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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1529	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	nomen dubium for Scotophilus trujilloi	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus	Scotophilus borbonicus		[MSW2] Included equivocally in leucogaster by Hayman and Hill (1971:50-51); but see Koopman (1975:414-416). Hill (1980b) considered African viridis and damarensis, and possibly leucogaster to be conspecific with borbonicus, but C. B. Robbins et al. (1985) rejected any affinity with African mainland species. Also see comment under leucogaster. Koopman (1986), included viridis, damarensis, and nigritellus (but not leucogaster) in this species.; [MSW3] Hill (1980b) considered African viridis and damarensis, and possibly leucogaster, to be conspecific with borbonicus, and Koopman (1986) included viridis, damarensis, and nigritellus (but not leucogaster) in this species. However, in a comprehensive revision of the African forms C. B. Robbins et al. (1985) rejected any affinity of borbonicus sensu stricto with African mainland species. I therefore restrict usage of the name borbonicus to the Réunion Isl form, and follow C. B. Robbins et al. (1985) in using leucogaster and dinganii for the smaller mainland species.; [HMW] Vespertilio borbonicus E. Geoffroy Saint-Hilaire, 1803 , Ile Bourbon [= Réunion Island ], Réunion. See S. robustus . The name borbonicus is restricted to the Réunion Island form. Monotypic.; [batnames2022] Hill (1980 b ) considered African viridis and damarensis , and possibly leucogaster , to be conspecific with borbonicus , and Koopman (1986) included viridis, damarensis , and nigritellus (but not leucogaster ) in this species. However, in a comprehensive revision of the African forms C. B. Robbins et al. (1985) rejected any affinity of borbonicus sensu stricto with African mainland species. I therefore restrict usage of the name borbonicus to the Réunion Isl form, and follow C.B. Robbins et al. (1985) in using leucogaster and dinganii for the smaller mainland species. Goodman et al (2020) recently suggested that the name borbonicus be considered a nomen dubium . The species has not been capture since its description and only a single museum specimens remains. Genetic material from this specimens nests within the recently described trujilloi , over which the name borbonicus has priority. Given the pressing need for a revision of the African forms of the genus (see discussion in Demos et al. (2018), we prefer to retain the name borbonicus as a valid species.; [IUCN] The taxonomic status of Scotophilus borbonicus on Madagascar is unclear. A single specimen from Madagascar is referred to as Scotophilus cf. borbonicus by Goodman at al. (2005), but unequivocal determination was not possible because of the poor state of the lectotype from La Réunion.; [batnames2023] Hill (1980 b ) considered African viridis and damarensis , and possibly leucogaster , to be conspecific with borbonicus , and Koopman (1986) included viridis, damarensis , and nigritellus (but not leucogaster ) in this species. However, in a comprehensive revision of the African forms C. B. Robbins et al. (1985) rejected any affinity of borbonicus  sensu stricto with African mainland species. We therefore restrict usage of the name borbonicus to the Réunion Isl form, and follow C.B. Robbins et al. (1985) in using leucogaster and dinganii for the smaller mainland species. Goodman et al (2020) recently suggested that the name borbonicus be considered a nomen dubium . The species has not been captured since its description and only a single museum specimens remains. Genetic material from this specimen nests within the recently described trujilloi , over which the name borbonicus has priority. Given the pressing need for a revision of the African forms of the genus (see discussion in Demos et al., 2018), we prefer to retain the name borbonicus as a valid species.; [MDD2023] likely synonymous with S. trujilloi based on molecular data; was considered a nomen dubium by Goodman et al., 2020, although this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi; [MDD2025_2.0] likely synonymous with S. trujilloi based on molecular data; was