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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1546	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Scotophilus trujilloi	Scotophilus trujilloi	Scotophilus trujilloi	Scotophilus trujilloi	Scotophilus trujilloi	Scotophilus trujilloi	Scotophilus trujilloi	Scotophilus trujilloi	Scotophilus trujilloi		[HMW] Scotophilus trujilloi Brooks & Bickham, 2014 , “ Kenya : Coastal Province, Kwale District , Moana Marine Station, 1 km S, 2 km E Ukanda ( 4°18°S , 39°35°E ).” Scotophilus trujilloi was recovered as sister to dinganii from which it differs by 4-2% sequence at cytochrome-b. It differs from viridis by 115%. Monotypic.; [batnames2022] In need of revision. Lineage validity confirmed by Demos et al. (2018), but older names amy have priority.; [MDD2022] recently described; the name borbonicus based on specimens reported from Reunion Island likely represents this species; borbonicus is tentatively included under this species as a nomen dubium, as it was likely collected in Tanzania rather than Reunion based on genetic material from the lectotype of borbonicus; [IUCN] This is a newly described species <span class="msoIns">from five samples from coastal Kenya (Brooks and Bickham 2014). Further research is required to verify<span class="msoIns"> the</span><span class="msoIns"> taxonomy<span class="msoIns"> of this taxon</span><span class="msoIns">.</span>; [batnames2023] In need of revision. Lineage validity confirmed by Demos et al. (2018), but older names (e.g., borbonicus ) may have priority. See also Goodman et al. (2020).; [MDD2023] recently described; the name borbonicus based on specimens reported from Reunion Island likely represents this species and it was considered a nomen dubium by Goodman et al., 2020; however, this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi; [MDD2025_2.0] recently described; the name borbonicus based on specimens reported from Reunion Island likely represents this species and it was considered a nomen dubium by Goodman et al., 2020; however, this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi; [batnames2025_1.7] In need of revision. Lineage validity confirmed by Demos et al. (2018), but older names (e.g., borbonicus) may have priority. See also Goodman et al. (2020).; [MDD2025_2.2] recently described; the name borbonicus based on specimens reported from Reunion Island likely represents this species and it was considered a nomen dubium by Goodman et al., 2020; however, this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi														borbonicus, trujilloi	This is a newly described species <span class="msoIns">from five samples from coastal Kenya (Brooks and Bickham 2014). Further research is required to verify<span class="msoIns"> the</span><span class="msoIns"> taxonomy<span class="msoIns"> of this taxon</span><span class="msoIns">.</span>			trujilloi 	trujilloi 			trujilloi Brooks & Bickham, 2014						N/A																																								NA																											4C3D87E8FF7A6AC6FA51931C1D48B85F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	890	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF786AC0FA5791F71D01B918.xml	Scotophilus trujilloi	Vespertilionidae	Scotophilus	trujilloi	Brooks & Bickham	2014	Scotophile de Trujillo @fr | Trujillo-Hausfledermaus @de | Scotofilode Trujillo @es | Trujillo's House Bat @en	Scotophilus trujilloi Brooks & Bickham, 2014 , “ Kenya : Coastal Province, Kwale District , Moana Marine Station, 1 km S, 2 km E Ukanda ( 4°18°S , 39°35°E ).” Scotophilus trujilloi was recovered as sister to dinganii from which it differs by 4-2% sequence at cytochrome-b. It differs from viridis by 115%. Monotypic.	Known only from three localities in SE Kenya ; it might also occur in S Somalia and NE Tanzania .	