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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1543	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Scotophilus nux	Scotophilus nux	Scotophilus nux	Scotophilus leucogaster nux	Scotophilus nux	Scotophilus nux	Scotophilus nux	Scotophilus nux	Scotophilus nux	Scotophilus nux	Scotophilus nux	Scotophilus nux	Scotophilus nux	Scotophilus nux		[MSW2] S. nux has been most often recognized as a subspecies of nigrita, when nigrita was used for the species now called dinganii; see Allen (1939), Rosevear (1965), and Hayman and Hill (1971:50). Koopman et al. (1978:4-5) listed it as a subspecies of leucogaster.; [MSW3] Often treated as a subspecies of dinganii or leucogaster (or nigrita, when that name was misapplied to the former species), see G. M. Allen (1939), Rosevear (1965), and Hayman and Hill (1971), Koopman et al. (1978) and Koopman (1994). However, nux appears to be distinct from all of the above species; see C. B. Robbins et al. (1985).; [HMW] Scotophilus nigrita nux Thomas, 1904 , “Efulen, Cameroons [= Cameroon ].” Scotophilus nux has been treated as a subspecies of nigrita (when nigrita was used for the species now called S. dinganii ) or a subspecies of S. leucogaster . Based on multivariate analyses of forearm and cranial measurements, it is treated as a distinct species. Monotypic.; [batnames2022] Often treated as a subspecies of dinganii or leucogaster (or nigrita , when that name was misapplied to the formerspecies), see G. M. Allen (1939), Rosevear (1965), and Hayman and Hill (1971), Koopman et al. (1978) and Koopman (1994). However, nux appears to be distinct from all of the above species; see C. B. Robbins et al. (1985).; [IUCN] Entire genus needs revision.; [batnames2023] Often treated as a subspecies of dinganii or leucogaster (or nigrita , when that name was misapplied to the formerspecies), see G. M. Allen (1939), Rosevear (1965), and Hayman and Hill (1971), Koopman et al. (1978) and Koopman (1994). However, nux appears to be distinct from all of the above species; see C. B. Robbins et al. (1985).; [batnames2025_1.7] Often treated as a subspecies of dinganii or leucogaster (or nigrita, when that name was misapplied to the formerspecies), see G. M. Allen (1939), Rosevear (1965), and Hayman and Hill (1971), Koopman et al. (1978) and Koopman (1994). However, nuxappears to be distinct from all of the above species; see C. B. Robbins et al. (1985).														nux	Entire genus needs revision.			nux	nux			nux O. Thomas, 1904					Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Scotophilus nux	Cameroon, Efulen.	Thomas	1904	Ann. Mag. Nat. Hist., ser. 7, 13:208.			Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Sierra Leone – Kenya; ref. 4.138, 139	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1904	Ann. Mag. Nat. Hist., ser. 7, 13:208.	S. nux has been most often recognized as a subspecies of nigrita, when nigrita was used for the species now called dinganii; see Allen (1939), Rosevear (1965), and Hayman and Hill (1971:50). Koopman et al. (1978:4-5) listed it as a subspecies of leucogaster.	Sierra Leone to Kenya.	Cameroon, Efulen.																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nycticeiini	Scotophilus nux	Scotophilus		nux	Thomas		1904		Ann. Mag. Nat. Hist., ser. 7	13		208		Nut-colored House Bat	Cameroon, Efulen.	High forest zones from Sierra Leone to Kenya.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Often treated as a subspecies of dinganii or leucogaster (or nigrita, when that name was misapplied to the former species), see G. M. Allen (1939), Rosevear (1965), and Hayman and Hill (1971), Koopman et al. (1978) and Koopman (1994). However, nux appears to be distinct from all of the above species; see C. B. Robbins et al. (1985).	