considered a nomen dubium by Goodman et al., 2020, although this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi; [batnames2025_1.7] Hill (1980b) considered African viridis and damarensis, and possibly leucogaster, to be conspecific with borbonicus, and Koopman (1986) included viridis, damarensis, and nigritellus (but not leucogaster) in this species. However, in a comprehensive revision of the African forms C. B. Robbins et al. (1985) rejected any affinity of borbonicus sensu stricto with African mainland species. We therefore restrict usage of the name borbonicus to the Réunion Isl form, and follow C.B. Robbins et al. (1985) in using leucogaster and dinganii for the smaller mainland species. Goodman et al (2020) recently suggested that the name borbonicus be considered a nomen dubium. The species has not been captured since its description and only a single museum specimens remains. Genetic material from this specimen nests within the recently described trujilloi, over which the name borbonicus has priority. Given the pressing need for a revision of the African forms of the genus (see discussion in Demos et al., 2018), we prefer to retain the name borbonicus as a valid species.; [MDD2025_2.2] likely synonymous with S. trujilloi based on molecular data; was considered a nomen dubium by Goodman et al., 2020, although this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi				leucogaster, viridis, damarensis			nigritellus, viridis, damarensis, borbonicus								The taxonomic status of Scotophilus borbonicus on Madagascar is unclear. A single specimen from Madagascar is referred to as Scotophilus cf. borbonicus by Goodman at al. (2005), but unequivocal determination was not possible because of the poor state of the lectotype from La Réunion.			borbonicus	borbonicus, burbonicus			borbonicus (É. Geoffroy Saint-Hilaire, 1803)|bourbonicus (Wiedemann, 1818) [incorrect subsequent spelling]|burbonicus (H. R. Schinz, 1821) [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Reunion, Madagascar	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Scotophilus borbonicus	Reunion Isl. (France).	E. Geoffroy	1806	Ann. Mus. Hist. Nat. Paris, 8:201.	Distribution: Ranging from Gambia to Ethiopia and from Kenya to Namibia and Natal; also Reunion in the Mascarenes and probably Madagascar		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Reunion, Madagascar; refs. 4.17, 87, 138, 139	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	E. Geoffroy	1803	Ann. Mus. Hist. Nat. Paris, 8:201.	Included equivocally in leucogaster by Hayman and Hill (1971:50-51); but see Koopman (1975:414-416). Hill (1980b) considered African viridis and damarensis, and possibly leucogaster to be conspecific with borbonicus, but C. B. Robbins et al. (1985) rejected any affinity with African mainland species. Also see comment under leucogaster. Koopman (1986), included viridis, damarensis, and nigritellus (but not leucogaster) in this species.	Madagascar, Reunion Isl (Mascarene Isis). Records from Mauritius (Mascarene Isis) are erroneous, see Cheke and Dahl (1981).	Reunion Isl (France).		E. GEOFFROY	1803	Size relatively small (forearm length, 41 -53 mm). Cingula of upper incisors relatively broad. Thumb relatively short.	Distribution: Ranging from Gambia to Ethiopia and from Kenya to Namibia and Natal; also Reunion in the Mascarenes and probably Madagascar	Four subspecies are here recognized:	S. b. nigritellus (Gambia to Ethiopia), S. b. viridis (Kenya to Natal), S. b. damarensis (Zambia to Namibia), S. b. borbonicus (Réunion, where probably now extinct).	128	species	S. borbonicus	E. GEOFFROY	1803	Scotophilus	genus	Scotophilus borbonicus				Size relatively small (forearm length, 41 -53 mm). Cingula of upper incisors relatively broad. Thumb relatively short.	Four subspecies are here recognized:		2. S. borbonicus (E. GEOFFROY 1803).	2	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nycticeiini	Scotophilus borbonicus	Scotophilus		borbonicus	E. Geoffroy	y	1803		Cat. Mamm. Mus. Nat. d'Hist. Nat.			46		Réunion House Bat	Réunion Isl (France).	Réunion Isl (Mascarene Isls). Records from Mauritius (Mascarene Isls) are erroneous, see Cheke and Dahl (1981). Reports from Madagascar have not been confirmed; see Peterson et al. (1995).