Head-body 65-4-75- 2 mm , tail 37-3-43- 8 mm , ear 7-2-7- 9 mm , hindfoot 9-7-10- 4 mm , forearm 43-8-46- 2 mm . Dorsal fur is reddish mahogany; venter is orange, with grayish abdomen. Ears are separated, with semi-rounded tips. Ventral plagiopatagium is hairy proximal to body and forearm. Dorsal plagiopatagium, uropatagium, dactylopatagium, tail, legs, and feet are naked. Skull is broad, with rounded orbits; premaxillae is deeply notched; sagittal crest is present but low to medium; zygomatic arch is thin; vomer has well-developed central process; tympanic bullae are spherical and well developed; foramen magnum is round; and occipital condyles are well developed. All mandibular processes are well developed; coronoid processis low and triangular; and angular process extends to same level as mandibular condyle. I? is bilobed, with inner cusp longer and wider than outer cusp; paracone of P* is much longer than metacone, which is longer than hypocone; P* has smaller diameter but similar metacone length compared to M' and M? M! and M? are similar in size and structure, with triangular outline that is notched exteriorly in occlusal view, and interior edgeis shortest; metacone of M' and M? is slightly longer than paracone, and both are longer than hypocone; M* is highly reduced, similar in appearance to metacone and proximal hypocone of M?*, with ellipsoidal outline narrowing exteriorly in occlusal view; lower incisors are small, with I, and I, weakly developed and trilobed; P, 1s shorter than C; lower molars are similar in structure with trapezoid outline in occlusal view, exterior edge is shortest, and paracone is longer than metacone that is longer than hypocone; and M| and M, are similar in size and wider in diameter than M,.	Woodland and forest habitats of coastal Kenya from sea level up to elevations of ¢. 760 m .	No information.	No information.	No information.	No information.	Classified as Least Concern on The IUCN Red List. Trujillo’s Yellow Bat occurs in the Shimba Hills National Reserve.	Brooks & Bickham (2014) | Trujillo et al. (2009)	https://zenodo.org/record/6398420/files/figure.png	287. Trujillo's Yellow Bat Scotophilus trujilloi French: Scotophile de Trujillo / German: Trujillo-Hausfledermaus / Spanish: Scotofilo de Trujillo Other common names: Trujillo's House Bat Taxonomy. Scotophilus trujilloi Brooks & Bickham, 2014 , “ Kenya : Coastal Province, Kwale District , Moana Marine Station, 1 km S, 2 km E Ukanda ( 4°18°S , 39°35°E ).” Scotophilus trujilloi was recovered as sister to dinganii from which it differs by 4-2% sequence at cytochrome-b. It differs from viridis by 115%. Monotypic. Distribution. Known only from three localities in SE Kenya ; it might also occur in S Somalia and NE Tanzania . Descriptive notes. Head-body 65-4-75- 2 mm , tail 37-3-43- 8 mm , ear 7-2-7- 9 mm , hindfoot 9-7-10- 4 mm , forearm 43-8-46- 2 mm . Dorsal fur is reddish mahogany; venter is orange, with grayish abdomen. Ears are separated, with semi-rounded tips. Ventral plagiopatagium is hairy proximal to body and forearm. Dorsal plagiopatagium, uropatagium, dactylopatagium, tail, legs, and feet are naked. Skull is broad, with rounded orbits; premaxillae is deeply notched; sagittal crest is present but low to medium; zygomatic arch is thin; vomer has well-developed central process; tympanic bullae are spherical and well developed; foramen magnum is round; and occipital condyles are well developed. All mandibular processes are well developed; coronoid processis low and triangular; and angular process extends to same level as mandibular condyle. I? is bilobed, with inner cusp longer and wider than outer cusp; paracone of P* is much longer than metacone, which is longer than hypocone; P* has smaller diameter but similar metacone length compared to M' and M? M! and M? are similar in size and structure, with triangular outline that is notched exteriorly in occlusal view, and interior edgeis shortest; metacone of M' and M? is slightly longer than paracone, and both are longer than hypocone; M* is highly reduced, similar in appearance to metacone and proximal hypocone of M?