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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1545	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Scotophilus tandrefana	Scotophilus tandrefana	Scotophilus tandrefana	Scotophilus tandrefana	Scotophilus tandrefana	Scotophilus tandrefana	Scotophilus tandrefana	Scotophilus tandrefana	Scotophilus tandrefana		[HMW] Scotophilus tandrefana Goodman, Jenkins & Ratrimomanarivo, 2005 , “ Madagascar , Province de Mahajanga ,just outside the limit of the Parc national de Bemaraha, 1.8 km SE from Bekopake and 0.6 km NE from Andadoany, 19°08.454’S , 44°48.732’E , 50 m above sea-level.” Cytochrome-b divergence of 0-6% occurs between S. tandrefana and S. marovaza , indicative of within-species divergence, but it does not provide support for recognition oftandrefana as a valid species; additional research is needed. Monotypic.; [MDD2022] recently described; [IUCN] Goodman et al. (2005) described this species based on three specimens which were collected in 1868, 1869 and 2003 (the latter being designated as the holotype). One of these (the specimen collected in 1869) was later reassessed as S. marovaza (Goodman et al . 2006).; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														trandrefana	Goodman et al. (2005) described this species based on three specimens which were collected in 1868, 1869 and 2003 (the latter being designated as the holotype). One of these (the specimen collected in 1869) was later reassessed as S. marovaza (Goodman et al . 2006).			trandrefana	trandrefana			tandrefana S. M. Goodman, R. K. B. Jenkins, & F. H. Ratrimomanarivo, 2005						N/A																																								NA																											4C3D87E8FF786AC0FA5791F71D01B918	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	886	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF846AC4FA549C211CBFBE2C.xml	Scotophilus tandrefana	Vespertilionidae	Scotophilus	tandrefana	Goodman, Jenkins & Ratrimomanarivo	2005	Scotophile d'Andadoany @fr | \Westmadagaskar Hausfledermaus @de | Scotofilo de Andadoany @es | \Vestern Yellow Bat @en	Scotophilus tandrefana Goodman, Jenkins & Ratrimomanarivo, 2005 , “ Madagascar , Province de Mahajanga ,just outside the limit of the Parc national de Bemaraha, 1.8 km SE from Bekopake and 0.6 km NE from Andadoany, 19°08.454’S , 44°48.732’E , 50 m above sea-level.” Cytochrome-b divergence of 0-6% occurs between S. tandrefana and S. marovaza , indicative of within-species divergence, but it does not provide support for recognition oftandrefana as a valid species; additional research is needed. Monotypic.	W Madagascar , confirmed from two localities near Tsingy de Bemaraha National Park and Sarodrano; possible record from Kirindy Forest.	Head-body ¢.65—- 68 mm , tail 43-46 mm , ear c. 13 mm , hindfoot c. 7 mm , forearm 44-47 mm ; weight c. 14 g . Dorsalfur is long, soft, and dark chocolatebrown, with basal parts distinctly lighter brown. Throat and ventral pelage are shorter and finer, lighter medium brown, with grayish brown bases. Wings and uropatagium are dark brownish black. Ears are black and short. Tragus is long, projecting forward, with slightly complex peduncle, and tip is slightly rounded. Muzzle is pronounced, relatively short, and rounded. Nostrils open slightly antero-laterally and are slightly elongated. Skull is small (greatest skull lengths 16-:9-17- 9 mm ) for Scotophilus ; rostrum is slightly short and broad, without expanded lacrimal processes; braincase is expanded; sagittal and lambdoidal crests are relatively well developed, forming helmet, but less prominent than in adults of most other species of Scotophilus ; posterior palatal extension terminates as acute spine; zygomatic arches are slightly flared; anterior emargination of palate is deep and broad; and pterygoids are expanded posteriorly and wing-shaped. I* is trifid; M' and M? have reduced mesostyle and distorted W-shaped cusp pattern; M* is greatly reduced in size; and trigonid of lower premolarsis larger than talonid.	Holotype from open grassy clearing adjacent to ricefields and disturbed deciduous forest and within 100 m of a limestone outcrop; original natural habitat dry semideciduous forest and capture site ¢. 200 m from relatively intact natural forest formations in Parc national de Bemaraha.	No information.	No information.	A Malagasy Yellow Bat in a flight cage emitted broadband FM/QCF calls at LDC, with frequency containing most energy at 48-2 kHz.	No information.	Classified as Data Deficient on The [UCN Red List.	