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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1552	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nycticeius balstoni [synonym of]	Nycticeius balstoni [synonym of]	Scotorepens sanborni	Nycticeius sanborni	Nycticeius sanborni	Scotorepens sanborni	Scotorepens sanborni	Scotorepens sanborni	Scotorepens sanborni	Scotorepens sanborni	Scotorepens sanborni	Scotorepens sanborni	Scotorepens sanborni	Scotorepens sanborni	Scotorepens sanborni		[MSW2] Subgenus Scotorepens. Included in N. balstoni by Koopman (1978fl), but see Kitchener and Caputi (1985).; [MSW3] Included in balstoni by Koopman (1978a), but see Kitchener and Caputi (1985). Reviewed by Kitchener et al. (1994c); also see Flannery (1995a, b) and Bonaccorso (1998).; [HMW] Scoteinus sanborni Troughton, 1937 , East Cape, Milne Bay Province , Papua New Guinea . Placement of Scotorepens and Scoteanax is currently uncertain. They seem to be closely related, but their relationship to other genera of Vespertilionidae is uncertain. They are tentatively included in Eptesicini here, but genetic data are needed to establish their position in the family. They might be related to other Australasian vespertilionids such as Falsistrellus , Vespadelus , Chalinolobus , and Nyctophilus . Scotorepens is in need of revision according to unpublished morphological data, and there is a well-known undescribed species of Scotorepens known as the “Central-eastern Broad-nosed Bat” (from south-eastern Queensland and north-eastern New South Wales ) that is sympatric with the three other species of Scotorepens : S. greyii , S. balstoni , and S. orion . Monotypic.; [batnames2022] Included in balstoni by Koopman (1978 a ), but see Kitchener and Caputi (1985). Reviewed by Kitchener et al. (1994 c );also see Flannery (1995 a , b ) and Bonaccorso (1998).; [IUCN] The western and eastern Australian populations have been shown to have significant genetic differences (T. Reardon pers. comm.).; [batnames2023] Included in balstoni by Koopman (1978 a ), but see Kitchener and Caputi (1985). Reviewed by Kitchener et al. (1994 c );also see Flannery (1995 a , b ) and Bonaccorso (1998).; [batnames2025_1.7] Included in balstoni by Koopman (1978a), but see Kitchener and Caputi (1985). Reviewed by Kitchener et al. (1994c);also see Flannery (1995a, b) and Bonaccorso (1998).														sanborni	The western and eastern Australian populations have been shown to have significant genetic differences (T. Reardon pers. comm.).			sanborni	sanborni			sanborni (Troughton, 1937)						N/A					Distribution: Confined to northwestern Australia, northeastern Australia and southeastern New Guinea.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Little northern broad-nosed N Australia, S, SE New bat	(Scotorepens greyi) Guinea	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Troughton	1937	Aust. Zool., 8:280.	Subgenus Scotorepens. Included in N. balstoni by Koopman (1978fl), but see Kitchener and Caputi (1985).	SE New Guinea; NE Queensland, Northern Territory, and N Western Australia (Australia).	New Guinea, Papua, Milne Bay prov.. East Cape.		TROUGHTON	1937	Size medium (forearm length, 31-35 mm). Rostrum usually fairly broad.	Distribution: Confined to northwestern Australia, northeastern Australia and southeastern New Guinea.	No subspecies.		126	species	N. sanborni	TROUGHTON	1937	Scotorepens	subgenus	Nycticeius sanborni				Size medium (forearm length, 31-35 mm). Rostrum usually fairly broad.	No subspecies.		4. N. sanborni (TROUGHTON 1937).	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nycticeiini	Scotorepens sanborni	Scotorepens		sanborni	Troughton	y	1937		Aust. Zool.	8		280		Northern Broad-nosed Bat	Papua New Guinea, Milne Bay Prov., East Cape.	W Timor; SE New Guinea; NE Queensland, Northern Territory, and N Western Australia (Australia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc) as Nycticeius sanborni.		Included in balstoni by Koopman (1978a), but see Kitchener and Caputi (1985). Reviewed by Kitchener et al. (1994c); also see Flannery (1995a, b) and Bonaccorso (1998).	