus Sturnira (Family Phyllostomatidae) from Northwestern South America. Life Sciences Occasional Papers, Royal Ontario Museum, 12, 1.	https://www.biodiversitylibrary.org/item/123473#page/5/mode/1up	USNM 395158		"2 km south of Pance (approximately 20 km southwest of Cali), Dept. of Valle, Colombia, at 1650 metres elevation."			aratathomasi R. L. Peterson & Tamsitt, 1968	NA	NA	Colombia|Venezuela|Ecuador|Peru	South America	Neotropic	LC	0	0	0	Sturnira_aratathomasi	0	sciname match	Sturnira_aratathomasi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	20949	Sturnira aratathomasi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Sturnira	aratathomasi	Peterson &; Tamsitt, 1968		20000000	Sturnira aratathomasi	Least Concern		2016	2016-05-24 00:00:00 UTC	3.1	English	Listed as Least Concern because of its large geographic distribution; although it might be locally rare, there is no information on specific requirements for its occurrence at a local scale. Its habitat is heavily fragmented and in portions of its range forests is rapidly being converted for illicit crops (poppy). Some populations in Colombia are more threatened than others. At the present, these impacts are not noticeable on their populations or distribution.	This species is rare and poorly known. This bat is mainly frugivorous (Soriano and Molinari 1987) and possibly feeds on pollen and nectar. A single young is born annually (Eisenberg and Redford, 1999). Associtaed to habitats of dry interandean, and lower and upper montane forests from 1,650 to 3,165 m a.s.l. (Pacheco and Hocking 2006).	None known, however, Andean habitats like the ones used by this species are being rapidly degraded and heavily fragmented. In portions of its geographic range, forests are rapidly being converted for illicit crops (coca, poppy).	This is one of the less known species in the genus, and is rare to unknown through its geographic range (Linares 1998, Pacheco and Hocking 2006).	Unknown	This Andean species occurs in Venezuela, Colombia, Ecuador and Peru (Gardner 2008). The records from Ecuador have no precise locality but correspond to this species (Peterson and Tamsitt 1968). A recent record extend this range to southern Peru (Pacheco and Hocking 2006).		Terrestrial	Occurs in a few protected areas in Venezuela, Colombia and Peru; however, much of the critical areas connecting the scattered localities where the species is recorded are unprotected and facing a number of threats.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Sturnira	Sturnira	aratathomasi	Peterson & Tamsitt	1968	0	R. Ontario Mus. Life Sci. Occas. Pap.	12:01	Aratathomas' Yellow-shouldered Bat	None.	Colombia, Valle, 2 km S Pance (ca. 20 km SW Cali), 1,650 m.	Colombia, Ecuador, NW Venezuela, Peru.	Not listed.	Least Concern	Subgenus Sturnira .  See Soriano and Molinari (1987).	Sturnira aratathomasi	1005072	23	Arata-Thomas Yellow-shouldered Bat	Arata Yellow-shouldered Bat|Giant Yellow-shouldered Bat|Thomas's Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STURNIRINI	Sturnira	Sturnira	aratathomasi	R. L. Peterson & Tamsitt	1968	0	Sturnira_aratathomasi	Peterson, R. L., & Tamsitt, J. R. (1968). A New Species of Bat of the Genus Sturnira (Family Phyllostomatidae) from Northwestern South America. Life Sciences Occasional Papers, Royal Ontario Museum, 12, 1.	https://www.biodiversitylibrary.org/item/123473#page/5/mode/1up	USNM 395158		"2 km south of Pance (approximately 20 km southwest of Cali), Dept. of Valle, Colombia, at 1650 metres elevation."			aratathomasi R. L. Peterson & Tamsitt, 1968	NA	NA				Colombia|Venezuela|Ecuador|Peru	South America	Neotropic	LC	0	0	0	Sturnira_aratathomasi	0	sciname match	Sturnira_aratathomasi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Sturnira_aratathomasi	1005072	23	Arata-Thomas Yellow-shouldered Bat	Arata Yellow-shouldered Bat|Giant Yellow-shouldered Bat|Thomas's Yellow-shouldered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Sturnirini	Sturnira	Sturnira	aratathomasi	R. L. Peterson & Tamsitt	0	Sturnira aratathomasi	Peterson, R.L. and Tamsitt, J.R. 1968-11-15. A new species of bat of the genus _Sturnira_ (family Phyllostomatidae) from northwestern South America. Royal Ontario Museum, Life Sciences Occasional Papers 12:1-8.	https://www.biodiversitylibrary.org/page/39966441	USNM:MAMM:395158	holotype	http://n2t.net/ark:/65665/372862983-d75b-491e-a57b-247676fd7841	"2 km south of Pance (approximately 20 km southwest of Cali), Dept. of Valle, Colombia, at 1650 metres elevation."			NA	NA				Colombia|Venezuela|Ecuador|Peru	South America	Neotropic	LC	0	0	0	Sturnira_aratathomasi	0	sciname match	Sturnira_aratathomasi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Sturnira	Sturnira	aratathomasi	Peterson & Tamsitt	1968	0	R. Ontario Mus. Life Sci. Occas. Pap.	12:01	Aratathomas' Yellow-shouldered Bat	None.	Colombia, Valle, 2 km S Pance (ca. 20 km SW Cali), 1,650 m.	Colombia, Ecuador, NW Venezuela, Peru.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/20949/22052176/' target='_blank'>Least Concern</a>	Subgenus Sturnira. See Soriano and Molinari (1987).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Sturnira aratathomasi; Sturnira aratathomasi; Sturnira aratathomasi; Sturnira aratathomasi; Sturnira aratathomasi; Sturnira aratathomasi; aratathomasi; Sturnire Arata-Thomas; Arata-Thomas-Gelbschulterfledermaus; Sturniro de Arata-Thomas; Arata Yellow-shouldered Bat; Giant Yellow-shouldered Bat; Thomas's Yellow-shouldered Bat; Arata-Thomas Yellow-shouldered Bat; Arata Yellow-shouldered Bat; Giant Yellow-shouldered Bat; Thomas's Yellow-shouldered Bat; Aratathomas's Yellow-shouldered Bat; Aratathomas' Yellow-shouldered Bat; S. aratathomasi