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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1550	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nycticeius greyi	Nycticeius greyii	Scotorepens greyi	Nycticeius greyii	Nycticeius greyii	Scotorepens greyii	Scotorepens greyii	Scotorepens greyii	Scotorepens greyii	Scotorepens greyii	Scotorepens greyii	Scotorepens greyii	Scotorepens greyii	Scotorepens greyii	Scotorepens greyii		[MSW2] Subgenus Scotorepens. See Kitchener and Caputi (1985) for content.; [MSW3] Reviewed by Kitchener and Caputi (1985).; [HMW] Scotophilus greyii]. E. Gray, 1842 , Port Essington, Northern Territory , Australia . This species is monotypic.; [batnames2022] Reviewed by Kitchener and Caputi (1985).; [batnames2023] Reviewed by Kitchener and Caputi (1985).; [batnames2025_1.7] Reviewed by Kitchener and Caputi (1985).		(Grey's bat)			(caprenus) (aquilo)	aquilo, caprenus.			aqeilo, caprenus			greyii	greyii - aqeilo, caprenus	greyii, aquilo, caprenus		greyii	greyii - aqeilo, caprenus	greyii, aquilo, caprenus	greyii, greyii, aquilo, caprenus, greyi, aqeilo	greyii 	greyii - aqeilo, caprenus	greyii (J. E. Gray, 1843) [nomen nudum]|greyii (J. E. Gray, 1844)|greyi (J. D. Ogilby, 1892) [incorrect subsequent spelling]|aquilo (Troughton, 1937)|caprenus (Troughton, 1937)|aqeilo Simmons, 2005 [incorrect subsequent spelling | not used as valid]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Little broad-nosed bat	W, N Australia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nycticeius greyii	Australia, Northern Territory, Port Essington.	Gray	1842	Zool. Voy. "Erebus" and "Terror," pl. 20.	Distribution: Ranging across Australia from northern Western Australia to New South Wales, but absent from the Cape York peninsula.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Little broad-nosed bat (Grey's bat)	Australia except SW, NE, SE	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Gray	1842	Zool. Voy. H.M.S. "Erebus" and "Terror," pl. 20.	Subgenus Scotorepens. See Kitchener and Caputi (1985) for content.	Western Australia (excluding the south). Northern Territory, South Australia, New South Wales, and Queensland (Australia). Records from Victoria refer to balstoni.	Australia, Northern Territory, Port Essington.		GRAY	1843	Size relatively small (forearm length, 27-33 mm). Rostrum usually fairly slender.	Distribution: Ranging across Australia from northern Western Australia to New South Wales, but absent from the Cape York peninsula.	No currently recognized subspecies.		126	species	N. greyii	GRAY	1843	Scotorepens	subgenus	Nycticeius greyii				Size relatively small (forearm length, 27-33 mm). Rostrum usually fairly slender.	No currently recognized subspecies.		3. N. greyii (GRAY 1843) ( = caprenus TROUGHTON 1937; aquilo TROUGHTON 1937).	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nycticeiini	Scotorepens greyii	Scotorepens		greyii	Gray	y	1842		Zool. Voy. H.M.S. "Erebus" and "Terror,"			pl. 20		Little Broad-nosed Bat	Australia, Northern Territory, Port Essington.	Western Australia (excluding the south), Northern Territory, South Australia, New South Wales, and Queensland (Australia). Records from Victoria refer to balstoni.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc) as Nycticeius greyii.	aqeilo Troughton, 1937; caprenus Troughton, 1937.	Reviewed by Kitchener and Caputi (1985).	4C3D87E8FFAA6A15FF7C9037179ABA50	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	838	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFAB6A14FF579F0218D3BCB5.xml	Scotorepens greyii	Vespertilionidae	Scotorepens	greyii		1844	Sérotine de Grey @fr | Grey-Breitnasenfledermaus @de | Scotorepo de Grey @es | Grey's Broad-nosed Bat @en	Scotophilus greyii]. E. Gray, 1842 , Port Essington, Northern Territory , Australia . This species is monotypic.	N Western Australia , Northern Territory (including Bathurst I), Queensland except Cape York Peninsula, N & E South Australia , and New South Wales , Australia .	Head-body 37-2-53 mm, tail 25-2-48.-5 mm, ear 9-7-12: 9 mm , forearm 27-3-35 mm; weight 4-8-5 g. The Little Broad-nosed Bat is almost identical to the Northern Broad-nosed Bat (S. sanborni ). Forearm size increases from east to west and from north to south. Dorsal pelage is brown to grayish brown; ventral pelageis slightly lighter. Ears are relatively broad and subtriangular, anterior edge being smoothly convex with rounded tip; tragus is evenly curved upward, with concave anterior edge, moderately convex posterior edge, and narrow and pointed tip. Muzzle is broad, with square shape and inflated glands on each side; nostrils are simple and rounded, diverging from one another. Bare parts of face, ears, and membranes are dark brown. Wings attach at base of fifth toe, and uropatagium extends from long