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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L154	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus	Choeroniscus periosus		[MSW3] Includes ponsi; see Koopman (1994).; [HMW] Choeroniscus periosus Handley, 1966 , “Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca ], Colombia .” This species is monotypic.; [batnames2022] Includes ponsi ; see Koopman (1994).; [IUCN] Simmons (2005) used the name C. ponsi which may have lead to confusion about occurrences in Venezuela, thus more work is necessary on this species.; [batnames2023] Includes ponsi ; see Koopman (1994).; [batnames2025_1.7] Includes ponsi; see Koopman (1994).							periosus, ponsi	periosus, ponsi				periosus, ponsi 		periosus	Simmons (2005) used the name C. ponsi which may have lead to confusion about occurrences in Venezuela, thus more work is necessary on this species.	periosus, ponsi 		periosus 	periosus 	periosus, ponsi		periosus Handley, 1966		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		W coast Colombia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Choeroniscus periosus	Colombia, Valle, 27 km S. Buenaventura, Rio Raposo.	Handley	1966	Proc. Biol. Soc. Wash., 79:84.	Distribution: Known only from western Colombia and northwestern Venezuela.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		W Colombia	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Handley	1966	Proc. Biol. Soc. Washington, 79:84.		W Colombia, W Ecuador.	Colombia, Valle, 27 km S Buenaventura, Rio Raposo.		HANDLEY	1966	Size relatively large (forearm length, 40-42 mm; condylobasal length, 29-30 mm). Postero-lateral margin of palate unnotched. Cranium relatively low. Rostrum relatively long and robust. Pterygoid inflation pronounced.	Distribution: Known only from western Colombia and northwestern Venezuela.	Two subspecies are here recognized:	C. p. periosus (western Colombia), C. p. ponsi (northwestern Venezuela). C. p. ponsi was originally described as a subspecies of Choeronycteris but is best placed here.	83	species	C. periosus	HANDLEY	1966	Choeroniscus	genus	Choeroniscus periosus				Size relatively large (forearm length, 40-42 mm; condylobasal length, 29-30 mm). Postero-lateral margin of palate unnotched. Cranium relatively low. Rostrum relatively long and robust. Pterygoid inflation pronounced.	Two subspecies are here recognized:		4. C. periosus HANDLEY 1966.	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Glossophaginae	Glossophagini	Choeroniscus periosus	Choeroniscus		periosus	Handley		1966		Proc. Biol. Soc. Wash.	79		84		Greater Long-tongued Bat	Colombia, Valle, 27 km S Buenaventura, Río Raposo.	NW Venezuela, W Colombia, W Ecuador.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.	ponsi Pirlot, 1967.	Includes ponsi; see Koopman (1994).	03A687BCFF96FF961399F398F9B5F7B9	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	523	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF96FF961399F398F9B5F7B9.xml	Choeroniscus periosus	Phyllostomidae	Choeroniscus	periosus	Handley	1966	Grande Oxyrhine @fr | Gro Re Langschwanz-Blumenfledermaus @de | Coeroniscogrande @es | Handley's Long-tailed Bat @en	Choeroniscus periosus Handley, 1966 , “Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca ], Colombia .” This species is monotypic.	Pacific lowlands of W Colombia (Choco S to Narino departments) and NW Ecuador ( Esmeraldas Province ).	Head-body 70 mm , tail 8= 10 mm , ear 13-17 mm, hindfoot 7-11 mm, 40-1-41-2 mm; weight 14 g . Dorsal fur of Greater Long-tailed Bat is dark brown, with bicolored hairs having orange-brown bases and dark brown tips. Underparts are slightly lighter than dorsum. Rostrum is long, with conspicuous vibrissae on muzzle and chin. Lower lip is notched. Ears are round and short, and antitragus is well defined. Noseleaf is blackish and relatively small, with no central rib. Wing membranes and uropatagium are blackish. Uropatagium is naked and long, reaching distal part oftibia. Calcar is smaller than foot. Tail is short and completely inserted in uropatagium. Rostrum is longer than braincase. Postorbital processes are absent. Basisphenoid pits are moderately deep, separated by wide septum. Zygomatic arches are incomplete. Posterolateral margin of palate is unnotched. Hamular process of pterygoid is inflated and touches auditory bulla. Upperincisors are small and separated as pairs in each side of maxillae. P, contacts C,.	Humid forests of the Chocé ecoregion from near sea level to elevations of ¢. 