4C3D87E8FFFD6A42FF4E935E16DFB95A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	837	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFB46A14FA829CF91C69B221.xml	Scotorepens sanborni	Vespertilionidae	Scotorepens	sanborni	Troughton	1937	Sérotine de Sanborn @fr | Sanborn-Breitnasenfledermaus @de | Scotorepo de Sanborn @es	Scoteinus sanborni Troughton, 1937 , East Cape, Milne Bay Province , Papua New Guinea . Placement of Scotorepens and Scoteanax is currently uncertain. They seem to be closely related, but their relationship to other genera of Vespertilionidae is uncertain. They are tentatively included in Eptesicini here, but genetic data are needed to establish their position in the family. They might be related to other Australasian vespertilionids such as Falsistrellus , Vespadelus , Chalinolobus , and Nyctophilus . Scotorepens is in need of revision according to unpublished morphological data, and there is a well-known undescribed species of Scotorepens known as the “Central-eastern Broad-nosed Bat” (from south-eastern Queensland and north-eastern New South Wales ) that is sympatric with the three other species of Scotorepens : S. greyii , S. balstoni , and S. orion . Monotypic.	Timor I, SE New Guinea (as well as Dolak I), and NE Western Australia , NW Northern Territory , and NE Queensland , Australia .	Head-body 36-8-57 mm, tail 27-5-38-9 mm, ear 9-1-13-5 mm, hindfoot 7-1-9 mm, forearm 27-9-37 mm; weight 5-7-9-1 g. The Northern Broadnosed Bat is almost identical to the Little Broad-nosed Bat ( S. greyii ), which can only be identified by genetics, skull measurements, and plotting braincase width against palatal length. Forearm length increases from west to east and from south to north. Dorsal pelage of the Northern Broad-nosed Bat is tawny brown (hairs brown with light reddish tinge and dark bases); ventral pelage is slightly lighter. Ears are relatively slender and subtriangular, anterior edge being smoothly convex with rounded tip; tragus is evenly curved upward, with concave anterior edge, moderately convex posterior edge, and rounded tip. Muzzle is broad, with square-shape and inflated glands on each side, and nostrils are simple and rounded, diverging from one another. Bare parts offace, ears, and membranes are dark brown. Wing attaches at base offifth toe, and uropatagium extends from long calcar to tail tip; calcar stretches about halfway to tail and has small calcaneal lobe. Glans penis has up to ten spines on head, mainly in two long rows. Baculum is short (2-4-2-6 mm), with relatively robust shaft thatis slightly curved in lateral view; distal head is bifurcated, with two short narrow prongs behind which shaft occasionally narrows slightly before enlarging into slight lateral flanges; base is moderately wide, gently curved posterodorsally, and occasionally more squared in line of shaft; there is a groove on caudal and cranial surface of base that is shallow and wide and meets as a notch at apex. Skull is short, with relatively wide cranium; sagittal and lambdoidal crests are absent; anterior palatal notch is semicircular, usually ending in line with anterior (occasionally posterior) edge of P* I? is unicuspid and does not touch C'; P* is one-half to two-thirds the height of C' and touchesit; lower incisors are tricuspid; and P, is small and less than one-half the height of P,. Dental formula for all species of Scotorepensis11/3,C1/1,P1/2,M 3/3 (x2) = 30.	Various habitats including monsoon forests, open woodlands, and heathlands and apparent preference for mangrove and floodplain areas adjacent to paperbark forests (Kimberley region of north-western Australia ) from sea level up to ¢. 2200 m .	The Northern Broad-nosed Bat forages close to cluttered vegetation by aerial-hawking and has been observed foraging 4-5 m aboveground in the forest understory of Queensland . In the Top End region, fecal and stomach samples mainly contained beetles (44-2% by volume), true bugs (27-7%), and hymenopterans (17-7%), along with smaller amounts of lepidopterans, cockroaches, termites, neuropterans, and flies (although individuals were not differentiated from the Little Broad-nosed Bat).	