4C3D87E8FF7A6AC5FF4E90D81DA2B755	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	886	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF846A3BFA5E956F198FB11D.xml	Scotophilus nux	Vespertilionidae	Scotophilus	nux	Thomas	1904	Scotophile noisette @fr | Nussfarbene Hausfledermaus @de | Scotofilocastano @es | Dark House Bat @en | Nut-colored House Bat @en | Nux Yellow House Bat @en	Scotophilus nigrita nux Thomas, 1904 , “Efulen, Cameroons [= Cameroon ].” Scotophilus nux has been treated as a subspecies of nigrita (when nigrita was used for the species now called S. dinganii ) or a subspecies of S. leucogaster . Based on multivariate analyses of forearm and cranial measurements, it is treated as a distinct species. Monotypic.	Patchily recorded in Sierra Leone , Guinea , Liberia , Ivory Coast , Ghana , Nigeria , Cameroon , DR Congo , Uganda , and Kenya ; possibly Equatorial Guinea and Gabon .	Head—body ¢. 67-89 mm , tail 44-54 mm , ear 15-19 mm , hindfoot 13-15 mm , forearm 53-61 mm ; weight 25-37 g . Pelage is smooth, sleek, and glossy. Dorsum is rusty brown, dark rusty brown, or blackish brown; hairs are unicolored. Mid-dorsal hairs are 6-7 mm . Venteris slightly paler and dark brown to reddish orange. Wings and uropatagium are blackish brown. Ears are dark brown, short, and separated; inner margin is strongly convex, with lobe at base; and outer margin is fairly straight, with semicircular fleshy antitragus. Tragus tapers to bluntly rounded tip, with anterior margin concave. Skull has well-developed sagittal crest and occipital helmet. Dorsal profile ofskull is slightly concave at rostrumand slopes gradually upward to occiput. Jaws and dentition are robust. I* is unicuspid; M' and M* have concave surfaces and indistinct ridges and appear worn; and M? is very short and has only two ridges. Chromosomal complement has 2n = 36 and FNa = 50 in Cameroon .	[Lowland rainforest and clearings.	The Nut-colored Yellow Bat apparently forages over water.	Two Nut-colored Yellow Bats with one fetus each were recorded in March in DR Congo .	The Nut-colored Yellow Bat roosts in hollow trees and in small groups in roofs of houses. In western Uganda , echolocation calls had minimum frequency of 40-7 kHz, maximum frequency of 54-1 kHz, characteristic frequency of 43 kHz, frequency of the knee of 45-1 kHz, and duration of 2 milliseconds. In Gabon , mean minimum frequency was 38-5 kHz (37-8-40), mean maximum frequency was 59-5 kHz (52-5-75-8), mean dominant frequency was 44-5 kHz (41:1-47-4), mean duration was 1-9 milliseconds (1-6-2-5), and mean interpulse interval was 57-1 milliseconds (16-2-108-6).	No information.	Classified as Least Concern on The IUCN Red Lust.	Monadjem et al. (2011) | Peereboom & Van Lieshout (2015) | Robbins et al. (1985) | Simmons (2005) | Van Cakenberghe & Happold (20139)	https://zenodo.org/record/6398390/files/figure.png	273. Nut-colored Yellow Bat Scotophilus nux French: Scotophile noisette / German: Nussfarbene Hausfledermaus / Spanish: Scotofilo castano Other common names: Dark House Bat , Nut-colored House Bat , Nux Yellow House Bat Taxonomy. Scotophilus nigrita nux Thomas, 1904 , “Efulen, Cameroons [= Cameroon ].” Scotophilus nux has been treated as a subspecies of nigrita (when nigrita was used for the species now called S. dinganii ) or a subspecies of S. leucogaster . Based on multivariate analyses of forearm and cranial measurements, it is treated as a distinct species. Monotypic. Distribution. Patchily recorded in Sierra Leone , Guinea , Liberia , Ivory Coast , Ghana , Nigeria , Cameroon , DR Congo , Uganda , and Kenya ; possibly Equatorial Guinea and Gabon . Descriptive notes. Head—body ¢. 