calcar to tail tip; calcar stretches about halfway to tail and has small calcaneal lobe. Glans penis has up to ten spines on head, mainly in two long rows. Baculum is short (2:2-3-1 mm), with relatively robust shaft thatis slightly curved in lateral view; distal head is bifurcated, with two short narrow prongs behind which shaft occasionally narrows slightly before enlarging into slight lateral flanges; base is moderately wide, gently curved posterodorsally, and occasionally more square in line of shaft; groove on caudal and cranial surface of base is shallow and wide and meets as notch at apex. Skull is short, with relatively wide cranium; cranium profile is generally less inflated than in the Northern Broad-nosed Bat; sagittal crest is absent, and lambdoidal crest is moderately developed; anterior palatal notch is semicircular (occasionally spatulated), usually ending at mid-point of P*; I* is unicuspid and does not touch C'; P* is one-half to two-thirds the height of C' and touches it; lower incisors are tricuspid; and P, is small and less than one-half the height of P,.	Monsoon forests, Melaleuca ( Myrtaceae ) forests, tall open forests, open forests, open woodlands, mulga shrublands, mixed shrublands, escarpments, river red gum lined streams, and grasslands. The Little Broad-nosed Bat is common around waterholes and other water sources and tends to be located near permanent water.	Little Broad-nosed Bats forage in canopy within 2 m of foliage and often forage over open spaces and water. They are moderately fast and agile fliers, making abrupt horizontal turns where they roll to near vertical bank angle. They feed on various insects but seem to prefer beetles, true bugs, and ants; they eat moths, termites, cockroaches, katydids, crickets, flies, and lacewings. In the Top End region, fecal and stomach samples mainly contained beetles (44-2% by volume), true bugs (27-7%), and hymenopterans (17-7%), with smaller amounts of lepidopterans, cockroaches, termites, neuropterans, and flies (but individuals used in that study were not differentiated from the Northern Broad-nosed Bat). Although Little Broad-nosed Bats feed mostly aerially, presence offlightless insects in diets suggests that they glean insects off foliage.	Pregnant Little Broad-nosed Bats are known from late August to early November, and twins (sometimes single young) are born in October-November. Volant young appear in mid-December, at which time femalesarestill lactating. Not all females breed every year. All males had enlarged testes in April that were regressed by November.	The Little Broad-nosed Bat is nocturnal. It generally roosts in tree hollows, but it has been reported roosting in hollows in fence posts, space under metal caps of telephone poles, and abandoned buildings. They can enter torpor during the day to conserve energy. In New South Wales , they entered torpor on 83-3% of days for c.7 hours/day, and 44-8% of individuals rewarmed daily using entirely passive rewarming. Call shape is steep FM/QCF sweep, and in Queensland , average minimum frequency was 38-4 kHz, maximum frequency was 64-5 kHz, peak frequency was 40-1 kHz, duration was 7-1 milliseconds, and interpulse interval was 104-5 milliseconds.	Little Broad-nosed Bats roost in colonies of 2-20 individuals.	Classified as Least Concern on The IUCN Red Lust.	Baverstock et al. (1987) | Bondarenco et al. (2016) | Bullen & McKenzie (2002b) | Churchill (2008) | Hall (2009) | Jones & Corben (1993) | Kitchener & Caputi (1985) | Lumsden & Bennett (2005) | McKenzie & Bullen (2012) | Milne (2002) | Milne, Armstrong et al. (2004) | Milne, Burwell & Pavey (2016) | Richards, Reardon & Pennay (2008) | Young & Ford (2000)	https://zenodo.org/record/6398130/files/figure.png	166. Little Broad-nosed Bat Scotorepens greyii French: Sérotine de Grey / German: Grey-Breitnasenfledermaus / Spanish: Scotorepo de Grey Other common names: Grey's Broad-nosed Bat Taxonomy. Scotophilus greyii]. E. Gray, 1842 , Port Essington, Northern Territory , Australia . This species is monotypic. Distribution. N Western Australia , Northern Territory (including Bathurst I), Queensland except Cape York Peninsula, N & E South Australia , and New South Wales , Australia . Descriptive notes. Head-body 37-2-53 mm, tail 25-2-48.