*, with ellipsoidal outline narrowing exteriorly in occlusal view; lower incisors are small, with I, and I, weakly developed and trilobed; P, 1s shorter than C; lower molars are similar in structure with trapezoid outline in occlusal view, exterior edge is shortest, and paracone is longer than metacone that is longer than hypocone; and M| and M, are similar in size and wider in diameter than M,. Habitat. Woodland and forest habitats of coastal Kenya from sea level up to elevations of ¢. 760 m . Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. Trujillo’s Yellow Bat occurs in the Shimba Hills National Reserve. Bibliography. Brooks & Bickham (2014), Trujillo et al. (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scotophilus trujilloi	Scotophilus		trujilloi	Brooks & Bickham	2014	0	Occas. Pap. Mus. Texas Tech Univ.	######	Trujillo's House Bat	None.	Kenya, Coastal Province, Kwale District, Moana Marine Station, 1 km S 2 km E Ukunda	Coastal Province, Kenya.	Not listed.	Least Concern	In need of revision. Lineage validity confirmed by Demos et al. (2018), but older names amy have priority.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scotophilus trujilloi	23	Trujillo's Yellow Bat	Trujillo's House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	SCOTOPHILINI	Scotophilus	NA	trujilloi	Brooks & Bickham	2014	0	Scotophilus_trujilloi	Brooks, D. M. & Bickham, J. W. (2014). New species of Scotophilus (Chiroptera: Vespertilionidae) from sub-saharan Africa. Occasional papers, Museum of Texas Tech University, 326, 16.	https://www.biodiversitylibrary.org/item/264372#page/16/mode/1up	CM 98038		"Kenya: Coastal Province, Kwale District, Moana Marine Station, 1 km S, 2 km E Ukunda (4Â°18'S, 39Â°35'E)."	-4.3	39.583	borbonicus (Ã‰. Geoffroy Saint-Hilaire, 1803) [nomen dubium]|trujilloi Brooks & Bickham, 2014	recently described; the name borbonicus based on specimens reported from Reunion Island likely represents this species; borbonicus is tentatively included under this species as a nomen dubium, as it was likely collected in Tanzania rather than Reunion based on genetic material from the lectotype of borbonicus	Brooks, D. M., & Bickham, J. W. (2014). New species of Scotophilus (Chiroptera: Vespertilionidae) from sub-Saharan Africa. Occasional Papers, Museum of Texas Tech University, 326, 1-21.|Goodman, S. M., Fratpietro, S., & Tortosa, P. (2020). Insight into the Identity and Origin of Scotophilus borbonicus (E. Geoffroy, 1803). Acta Chiropterologica, 22(1), 41-47.	Kenya|Somalia?|Tanzania?	Africa	Afrotropic	LC	0	0	0	Scotophilus_trujilloi	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Scotophilus trujilloi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scotophilus	trujilloi	Brooks &; Bickham, 2014	This is a newly described species <span class="msoIns">from five samples from coastal Kenya (Brooks and Bickham 2014). Further research is required to verify<span class="msoIns"> the</span><span class="msoIns"> taxonomy<span class="msoIns"> of this taxon</span><span class="msoIns">.</span>	80000000	Scotophilus trujilloi	Least Concern		2017	2016-04-25 00:00:00 UTC	3.1	English	This newly described species is known from three localities in coastal Kenya including one protected area (Brooks and Bickham 2014). It may occur in close association with human altered landscapes.	This species appears to occur in woodland and forest habitats of coastal Kenya. Its ecology is poorly known, but it may utilize human-altered landscapes.	The major threat is persecution of roosting sites.	There is no information about population size and trend.	Unknown	S. trujilloi is currently only known from coastal south-east Kenya, but may also occur in north-east Tanzania and southern Somalia.		Terrestrial	This species is known to occur in the Shimba Hills National Park and may occur in the other protected areas in Kenya.