Goodman, Jenkins & Ratrimomanarivo (2005) | Goodman, Ratrimomanarivo & Randrianandrianina (2006) | Kofoky et al. (2009) | Trujillo et al. (2009)	https://zenodo.org/record/6398392/files/figure.png	274. Malagasy Yellow Bat Scotophilus tandrefana French: Scotophile d'Andadoany / German: \WestmadagaskarHausfledermaus / Spanish: Scotofilo de Andadoany Other common names: \Vestern Yellow Bat Taxonomy. Scotophilus tandrefana Goodman, Jenkins & Ratrimomanarivo, 2005 , “ Madagascar , Province de Mahajanga ,just outside the limit of the Parc national de Bemaraha, 1.8 km SE from Bekopake and 0.6 km NE from Andadoany, 19°08.454’S , 44°48.732’E , 50 m above sea-level.” Cytochrome-b divergence of 0-6% occurs between S. tandrefana and S. marovaza , indicative of within-species divergence, but it does not provide support for recognition oftandrefana as a valid species; additional research is needed. Monotypic. Distribution. W Madagascar , confirmed from two localities near Tsingy de Bemaraha National Park and Sarodrano; possible record from Kirindy Forest. Descriptive notes. Head-body ¢.65—- 68 mm , tail 43-46 mm , ear c. 13 mm , hindfoot c. 7 mm , forearm 44-47 mm ; weight c. 14 g . Dorsalfur is long, soft, and dark chocolatebrown, with basal parts distinctly lighter brown. Throat and ventral pelage are shorter and finer, lighter medium brown, with grayish brown bases. Wings and uropatagium are dark brownish black. Ears are black and short. Tragus is long, projecting forward, with slightly complex peduncle, and tip is slightly rounded. Muzzle is pronounced, relatively short, and rounded. Nostrils open slightly antero-laterally and are slightly elongated. Skull is small (greatest skull lengths 16-:9-17- 9 mm ) for Scotophilus ; rostrum is slightly short and broad, without expanded lacrimal processes; braincase is expanded; sagittal and lambdoidal crests are relatively well developed, forming helmet, but less prominent than in adults of most other species of Scotophilus ; posterior palatal extension terminates as acute spine; zygomatic arches are slightly flared; anterior emargination of palate is deep and broad; and pterygoids are expanded posteriorly and wing-shaped. I* is trifid; M' and M? have reduced mesostyle and distorted W-shaped cusp pattern; M* is greatly reduced in size; and trigonid of lower premolarsis larger than talonid. Habitat. Holotype from open grassy clearing adjacent to ricefields and disturbed deciduous forest and within 100 m of a limestone outcrop; original natural habitat dry semideciduous forest and capture site ¢. 200 m from relatively intact natural forest formations in Parc national de Bemaraha. Food and Feeding. No information. Breeding. No information. Activity patterns. A Malagasy Yellow Bat in a flight cage emitted broadband FM/QCF calls at LDC, with frequency containing most energy at 48-2 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The [UCN Red List. Bibliography. Goodman, Jenkins & Ratrimomanarivo (2005), Goodman, Ratrimomanarivo & Randrianandrianina (2006), Kofoky et al. (2009), Trujillo et al. (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scotophilus tandrefana	Scotophilus		tandrefana	Goodman, Jenkins & Ratrimomanarivo	2005	0	Zoosystema	27(4): 875	Western House Bat	None.	outside Parc National de Bemaraha, Province de Mahajanga, Madagascar	W Madagascar	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scotophilus tandrefana	23	Malagasy Yellow Bat	Western Yellow Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	SCOTOPHILINI	Scotophilus	NA	tandrefana	Goodman, Jenkins, & Ratrimomanarivo	2005	0	Scotophilus_tandrefana	Goodman, S. M., Jenkins, R. K., & Ratrimomanarivo, F. H. (2005). A review of the genus Scotophilus (Mammalia, Chiroptera, Vespertilionidae) on Madagascar, with the description of a new species. Zoosystema, 27(4), 875.	https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/z2005n4a8.pdf	UADBA 46923		"Madagascar, Province de Mahajanga, just outside the limit of the Parc national de Bemaraha, 1.8 km SE from Bekopake and 0.6 km NE from Andadoany, 19°08.454'S, 44°48.732'E, 50 m above sea-level."	-19.14	44.812	trandrefana Goodman, Jenkins, & Ratrimomanarivo, 2005	recently described	Goodman, S. M., Jenkins, R. K., & Ratrimomanarivo, F. H. (2005). A review of the genus Scotophilus (Mammalia, Chiroptera, Vespertilionidae) on Madagascar, with the description of a new species. Zoosystema, 27(4), 867.	Madagascar	Africa	Afrotropic	DD	0	0	0	Scotophilus_tandrefana	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136675	Scotophilus tandrefana	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scotophilus	tandrefana	Goodman, Jenkins &; Ratrimomanarivo, 2005	Goodman et al. (2005) described this species based on three specimens which were collected in 1868, 1869 and 2003 (the latter being designated as the holotype). One of these (the specimen collected in 1869) was later reassessed as S. marovaza (Goodman et al . 