-5 mm, ear 9-7-12: 9 mm , forearm 27-3-35 mm; weight 4-8-5 g. The Little Broad-nosed Bat is almost identical to the Northern Broad-nosed Bat (S. sanborni ). Forearm size increases from east to west and from north to south. Dorsal pelage is brown to grayish brown; ventral pelageis slightly lighter. Ears are relatively broad and subtriangular, anterior edge being smoothly convex with rounded tip; tragus is evenly curved upward, with concave anterior edge, moderately convex posterior edge, and narrow and pointed tip. Muzzle is broad, with square shape and inflated glands on each side; nostrils are simple and rounded, diverging from one another. Bare parts of face, ears, and membranes are dark brown. Wings attach at base of fifth toe, and uropatagium extends from long calcar to tail tip; calcar stretches about halfway to tail and has small calcaneal lobe. Glans penis has up to ten spines on head, mainly in two long rows. Baculum is short (2:2-3-1 mm), with relatively robust shaft thatis slightly curved in lateral view; distal head is bifurcated, with two short narrow prongs behind which shaft occasionally narrows slightly before enlarging into slight lateral flanges; base is moderately wide, gently curved posterodorsally, and occasionally more square in line of shaft; groove on caudal and cranial surface of base is shallow and wide and meets as notch at apex. Skull is short, with relatively wide cranium; cranium profile is generally less inflated than in the Northern Broad-nosed Bat; sagittal crest is absent, and lambdoidal crest is moderately developed; anterior palatal notch is semicircular (occasionally spatulated), usually ending at mid-point of P*; I* is unicuspid and does not touch C'; P* is one-half to two-thirds the height of C' and touches it; lower incisors are tricuspid; and P, is small and less than one-half the height of P,. Habitat. Monsoon forests, Melaleuca ( Myrtaceae ) forests, tall open forests, open forests, open woodlands, mulga shrublands, mixed shrublands, escarpments, river red gum lined streams, and grasslands. The Little Broad-nosed Bat is common around waterholes and other water sources and tends to be located near permanent water. Food and Feeding. Little Broad-nosed Bats forage in canopy within 2 m of foliage and often forage over open spaces and water. They are moderately fast and agile fliers, making abrupt horizontal turns where they roll to near vertical bank angle. They feed on various insects but seem to prefer beetles, true bugs, and ants; they eat moths, termites, cockroaches, katydids, crickets, flies, and lacewings. In the Top End region, fecal and stomach samples mainly contained beetles (44-2% by volume), true bugs (27-7%), and hymenopterans (17-7%), with smaller amounts of lepidopterans, cockroaches, termites, neuropterans, and flies (but individuals used in that study were not differentiated from the Northern Broad-nosed Bat). Although Little Broad-nosed Bats feed mostly aerially, presence offlightless insects in diets suggests that they glean insects off foliage. Breeding. Pregnant Little Broad-nosed Bats are known from late August to early November, and twins (sometimes single young) are born in October-November. Volant young appear in mid-December, at which time femalesarestill lactating. Not all females breed every year. All males had enlarged testes in April that were regressed by November. Activity patterns. The Little Broad-nosed Bat is nocturnal. It generally roosts in tree hollows, but it has been reported roosting in hollows in fence posts, space under metal caps of telephone poles, and abandoned buildings. They can enter torpor during the day to conserve energy. In New South Wales , they entered torpor on 83-3% of days for c.7 hours/day, and 44-8% of individuals rewarmed daily using entirely passive rewarming. Call shape is steep FM/QCF sweep, and in Queensland , average minimum frequency was 38-4 kHz, maximum frequency was 64-5 kHz, peak frequency was 40-1 kHz, duration was 7-1 milliseconds, and interpulse interval was 104-5 milliseconds. Movements, Home range and Social organization. Little Broad-nosed Bats roost in colonies of 2-20 individuals. Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Baverstock et al. (1987), Bondarenco et al. (2016), Bullen & McKenzie (2002b), Churchill (2008), Hall (2009), Jones & Corben (1993), Kitchener & Caputi (1985), Lumsden & Bennett (2005), McKenzie & Bullen (2012), Milne (2002), Milne, Armstrong et al. (2004), Milne, Burwell & Pavey (2016), Richards, Reardon & Pennay (2008), Young & Ford (2000).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scotorepens greyii	Scotorepens		greyii	Gray	1842	1	Zool. Voy. H.M.S. "Erebus" and "Terror,"	pl. 20	Little Broad-nosed Bat	 aqeilo Troughton, 1937; caprenus Troughton, 1937.	Australia, Northern Territory, Port Essington.	Western Australia (excluding the south), Northern Territory, South Australia, New South Wales, and Queensland (Australia). Records from Victoria refer to balstoni.	Not listed.	Least Concern	Reviewed by Kitchener and Caputi (1985).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scotorepens greyii	23	Little Broad-nosed Bat	Grey's Broad-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Scotorepens	NA	greyii	J. E. Gray	1842	1						Port Essington, Northern Territory, Australia.			