67-89 mm , tail 44-54 mm , ear 15-19 mm , hindfoot 13-15 mm , forearm 53-61 mm ; weight 25-37 g . Pelage is smooth, sleek, and glossy. Dorsum is rusty brown, dark rusty brown, or blackish brown; hairs are unicolored. Mid-dorsal hairs are 6-7 mm . Venteris slightly paler and dark brown to reddish orange. Wings and uropatagium are blackish brown. Ears are dark brown, short, and separated; inner margin is strongly convex, with lobe at base; and outer margin is fairly straight, with semicircular fleshy antitragus. Tragus tapers to bluntly rounded tip, with anterior margin concave. Skull has well-developed sagittal crest and occipital helmet. Dorsal profile ofskull is slightly concave at rostrumand slopes gradually upward to occiput. Jaws and dentition are robust. I* is unicuspid; M' and M* have concave surfaces and indistinct ridges and appear worn; and M? is very short and has only two ridges. Chromosomal complement has 2n = 36 and FNa = 50 in Cameroon . Habitat. [Lowland rainforest and clearings. Food and Feeding. The Nut-colored Yellow Bat apparently forages over water. Breeding. Two Nut-colored Yellow Bats with one fetus each were recorded in March in DR Congo . Activity patterns. The Nut-colored Yellow Bat roosts in hollow trees and in small groups in roofs of houses. In western Uganda , echolocation calls had minimum frequency of 40-7 kHz, maximum frequency of 54-1 kHz, characteristic frequency of 43 kHz, frequency of the knee of 45-1 kHz, and duration of 2 milliseconds. In Gabon , mean minimum frequency was 38-5 kHz (37-8-40), mean maximum frequency was 59-5 kHz (52-5-75-8), mean dominant frequency was 44-5 kHz (41:1-47-4), mean duration was 1-9 milliseconds (1-6-2-5), and mean interpulse interval was 57-1 milliseconds (16-2-108-6). Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Monadjem et al. (2011), Peereboom & Van Lieshout (2015), Robbins et al. (1985), Simmons (2005), Van Cakenberghe & Happold (20139).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scotophilus nux	Scotophilus		nux	Thomas	1904	0	Ann. Mag. Nat. Hist.	ser. 7, 13: 208	Nut-colored House Bat	None.	Cameroon, Efulen.	High forest zones from Sierra Leone to Kenya.	Not listed.	Least Concern	Often treated as a subspecies of dinganii or leucogaster (or nigrita , when that name was misapplied to the formerspecies), see G. M. Allen (1939), Rosevear (1965), and Hayman and Hill (1971), Koopman et al. (1978) and Koopman (1994). However, nux appears to be distinct from all of the above species; see C. B. Robbins et al. (1985).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scotophilus nux	23	Nut-colored Yellow Bat	Dark House Bat|Nut-colored House Bat|Nux Yellow House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	SCOTOPHILINI	Scotophilus	NA	nux	O. Thomas	1904	0	Scotophilus_nigrita_nux	Thomas, O. (1904). New bats from British East Africa collected by Mrs. Hinde, and from the Cameroons by Mr. G. L. Bates. Annals and Magazine of Natural History, ser. 7, 13, 208.	https://www.biodiversitylibrary.org/item/54611#page/228/mode/1up	BM 1903.2.4.5		"Efulen, Cameroons [= Cameroon]."			nux O. Thomas, 1904	NA	NA	Sierra Leone|Guinea|Liberia|Côte d'Ivoire|Ghana|Nigeria|Cameroon|Democratic Republic of the Congo|Uganda|Kenya|Equatorial Guinea?|Gabon?	Africa	Afrotropic	LC	0	0	0	Scotophilus_nux	0	sciname match	Scotophilus_nux	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	20071	Scotophilus nux	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scotophilus	nux	Thomas, 1904	Entire genus needs revision.	20000000	Scotophilus nux	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species appears to be associated with tropical lowland forest (Robbins et al. 1985; Happold, 1987; Grubb et al. 1998). Little additional information is available on the natural history of this species.	There appear to be no major threats to this species. Some populations may be impacted by general deforestation, however, this needs to be confirmed.	Little information is available on the population abundance or size of this species.	