Pregnant Northern Broad-nosed Bats have been caught in August-September. One or two young (generally one) are born in late September or early December.	The Northern Broad-nosed Bat roosts in hollow trees and buildings. It forages along rivers and edges of quiet bays, along paths and roads, and around lights in urban areas. Call shape is steep FM/QCF sweep. In Kimberley, characteristic frequency averaged 41-1 kHz, with durations of 6-12 milliseconds. Another study in the Top End region found characteristic frequencies of 34-6—41-4 kHz (mean 38 kHz), but these were not differentiable from the Little Broad-nosed Bat.	Northern Broad-nosed Bats roost in colonies of a few individuals (often ¢.20 individuals) in tree hollows up to several hundred individuals in buildings.	Classified as Least Concern on The IUCN Red Lust.	Baverstock et al. (1987) | Bonaccorso (1998) | Churchill (2008) | Flannery (1995a, 1995b) | Hall (2008d) | Hutson, Schlitter, Csorba, Bonaccorso & McKenzie (2008) | Kitchener & Caputi (1985) | Kitchener, Adams & Boeadi (1994) | McKenzie & Bullen (2012) | McKenzie & Rolfe (1986) | Milne (2002) | Milne, Armstrong et al. (2004) | Milne, Burwell & Pavey (2016)	https://zenodo.org/record/6398128/files/figure.png	165. Northern Broad-nosed Bat Scotorepens sanborni French: Sérotine de Sanborn / German: Sanborn-Breitnasenfledermaus / Spanish: Scotorepo de Sanborn Taxonomy. Scoteinus sanborni Troughton, 1937 , East Cape, Milne Bay Province , Papua New Guinea . Placement of Scotorepens and Scoteanax is currently uncertain. They seem to be closely related, but their relationship to other genera of Vespertilionidae is uncertain. They are tentatively included in Eptesicini here, but genetic data are needed to establish their position in the family. They might be related to other Australasian vespertilionids such as Falsistrellus , Vespadelus , Chalinolobus , and Nyctophilus . Scotorepens is in need of revision according to unpublished morphological data, and there is a well-known undescribed species of Scotorepens known as the “Central-eastern Broad-nosed Bat” (from south-eastern Queensland and north-eastern New South Wales ) that is sympatric with the three other species of Scotorepens : S. greyii , S. balstoni , and S. orion . Monotypic. Distribution. Timor I, SE New Guinea (as well as Dolak I), and NE Western Australia , NW Northern Territory , and NE Queensland , Australia . Descriptive notes. Head-body 36-8-57 mm, tail 27-5-38-9 mm, ear 9-1-13-5 mm, hindfoot 7-1-9 mm, forearm 27-9-37 mm; weight 5-7-9-1 g. The Northern Broadnosed Bat is almost identical to the Little Broad-nosed Bat ( S. greyii ), which can only be identified by genetics, skull measurements, and plotting braincase width against palatal length. Forearm length increases from west to east and from south to north. Dorsal pelage of the Northern Broad-nosed Bat is tawny brown (hairs brown with light reddish tinge and dark bases); ventral pelage is slightly lighter. Ears are relatively slender and subtriangular, anterior edge being smoothly convex with rounded tip; tragus is evenly curved upward, with concave anterior edge, moderately convex posterior edge, and rounded tip. Muzzle is broad, with square-shape and inflated glands on each side, and nostrils are simple and rounded, diverging from one another. Bare parts offace, ears, and membranes are dark brown. Wing attaches at base offifth toe, and uropatagium extends from long calcar to tail tip; calcar stretches about halfway to tail and has small calcaneal lobe. Glans penis has up to ten spines on head, mainly in two long rows. Baculum is short (2-4-2-6 mm), with relatively robust shaft thatis slightly curved in lateral view; distal head is bifurcated, with two short narrow prongs behind which shaft occasionally narrows slightly before enlarging into slight lateral flanges; base is moderately wide, gently curved posterodorsally, and occasionally more squared