greyii (J. E. Gray, 1843)|aquilo (Troughton, 1937)|caprenus (Troughton, 1937)	NA	NA	Australia	Oceania	Australasia/Oceania	LC	0	0	0	Scotorepens_greyii	0	sciname match	Scotorepens_greyii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14943	Scotorepens greyii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scotorepens	greyii	(Gray, 1843)		200000000	Scotorepens greyii	Least Concern		2021	2019-07-07 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, abundance, occurrence in numerous protected areas and habitats. It is assumed that these features ensure the stability of the species, and if there is, or has been a recent, population decline that the magnitude or rate of this decline is not sufficient to meet IUCN criteria for a threat category. However, it is important to note that due to a lack of broad-scale, long-term population monitoring, there are no hard data to support or reject this assumption.	This small nocturnal insectivorous bat occurs in dry grassland and woodland, sandy deserts, riparian areas along inland rivers, monsoon forest and paperbark swamps. It roosts in colonies of a few individuals to around 20 animals in tree holes, disused buildings, and other available structures (Richards et al. 2008). Its diet includes beetles, bugs, flies, moths, crickets and wingless ants (Richards et al. 2008).	The main threats are likely to be habitat loss and degradation. Feral and domestic cats are likely to prey on the species.	It is a common species (Richards et al. 2008). The population is currently assumed to be relatively stable as there is no evidence of a decline, although there is loss of hollow-bearing trees throughout parts of its range and there are no long-term monitoring programs to confirm status or trends.	Stable	This species is endemic to Australia where it is widespread in northern, central, and eastern parts of the country, mainly in the drier regions, including the arid centre but extends to high rainfall country of the Top End (Richards et al. 2008).		Terrestrial	This species is present in numerous protected areas.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scotorepens		greyii	Gray	1842	1	Zool. Voy. H.M.S. "Erebus" and "Terror,"	pl. 20	Little Broad-nosed Bat	 aqeilo Troughton, 1937; caprenus Troughton, 1937.	Australia, Northern Territory, Port Essington.	Western Australia (excluding the south), Northern Territory, South Australia, New South Wales, and Queensland (Australia). Records from Victoria refer to balstoni.	Not listed.	Least Concern	Reviewed by Kitchener and Caputi (1985).	Scotorepens greyii	1005566	23	Little Broad-nosed Bat	Grey's Broad-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Scotorepens	NA	greyii	J. E. Gray	1842	1						Port Essington, Northern Territory, Australia.			greyii (J. E. Gray, 1843)|aquilo (Troughton, 1937)|caprenus (Troughton, 1937)	NA	NA				Australia	Oceania	Australasia/Oceania	LC	0	0	0	Scotorepens_greyii	0	sciname match	Scotorepens_greyii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotorepens_greyii	1005566	23	Little Broad-nosed Bat	Grey's Broad-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Scotorepens	NA	greyii	J. E. Gray	1	Scotophilus greyii	Gray, J.E. 1844-12-02. Part IV. Pp. 5–8 in Richardson, J. and Gray, J.E. 1844-1875. The Zoology of the Voyage of H.M.S. Erebus & Terror, under the command of Captain Sir James Clark Ross, R.N., F.R.S., during the years 1839 to 1843. Vol. I. Mammalia, Birds. E. W. Janson, London, not continuously paginated pp.	https://www.biodiversitylibrary.org/page/6957560	BMNH:Mamm:1842.8.17.12	lectotype	https://data.nhm.ac.uk/object/ffad2ad1-e8c1-41ea-bd42-2af1f223020e	Port Essington, Northern Territory, Australia.			NA	NA				Australia	Oceania (Continent)	Australasia	LC	0	0	0	Scotorepens_greyii	0	sciname match	Scotorepens_greyii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scotorepens		greyii	Gray	1842	1	Zool. Voy. H.M.S. "Erebus" and "Terror,"	pl. 20	Little Broad-nosed Bat	aqeilo Troughton, 1937; caprenus Troughton, 1937.	Australia, Northern Territory, Port Essington.	Western Australia (excluding the south), Northern Territory, South Australia, New South Wales, and Queensland (Australia). Records from Victoria refer to balstoni.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14943/209531715/' target='_blank'>Least Concern</a>	Reviewed by Kitchener and Caputi (1985).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotorepens greyii; Scotorepens greyii; Scotorepens greyii; Scotorepens greyii; Scotorepens greyii; Scotorepens greyii; aqeilo; caprenus; aqeilo; caprenus; greyii; aquilo; caprenus; Sérotine de Grey; Grey-Breitnasenfledermaus; Scotorepo de Grey; Grey's Broad-nosed Bat; Little Broad-nosed Bat; Grey's Broad-nosed Bat; Little Broad-nosed Bat; Little Broad-nosed Bat; Nycticeius greyii; S. greyii