Unknown	This sub-Saharan species has been patchily recorded from West Africa (Sierra Leone, Guinea, Liberia Côte d'Ivoire, Ghana and Nigeria), Central Africa (Cameroon and Democratic Republic of the Congo) and East Africa (Uganda and Kenya), and is believed to likely occur more widely than is currently known. It is possibly present in Equatorial Guinea, however, this needs to be confirmed.		Terrestrial	There appear to be no direct conservation measures in place. It is not known if the species is present in any protected areas.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scotophilus		nux	Thomas	1904	0	Ann. Mag. Nat. Hist.	ser. 7, 13: 208	Nut-colored House Bat	None.	Cameroon, Efulen.	High forest zones from Sierra Leone to Kenya.	Not listed.	Least Concern	Often treated as a subspecies of dinganii or leucogaster (or nigrita , when that name was misapplied to the formerspecies), see G. M. Allen (1939), Rosevear (1965), and Hayman and Hill (1971), Koopman et al. (1978) and Koopman (1994). However, nux appears to be distinct from all of the above species; see C. B. Robbins et al. (1985).	Scotophilus nux	1005694	23	Nut-colored Yellow Bat	Dark House Bat|Nut-colored House Bat|Nux Yellow House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	SCOTOPHILINI	Scotophilus	NA	nux	O. Thomas	1904	0	Scotophilus_nigrita_nux	Thomas, O. (1904). New bats from British East Africa collected by Mrs. Hinde, and from the Cameroons by Mr. G. L. Bates. Annals and Magazine of Natural History, ser. 7, 13, 208.	https://www.biodiversitylibrary.org/item/54611#page/228/mode/1up	BM 1903.2.4.5		"Efulen, Cameroons [= Cameroon]."			nux O. Thomas, 1904	NA	NA				Sierra Leone|Guinea|Liberia|Côte d'Ivoire|Ghana|Nigeria|Cameroon|Democratic Republic of the Congo|Uganda|Kenya|Equatorial Guinea?|Gabon?	Africa	Afrotropic	LC	0	0	0	Scotophilus_nux	0	sciname match	Scotophilus_nux	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotophilus_nux	1005694	23	Nut-colored Yellow Bat	Dark House Bat|Nut-colored House Bat|Nux Yellow House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Scotophilini	Scotophilus	NA	nux	O. Thomas	0	Scotophilus nigrita nux	Thomas, O. 1904-03-01. New bats from British East Africa collected by Mrs. Hinde, and from the Cameroons by Mr. G. L. Bates. Annals and Magazine of Natural History (7)13(75):206-210.	https://www.biodiversitylibrary.org/page/16108294	BMNH:Mamm:1903.2.4.5	holotype	https://data.nhm.ac.uk/object/9969c604-fa88-442c-80b5-d1db6c25a269	"Efulen, Cameroons [= Cameroon]."	2.766667	10.7	NA	NA				Sierra Leone|Guinea|Liberia|Cote d'Ivoire|Ghana|Nigeria|Cameroon|Democratic Republic of the Congo|Uganda|Kenya|Equatorial Guinea?|Gabon?	Africa	Afrotropic	LC	0	0	0	Scotophilus_nux	0	sciname match	Scotophilus_nux	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scotophilus		nux	Thomas	1904	0	Ann. Mag. Nat. Hist.	ser. 7, 13: 208	Nut-colored House Bat	None.	Cameroon, Efulen.	High forest zones from Sierra Leone to Kenya.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/20071/22032438/' target='_blank'>Least Concern</a>	Often treated as a subspecies of dinganii or leucogaster (or nigrita, when that name was misapplied to the formerspecies), see G. M. Allen (1939), Rosevear (1965), and Hayman and Hill (1971), Koopman et al. (1978) and Koopman (1994). However, nuxappears to be distinct from all of the above species; see C. B. Robbins et al. (1985).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotophilus nux; Scotophilus nux; Scotophilus nux; Scotophilus nux; Scotophilus nux; Scotophilus nux; nux; Scotophile noisette; Nussfarbene Hausfledermaus; Scotofilocastano; Dark House Bat; Nut-colored House Bat; Nux Yellow House Bat; Nut-colored Yellow Bat; Dark House Bat; Nut-colored House Bat; Nux Yellow House Bat; Nut-colored House Bat; Nut-colored House Bat; S. nux