400 m . In Ecuador , a Greater Long-tailed Bat was captured in a marshy landscape dominated by Heliconia ( Heliconiaceae ).	Skull morphology and dentition of the Greater Long-tailed Bat suggest it is a nectarivore, similar to congeners.	A lactating Greater Long-tailed Bat was captured in September in Ecuador .	No information.	No information.	Classified as Vulnerable on The IUCN Red List. The Greater Long-tailed Bat is confined to a small area and its habitat, the Chocoan humid forests, suffers intense and large-scale deforestation.	Albuja & Mena (1999) | Griffiths & Gardner (2008a) | Handley (1966a) | Saavedra-Rodriguez & Rojas-Diaz (2011)	https://zenodo.org/record/6458754/files/figure.png	76. Greater Long-tailed Bat Choeroniscus periosus French: Grande Oxyrhine / German: GroRe Langschwanz-Blumenfledermaus / Spanish: Coeronisco grande Other common names: Handley's Long-tailed Bat Movements, Home range and Social organization. Mexican Long-tongued Bats apparently have latitudinal migrations. They have been found flying near columnar cacti fruits with long-nosed bats ( Leptonycteris spp.). In the northern part ofits distribution, Mexican Long-tongued Bats share roosts with Cave Myotis ( Myotis velifer), Big Brown Bats ( Eptesicus fuscus), Townsend's Big-eared Bats ( Plecotus townsendir), and Mexican Free-tailed Bats (Tadarida brasiliensis ). In the southern part ofits distribution, it shares roosts with other long-tongued bats ( Glossophaga spp.). Taxonomy. Choeroniscus periosus Handley, 1966 , “Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca ], Colombia .” This species is monotypic. Distribution. Pacific lowlands of W Colombia (Choco S to Narino departments) and NW Ecuador ( Esmeraldas Province ). Descriptive notes. Head-body 70 mm , tail 8= 10 mm , ear 13-17 mm, hindfoot 7-11 mm, 40-1-41-2 mm; weight 14 g . Dorsal fur of Greater Long-tailed Bat is dark brown, with bicolored hairs having orange-brown bases and dark brown tips. Underparts are slightly lighter than dorsum. Rostrum is long, with conspicuous vibrissae on muzzle and chin. Lower lip is notched. Ears are round and short, and antitragus is well defined. Noseleaf is blackish and relatively small, with no central rib. Wing membranes and uropatagium are blackish. Uropatagium is naked and long, reaching distal part oftibia. Calcar is smaller than foot. Tail is short and completely inserted in uropatagium. Rostrum is longer than braincase. Postorbital processes are absent. Basisphenoid pits are moderately deep, separated by wide septum. Zygomatic arches are incomplete. Posterolateral margin of palate is unnotched. Hamular process of pterygoid is inflated and touches auditory bulla. Upperincisors are small and separated as pairs in each side of maxillae. P, contacts C,. Habitat. Humid forests of the Chocé ecoregion from near sea level to elevations of ¢. 400 m . In Ecuador , a Greater Long-tailed Bat was captured in a marshy landscape dominated by Heliconia ( Heliconiaceae ). Food and Feeding. Skull morphology and dentition of the Greater Long-tailed Bat suggest it is a nectarivore, similar to congeners. Breeding. A lactating Greater Long-tailed Bat was captured in September in Ecuador . Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Vulnerable on The IUCN Red List. The Greater Long-tailed Bat is confined to a small area and its habitat, the Chocoan humid forests, suffers intense and large-scale deforestation. Bibliography. Albuja & Mena (1999), Griffiths & Gardner (2008a), Handley (1966a), Saavedra-Rodriguez & Rojas-Diaz (2011).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Choeroniscus periosus	Choeroniscus		periosus	Handley	1966	0	Proc. Biol. Soc. Wash.	80:24:00	Greater Long-tongued Bat	 <b>ponsi </b> Pirlot, 1967.	Colombia, Valle, 27 km S Buenaventura, Río Raposo.	NW Venezuela, W Colombia, W Ecuador.	Not listed.	Vulnerable	Includes ponsi ; see Koopman (1994).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Choeroniscus periosus	23	Greater Long-tailed Bat	Handley's Long-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	GLOSSOPHAGINAE	CHOERONYCTERINI	Choeroniscus	NA	periosus	Handley	1966	0	Choeroniscus_periosus	Handley, C. O., Jr. (1966). Descriptions of New Bats (Choeroniscus and Rhinophylla) from Colombia. Proceedings of the Biological Society of Washington, 79, 84.	https://www.biodiversitylibrary.org/page/34562753#page/440/mode/1up	USNM 344918		"Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca], Colombia."			