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scotophilus		trujilloi	Brooks & Bickham	2014	0	Occas. Pap. Mus. Texas Tech Univ.	326:16:00	Trujillo's House Bat	None.	Kenya, Coastal Province, Kwale District, Moana Marine Station, 1 km S 2 km E Ukunda	Coastal Province, Kenya.	Not listed.	Least Concern	In need of revision. Lineage validity confirmed by Demos et al. (2018), but older names (e.g., borbonicus ) may have priority. See also Goodman et al. (2020).	Scotophilus trujilloi	1005697	23	Trujillo's Yellow Bat	Trujillo's House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	SCOTOPHILINI	Scotophilus	NA	trujilloi	Brooks & Bickham	2014	0	Scotophilus_trujilloi	Brooks, D. M. & Bickham, J. W. (2014). New species of Scotophilus (Chiroptera: Vespertilionidae) from sub-saharan Africa. Occasional papers, Museum of Texas Tech University, 326, 16.	https://www.biodiversitylibrary.org/item/264372#page/16/mode/1up	CM 98038		"Kenya: Coastal Province, Kwale District, Moana Marine Station, 1 km S, 2 km E Ukunda (4Â°18'S, 39Â°35'E)."	-4.3	39.58333	trujilloi Brooks & Bickham, 2014	recently described; the name borbonicus based on specimens reported from Reunion Island likely represents this species and it was considered a nomen dubium by Goodman et al., 2020; however, this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi	Brooks, D. M., & Bickham, J. W. (2014). New species of Scotophilus (Chiroptera: Vespertilionidae) from sub-Saharan Africa. Occasional Papers, Museum of Texas Tech University, 326, 1-21.|Goodman, S. M., Fratpietro, S., & Tortosa, P. (2020). Insight into the Identity and Origin of Scotophilus borbonicus (E. Geoffroy, 1803). Acta Chiropterologica, 22(1), 41-47.				Kenya|Somalia?|Tanzania?	Africa	Afrotropic	LC	0	0	0	Scotophilus_trujilloi	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotophilus_trujilloi	1005697	23	Trujillo's Yellow Bat	Trujillo's House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Scotophilini	Scotophilus	NA	trujilloi	Brooks & Bickham	0	Scotophilus trujilloi	Brooks, D.M. and Bickham, J.W. 2014. New species of _Scotophilus_ (Chiroptera: Vespertilionidae) from Sub-Saharan Africa. Occasional Papers, Museum of Texas Tech University 326:1-21.	https://www.biodiversitylibrary.org/page/57806524	CM 98038	holotype		"Kenya: Coastal Province, Kwale District, Moana Marine Station, 1 km S, 2 km E Ukunda (4Â°18'S, 39Â°35'E)."	-4.3	39.58333	recently described; the name borbonicus based on specimens reported from Reunion Island likely represents this species and it was considered a nomen dubium by Goodman et al., 2020; however, this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi	Brooks, D. M., & Bickham, J. W. (2014). New species of Scotophilus (Chiroptera: Vespertilionidae) from sub-Saharan Africa. Occasional Papers, Museum of Texas Tech University, 326, 1-21.|Goodman, S. M., Fratpietro, S., & Tortosa, P. (2020). Insight into the Identity and Origin of Scotophilus borbonicus (E. Geoffroy, 1803). Acta Chiropterologica, 22(1), 41-47.				Kenya|Somalia?|Tanzania?	Africa	Afrotropic	LC	0	0	0	Scotophilus_trujilloi	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scotophilus		trujilloi	Brooks & Bickham	2014	0	Occas. Pap. Mus. Texas Tech Univ.	326:16:00	Trujillo's House Bat	None.	Kenya, Coastal Province, Kwale District, Moana Marine Station, 1 km S 2 km E Ukunda	Coastal Province, Kenya.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84466859/84466867/' target='_blank'>Least Concern</a>	In need of revision. Lineage validity confirmed by Demos et al. (2018), but older names (e.g., borbonicus) may have priority. See also Goodman et al. (2020).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotophilus trujilloi; Scotophilus trujilloi; Scotophilus trujilloi; Scotophilus trujilloi; Scotophilus trujilloi; borbonicus; trujilloi; Scotophile de Trujillo; Trujillo-Hausfledermaus; Scotofilode Trujillo; Trujillo's House Bat; Trujillo's Yellow Bat; Trujillo's House Bat; Trujillo's House Bat; S. trujilloi