in line of shaft; there is a groove on caudal and cranial surface of base that is shallow and wide and meets as a notch at apex. Skull is short, with relatively wide cranium; sagittal and lambdoidal crests are absent; anterior palatal notch is semicircular, usually ending in line with anterior (occasionally posterior) edge of P* I? is unicuspid and does not touch C'; P* is one-half to two-thirds the height of C' and touchesit; lower incisors are tricuspid; and P, is small and less than one-half the height of P,. Dental formula for all species of Scotorepensis11/3,C1/1,P1/2,M 3/3 (x2) = 30. Habitat. Various habitats including monsoon forests, open woodlands, and heathlands and apparent preference for mangrove and floodplain areas adjacent to paperbark forests (Kimberley region of north-western Australia ) from sea level up to ¢. 2200 m . Food and Feeding. The Northern Broad-nosed Bat forages close to cluttered vegetation by aerial-hawking and has been observed foraging 4-5 m aboveground in the forest understory of Queensland . In the Top End region, fecal and stomach samples mainly contained beetles (44-2% by volume), true bugs (27-7%), and hymenopterans (17-7%), along with smaller amounts of lepidopterans, cockroaches, termites, neuropterans, and flies (although individuals were not differentiated from the Little Broad-nosed Bat). Breeding. Pregnant Northern Broad-nosed Bats have been caught in August-September. One or two young (generally one) are born in late September or early December. Activity patterns. The Northern Broad-nosed Bat roosts in hollow trees and buildings. It forages along rivers and edges of quiet bays, along paths and roads, and around lights in urban areas. Call shape is steep FM/QCF sweep. In Kimberley, characteristic frequency averaged 41-1 kHz, with durations of 6-12 milliseconds. Another study in the Top End region found characteristic frequencies of 34-6—41-4 kHz (mean 38 kHz), but these were not differentiable from the Little Broad-nosed Bat. Movements, Home range and Social organization. Northern Broad-nosed Bats roost in colonies of a few individuals (often ¢.20 individuals) in tree hollows up to several hundred individuals in buildings. Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Baverstock et al. (1987), Bonaccorso (1998), Churchill (2008), Flannery (1995a, 1995b), Hall (2008d), Hutson, Schlitter, Csorba, Bonaccorso & McKenzie (2008), Kitchener & Caputi (1985), Kitchener, Adams & Boeadi (1994), McKenzie & Bullen (2012), McKenzie & Rolfe (1986), Milne (2002), Milne, Armstrong et al. (2004), Milne, Burwell & Pavey (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scotorepens sanborni	Scotorepens		sanborni	Troughton	1937	1	Aust. Zool.	0.5278	Northern Broad-nosed Bat	None.	Papua New Guinea, Milne Bay Prov., East Cape.	W Timor; SE New Guinea; NE Queensland, Northern Territory, and N Western Australia (Australia).	Not listed.	Least Concern	Included in balstoni by Koopman (1978 a ), but see Kitchener and Caputi (1985). Reviewed by Kitchener et al. (1994 c );also see Flannery (1995 a , b ) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scotorepens sanborni	23	Northern Broad-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Scotorepens	NA	sanborni	Troughton	1937	1	Scoteinus_sanborni	Troughton, E. L. G. (1937). Six new bats (Microchiroptera) from the Australasian region. Australian Zoologist, 8, 280.	https://www.biodiversitylibrary.org/item/119324#page/330/mode/1up	AM A.3176		East Cape, Milne Bay Province, Papua New Guinea.			sanborni (Troughton, 1937)	NA	NA	Indonesia|East Timor|Papua New Guinea|Australia	Asia|Oceania	Australasia/Oceania	LC	0	0	0	Scotorepens_sanborni	0	sciname match	Scotorepens_sanborni	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14947	Scotorepens sanborni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scotorepens	sanborni	(Troughton, 1937)	The western and eastern Australian populations have been shown to have significant genetic differences (T. Reardon pers. comm.).	