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Critically Endangered. May be extinct; see Cheke and Dahl (1981).		Hill (1980b) considered African viridis and damarensis, and possibly leucogaster, to be conspecific with borbonicus, and Koopman (1986) included viridis, damarensis, and nigritellus (but not leucogaster) in this species. However, in a comprehensive revision of the African forms C. B. Robbins et al. (1985) rejected any affinity of borbonicus sensu stricto with African mainland species. I therefore restrict usage of the name borbonicus to the Réunion Isl form, and follow C. B. Robbins et al. (1985) in using leucogaster and dinganii for the smaller mainland species.	4C3D87E8FF846A3BFF509DED1740B9C3	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	888	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF7A6AC5FF4B9AF518B7BFCF.xml	Scotophilus borbonicus	Vespertilionidae	Scotophilus	borbonicus		1803	Scotophile de Bourbon @fr | Kleine Hausfledermaus @de | Scotofilode la @es | slade Reunion @en | Lesser Yellow House Bat @en | Bourbon Bat @en	Vespertilio borbonicus E. Geoffroy Saint-Hilaire, 1803 , Ile Bourbon [= Réunion Island ], Réunion. See S. robustus . The name borbonicus is restricted to the Réunion Island form. Monotypic.	Confirmed from Réunion I; in Madagascar , known only from a single specimen collected in the SW, butthis record cannot be confirmed; records from Mauritius are erroneous.	Forearm 51 mm (one specimen). The Lesser Yellow Bat is known from only one individual in very poor condition. Dorsum of the specimen is apparently reddish brown, and venteris whitish, which matches original description by E. Geoffroy Saint-Hilaire in 1803.	No information.	No information.	No information.	Some echolocation calls of two unidentified bats were recorded during fieldwork on Réunion . Signals of one call from “ Chiroptera spl” (QFC = 60 kHz and FM = 61 kHz) were very similar to those of Madagascan and African continental species of Scotophilus , suggesting that it might have been the Lesser Yellow Bat.	No information.	Classified as Data Deficient on The IUCN Red Last.	ACR (2017) | Barataud & Giosa (2013a) | Dorst (1947) | Geoffroy Saint-Hilaire (1803) | Goodman, Jenkins & Ratrimomanarivo (2005) | Jentink (1888) | Hill (1980b) | ICZN (2002b) | Koopman (1986) | Robbins et al. (1985) | Simmons (2005)	https://zenodo.org/record/6398404/files/figure.png	279. Lesser Yellow Bat Scotophilus borbonicus French: Scotophile de Bourbon / German: Kleine Hausfledermaus / Spanish: Scotofilo de la Isla de Reunion Other common names: Lesser Yellow House Bat , Bourbon Bat Taxonomy. Vespertilio borbonicus E. Geoffroy Saint-Hilaire, 1803 , Ile Bourbon [= Réunion Island ], Réunion. See S. robustus . The name borbonicus is restricted to the Réunion Island form. Monotypic. Distribution. Confirmed from Réunion I; in Madagascar , known only from a single specimen collected in the SW, butthis record cannot be confirmed; records from Mauritius are erroneous. Descriptive notes. Forearm 51 mm (one specimen). The Lesser Yellow Bat is known from only one individual in very poor condition. Dorsum of the specimen is apparently reddish brown, and venteris whitish, which matches original description by E. Geoffroy Saint-Hilaire in 1803. Habitat. No information. Food and Feeding. No information. Breeding. No information. Activity patterns. Some echolocation calls of two unidentified bats were recorded during fieldwork on Réunion . Signals of one call from “ Chiroptera spl” (QFC = 60 kHz and FM = 61 kHz) were very similar to those of Madagascan and African continental species of Scotophilus , suggesting that it might have been the Lesser Yellow Bat. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red Last. Bibliography. ACR (2017), Barataud & Giosa (2013a), Dorst (1947), Geoffroy Saint-Hilaire (1803), Goodman, Jenkins & Ratrimomanarivo (2005), Jentink (1888), Hill (1980b), ICZN (2002b), Koopman (1986), Robbins et al. (1985), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scotophilus borbonicus	Scotophilus		borbonicus	E. Geoffroy	1803	1	Cat. Mamm. Mus. Nat. d'Hist. Nat.	p. 46	R&eacute;union House Bat	None.	Réunion Isl (France).	