2006).	20000000	Scotophilus tandrefana	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Data Deficient as the species is known from only two specimens (one collected as recently as 2003) and there is no information on its population status or threats.	This species is not considered to be a strict forest-dwelling species or known to be synanthropic (Goodman et al. 2005). There is no information on roosting or foraging habitats, but it appears to be associated with dry deciduous forest (Goodman et al . 2005).	The major threats to this species are not known.	There are no data on the population of this species and all existing information is based on two specimens that were collected in 1868 and 2003 (Goodman et al. 2005, and see Goodman et al. 2006).	Unknown	There is specimen evidence of this species from two localities: near Parc National Tsingy de Bemaraha in the west and Sarodrano in the south (Goodman et al. 2005). Photographs and forearm measurements of an individual from Kirindy (CFPF) forest in the west represents the third known locality for this species (Goodman et al . 2005).		Terrestrial	This species has been recorded from near Parc National Tsingy de Bemaraha and the managed forest of Kirindy CFPF (Goodman et al. 2005), but evidence is lacking that it actually roosts or feeds in these forests.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scotophilus		tandrefana	Goodman, Jenkins & Ratrimomanarivo	2005	0	Zoosystema	27(4): 875	Western House Bat	None.	outside Parc National de Bemaraha, Province de Mahajanga, Madagascar	W Madagascar	Not listed.	Data Deficient		Scotophilus tandrefana	1005696	23	Malagasy Yellow Bat	Western Yellow Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	SCOTOPHILINI	Scotophilus	NA	tandrefana	Goodman, Jenkins, & Ratrimomanarivo	2005	0	Scotophilus_tandrefana	Goodman, S. M., Jenkins, R. K., & Ratrimomanarivo, F. H. (2005). A review of the genus Scotophilus (Mammalia, Chiroptera, Vespertilionidae) on Madagascar, with the description of a new species. Zoosystema, 27(4), 875.	https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/z2005n4a8.pdf	UADBA 46923		"Madagascar, Province de Mahajanga, just outside the limit of the Parc national de Bemaraha, 1.8 km SE from Bekopake and 0.6 km NE from Andadoany, 19°08.454'S, 44°48.732'E, 50 m above sea-level."	-19.1409	44.8122	trandrefana Goodman, Jenkins, & Ratrimomanarivo, 2005	recently described	Goodman, S. M., Jenkins, R. K., & Ratrimomanarivo, F. H. (2005). A review of the genus Scotophilus (Mammalia, Chiroptera, Vespertilionidae) on Madagascar, with the description of a new species. Zoosystema, 27(4), 867.				Madagascar	Africa	Afrotropic	DD	0	0	0	Scotophilus_tandrefana	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotophilus_tandrefana	1005696	23	Malagasy Yellow Bat	Western Yellow Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Scotophilini	Scotophilus	NA	tandrefana	S. M. Goodman, R. K. B. Jenkins, & F. H. Ratrimomanarivo	0	Scotophilus tandrefana	Goodman, S.M., Jenkins, R.K.B. and Ratrimomanarivo, F.H. 2005. A review of the genus _Scotophilus_ (Mammalia, Chiroptera, Vespertilionidae) on Madagascar, with the description of a new species. Zoosystema 27(4):867-882.	https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/z2005n4a8.pdf	UAnt DBA 46923	holotype		"Madagascar, Province de Mahajanga, just outside the limit of the Parc national de Bemaraha, 1.8 km SE from Bekopake and 0.6 km NE from Andadoany, 19°08.454'S, 44°48.732'E, 50 m above sea-level."	-19.1409	44.8122	recently described	Goodman, S. M., Jenkins, R. K., & Ratrimomanarivo, F. H. (2005). A review of the genus Scotophilus (Mammalia, Chiroptera, Vespertilionidae) on Madagascar, with the description of a new species. Zoosystema, 27(4), 867.				Madagascar	Africa	Afrotropic	DD	0	0	0	Scotophilus_tandrefana	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scotophilus		tandrefana	Goodman, Jenkins & Ratrimomanarivo	2005	0	Zoosystema	27(4): 875	Western House Bat	None.	outside Parc National de Bemaraha, Province de Mahajanga, Madagascar	W Madagascar	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136675/22039268/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotophilus tandrefana; Scotophilus tandrefana; Scotophilus tandrefana; Scotophilus tandrefana; Scotophilus tandrefana; trandrefana; Scotophile d'Andadoany; \Westmadagaskar Hausfledermaus; Scotofilo de Andadoany; \Vestern Yellow Bat; Malagasy Yellow Bat; Western Yellow Bat; Western House Bat; S. tandrefana