20000000	Scotorepens sanborni	Least Concern		2020	2019-08-05 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, presumed large population, occurrence in a number of protected areas, and because it is unlikely to be declining fast enough to qualify for listing in a threatened category.	This species has been recorded from lowland tropical moist forest, sclerophyll forest, coastal forests, and savanna. In the Kimberley, it is only recorded in mangroves (N. McKenzie pers. comm.). It forages along rivers and edges of quiet bays, along paths and roads, and around lights in urban areas. It roosts in hollow trees and buildings. Colonies may consist of a few individuals to several hundred animals. Females gives birth to one or two young (Bonaccorso 1998, Hall 2008).	There appear to be no major threats to this species.	It is a fairly common species.	Stable	This species has been recorded from West Timor (Indonesia), south-eastern New Guinea (Papua New Guinea), Dolak Island (Papua Province, Indonesia), and in Australia where it occurs in north-eastern Queensland, Northern Territory, and northern Western Australia. It presumably ranges more widely in little-surveyed parts of Papua Province, Indonesia. The species occurs from sea level to 2,200 m asl in New Guinea, but is usually found below 100 m asl (Bonaccorso 1998).		Terrestrial	This species occurs in a number of protected areas. Local education programmes on bat species would benefit the conservation of this species. Research into the species ecology, habitat, threats, and population status is needed.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scotorepens		sanborni	Troughton	1937	1	Aust. Zool.	0.527778	Northern Broad-nosed Bat	None.	Papua New Guinea, Milne Bay Prov., East Cape.	W Timor; SE New Guinea; NE Queensland, Northern Territory, and N Western Australia (Australia).	Not listed.	Least Concern	Included in balstoni by Koopman (1978 a ), but see Kitchener and Caputi (1985). Reviewed by Kitchener et al. (1994 c );also see Flannery (1995 a , b ) and Bonaccorso (1998).	Scotorepens sanborni	1005568	23	Northern Broad-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Scotorepens	NA	sanborni	Troughton	1937	1	Scoteinus_sanborni	Troughton, E. L. G. (1937). Six new bats (Microchiroptera) from the Australasian region. Australian Zoologist, 8, 280.	https://www.biodiversitylibrary.org/item/119324#page/330/mode/1up	AM A.3176		East Cape, Milne Bay Province, Papua New Guinea.			sanborni (Troughton, 1937)	NA	NA				Indonesia|East Timor|Papua New Guinea|Australia	Asia|Oceania	Australasia/Oceania	LC	0	0	0	Scotorepens_sanborni	0	sciname match	Scotorepens_sanborni	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotorepens_sanborni	1005568	23	Northern Broad-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Scotorepens	NA	sanborni	Troughton	1	Scoteinus sanborni	Troughton, E.L.G. 1937. Six new bats (Microchiroptera) from the Australasian region. Australian Zoologist 8:274-281.	https://www.biodiversitylibrary.org/page/38709909	AM A.3176	holotype		East Cape, Milne Bay Province, Papua New Guinea.			NA	NA				Indonesia|East Timor|Papua New Guinea|Australia	Asia|Oceania (Continent)	Australasia	LC	0	0	0	Scotorepens_sanborni	0	sciname match	Scotorepens_sanborni	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scotorepens		sanborni	Troughton	1937	1	Aust. Zool.	0.527778	Northern Broad-nosed Bat	None.	Papua New Guinea, Milne Bay Prov., East Cape.	W Timor; SE New Guinea; NE Queensland, Northern Territory, and N Western Australia (Australia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14947/22008930/' target='_blank'>Least Concern</a>	Included in balstoni by Koopman (1978a), but see Kitchener and Caputi (1985). Reviewed by Kitchener et al. (1994c);also see Flannery (1995a, b) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotorepens sanborni; Scotorepens sanborni; Scotorepens sanborni; Scotorepens sanborni; Scotorepens sanborni; Scotorepens sanborni; sanborni; Sérotine de Sanborn; Sanborn-Breitnasenfledermaus; Scotorepo de Sanborn; Northern Broad-nosed Bat; Northern Broad-nosed Bat; Northern Broad-nosed Bat; S. sanborni