Réunion Isl (Mascarene Isls). Records from Mauritius (Mascarene Isls) are erroneous, see Cheke and Dahl (1981). Reports from Madagascar have not been confirmed; see Peterson et al. (1995).	Not listed.	Data Deficient	Hill (1980 b ) considered African viridis and damarensis , and possibly leucogaster , to be conspecific with borbonicus , and Koopman (1986) included viridis, damarensis , and nigritellus (but not leucogaster ) in this species. However, in a comprehensive revision of the African forms C. B. Robbins et al. (1985) rejected any affinity of borbonicus sensu stricto with African mainland species. I therefore restrict usage of the name borbonicus to the Réunion Isl form, and follow C.B. Robbins et al. (1985) in using leucogaster and dinganii for the smaller mainland species. Goodman et al (2020) recently suggested that the name borbonicus be considered a nomen dubium . The species has not been capture since its description and only a single museum specimens remains. Genetic material from this specimens nests within the recently described trujilloi , over which the name borbonicus has priority. Given the pressing need for a revision of the African forms of the genus (see discussion in Demos et al. (2018), we prefer to retain the name borbonicus as a valid species.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	nomen dubium for Scotophilus trujilloi																																													IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	20064	Scotophilus borbonicus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scotophilus	borbonicus	(É. Geoffroy, 1803)	The taxonomic status of Scotophilus borbonicus on Madagascar is unclear. A single specimen from Madagascar is referred to as Scotophilus cf. borbonicus by Goodman at al. (2005), but unequivocal determination was not possible because of the poor state of the lectotype from La Réunion.	20000000	Scotophilus borbonicus	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	There have been exhaustive surveys of Réunion and this species has not been recorded for nearly 140 years. However, there is not enough information to determine its taxonomic status, as the holotype is badly damaged and other possible specimens cannot be reliably assigned to this species. If the taxonomic status of this species is confirmed, then the species may qualify for a higher threat category and may well be extinct.	The natural history of this species is not known.	The threats to this species are not known.	This species was considered to be relatively common in the early 19th Century on La Réunion; however, it was not recorded in the 20th century or 21st century (Cheke and Dahl 1981).	Unknown	This holotype was described from two specimens from La Réunion and one of these was designated as the lectotype but is now in very poor condition (Goodman et al. 2005). In Madagascar, it is known only from a single specimen collected in 1868, at Sarodrano, near Toliara in the south (Goodman et al . 2005). The identity of the Madagascar specimen cannot be confirmed due to absence of reliable material from La Réunion. The area near Sarodrano has been relatively well surveyed recently, but no new specimens were obtained (Goodman et al . 2005).		Terrestrial	There are no conservation measures in place.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scotophilus		borbonicus	É. Geoffroy Saint-Hilaire	1803	1	Cat. Mamm. Mus. Nat. d'Hist. Nat.	p. 55	R&eacute;union House Bat	None.	Réunion Isl (France).	Réunion Isl (Mascarene Isls). Records from Mauritius (Mascarene Isls) are erroneous, see Cheke and Dahl (1981). Reports from Madagascar have not been confirmed; see Peterson et al. (1995).	Not listed.	Data Deficient	Hill (1980 b ) considered African viridis and damarensis , and possibly leucogaster , to be conspecific with borbonicus , and Koopman (1986) included viridis, damarensis , and nigritellus (but not leucogaster ) in this species. However, in a comprehensive revision of the African forms C. B. Robbins et al. (1985) rejected any affinity of borbonicus  sensu stricto with African mainland species. We therefore restrict usage of the name borbonicus to the Réunion Isl form, and follow C.B. Robbins et al. (1985) in using leucogaster and dinganii for the smaller mainland species. Goodman et al (2020) recently suggested that the name borbonicus be considered a nomen dubium . The species has not been captured since its description and only a single museum specimens remains. Genetic material from this specimen nests within the recently described trujilloi , over which the name borbonicus has priority. Given the pressing need for a revision of the African forms of the genus (see discussion in Demos et al., 2018), we prefer to retain the name borbonicus as a valid species.	