periosus Handley, 1966	NA	NA	Colombia|Ecuador	South America	Neotropic	VU	0	0	0	Choeroniscus_periosus	0	sciname match	Choeroniscus_periosus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	4775	Choeroniscus periosus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Choeroniscus	periosus	Handley, 1966	Simmons (2005) used the name C. ponsi which may have lead to confusion about occurrences in Venezuela, thus more work is necessary on this species.	20000000	Choeroniscus periosus	Vulnerable	A3c	2015	2014-07-31 00:00:00 UTC	3.1	English	This species is listed as Vulnerable because it is suspected that a population decline of ;&gt;30% will be met over the next three generations (with a generation length of six years; Pacifici et al. 2013), due to the rapid loss of habitat to agriculture, logging and illicit crops. This species is poorly known, restricted to forest habitats, and very susceptible to habitat conversion. This species needs taxonomic investigation and should be reviewed following an evaluation.	This species is limited to primary forest. Little is known of the behaviour of this species, but it is probably similar to other members of the genus. It feeds on nectar and small insects. The generation length is six years (Pacifici et al. 2013).	The habitat where this species occurs is rapidly being destroyed for agriculture, logging, road construction and illicit crops (Tirira 2011).	This species is poorly known and rare (Tirira 2007).	Unknown	This species if found in southwestern Colombia and northwestern Ecuador. It seems to be a Chocoan endemic and thus records from elsewhere are likely erroneous. It occurs from zero to 500 m asl in humid habitats.	This species is not used.	Terrestrial	It occurs in very few protected areas (e.g. Parque Nacional Natural Los Katíos, Colombia) between which the original habitat is rapidly being destroyed. Existing protected areas are not enforced and may offer little protection. ;This species needs taxonomic investigation	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Choeroniscus		periosus	Handley	1966	0	Proc. Biol. Soc. Wash.	80:24:00	Greater Long-tongued Bat	 <b>ponsi </b> Pirlot, 1967.	Colombia, Valle, 27 km S Buenaventura, Río Raposo.	NW Venezuela, W Colombia, W Ecuador.	Not listed.	Vulnerable	Includes ponsi ; see Koopman (1994).	Choeroniscus periosus	1004901	23	Greater Long-tailed Bat	Handley's Long-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	GLOSSOPHAGINAE	CHOERONYCTERINI	Choeroniscus	NA	periosus	Handley	1966	0	Choeroniscus_periosus	Handley, C. O., Jr. (1966). Descriptions of New Bats (Choeroniscus and Rhinophylla) from Colombia. Proceedings of the Biological Society of Washington, 79, 84.	https://www.biodiversitylibrary.org/page/34562753#page/440/mode/1up	USNM 344918		"Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca], Colombia."			periosus Handley, 1966	NA	NA				Colombia|Ecuador	South America	Neotropic	VU	0	0	0	Choeroniscus_periosus	0	sciname match	Choeroniscus_periosus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Choeroniscus_periosus	1004901	23	Greater Long-tailed Bat	Handley's Long-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Glossophaginae	Choeronycterini	Choeroniscus	NA	periosus	Handley	0	Choeroniscus periosus	Handley, C.O., Jr. 1966-05-23. Descriptions of new bats (_Choeroniscus_ and _Rhinophylla_) from Colombia. Proceedings of the Biological Society of Washington 79:83-88.	https://www.biodiversitylibrary.org/page/34562754	USNM:MAMM:344918	holotype	http://n2t.net/ark:/65665/38da44a99-6b19-49ac-9e14-db9e744f1266	"Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca], Colombia."			NA	NA				Colombia|Ecuador	South America	Neotropic	VU	0	0	0	Choeroniscus_periosus	0	sciname match	Choeroniscus_periosus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Choeroniscus		periosus	Handley	1966	0	Proc. Biol. Soc. Wash.	80:24:00	Greater Long-tongued Bat	ponsi Pirlot, 1967.	Colombia, Valle, 27 km S Buenaventura, Río Raposo.	NW Venezuela, W Colombia, W Ecuador.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/4775/22042360/' target='_blank'>Vulnerable</a>	Includes ponsi; see Koopman (1994).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Choeronycterina	Choeroniscus periosus; Choeroniscus periosus; Choeroniscus periosus; Choeroniscus periosus; Choeroniscus periosus; Choeroniscus periosus; periosus; ponsi; ponsi; periosus; Grande Oxyrhine; Gro Re Langschwanz-Blumenfledermaus; Coeroniscogrande; Handley's Long-tailed Bat; Greater Long-tailed Bat; Handley's Long-tailed Bat; Greater Long-tongued Bat; Greater Long-tongued Bat; C. periosus