Scotophilus borbonicus	1006739	23	Réunion Yellow Bat	Réunion House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	SCOTOPHILINI	Scotophilus	NA	borbonicus	É. Geoffroy Saint-Hilaire	1803	1						Ile Bourbon [=Réunion Island].			borbonicus (É. Geoffroy Saint-Hilaire, 1803)	likely synonymous with S. trujilloi based on molecular data; was considered a nomen dubium by Goodman et al., 2020, although this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi	Goodman, S. M., Fratpietro, S., & Tortosa, P. (2020). Insight into the Identity and Origin of Scotophilus borbonicus (E. Geoffroy, 1803). Acta Chiropterologica, 22(1), 41-47.				Reunion	Africa	Afrotropic	DD	0	0	0	NA	1	sciname match	Scotophilus_borbonicus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotophilus_borbonicus	1006739	23	Réunion Yellow Bat	Réunion House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Scotophilini	Scotophilus	NA	borbonicus	É. Geoffroy Saint-Hilaire	1	Vespertilio borbonicus	Geoffroy Saint-Hilaire, É. 1803. Catalogue des Mammifères du Muséum national d'Histoire naturelle. Privately published, Paris, 272 pp.	https://www.digitale-sammlungen.de/en/details/bsb10482289	RMNH.MAM.28508	lectotype	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.28508.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.28508.b	Ile Bourbon [=Réunion Island].			likely synonymous with S. trujilloi based on molecular data; was considered a nomen dubium by Goodman et al., 2020, although this seems premature and the name borbonicus is likely better treated as the senior name of S. trujilloi	Goodman, S. M., Fratpietro, S., & Tortosa, P. (2020). Insight into the Identity and Origin of Scotophilus borbonicus (E. Geoffroy, 1803). Acta Chiropterologica, 22(1), 41-47.				Réunion	Africa	Afrotropic	DD	0	0	0	NA	1	sciname match	Scotophilus_borbonicus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scotophilus		borbonicus	É. Geoffroy Saint-Hilaire	1803	1	Cat. Mamm. Mus. Nat. d'Hist. Nat.	p. 55	R&eacute;union House Bat	None.	Réunion Isl (France).	Réunion Isl (Mascarene Isls). Records from Mauritius (Mascarene Isls) are erroneous, see Cheke and Dahl (1981). Reports from Madagascar have not been confirmed; see Peterson et al. (1995).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/20064/22024708/' target='_blank'>Data Deficient</a>	Hill (1980b) considered African viridis and damarensis, and possibly leucogaster, to be conspecific with borbonicus, and Koopman (1986) included viridis, damarensis, and nigritellus (but not leucogaster) in this species. However, in a comprehensive revision of the African forms C. B. Robbins et al. (1985) rejected any affinity of borbonicus sensu stricto with African mainland species. We therefore restrict usage of the name borbonicus to the Réunion Isl form, and follow C.B. Robbins et al. (1985) in using leucogaster and dinganii for the smaller mainland species. Goodman et al (2020) recently suggested that the name borbonicus be considered a nomen dubium. The species has not been captured since its description and only a single museum specimens remains. Genetic material from this specimen nests within the recently described trujilloi, over which the name borbonicus has priority. Given the pressing need for a revision of the African forms of the genus (see discussion in Demos et al., 2018), we prefer to retain the name borbonicus as a valid species.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotophilus borbonicus; Scotophilus borbonicus; Scotophilus borbonicus; nomen dubium for Scotophilus trujilloi; Scotophilus borbonicus; Scotophilus borbonicus; Scotophile de Bourbon; Kleine Hausfledermaus; Scotofilode la; slade Reunion; Lesser Yellow House Bat; Bourbon Bat; Réunion House Bat; R&eacute;union House Bat; S. borbonicus