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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1526	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri	Scotonycteris zenkeri		[MSW3] Current subspecific nomenclature does not adequately describe the known range of variation in this species; see Bergmans (1990).; [HMW] Scotonycteris zenkeri Matschie, 1894 , “Yaunde [= Yaoundé] Station,” Centre Region, Cameroon . Until recently, S. zenkeri was considered to have a relatively wide but disjunct distribution; however, mtDNA analyses distinguished four clades that resulted in raising subspecies occidentalis to the species level for the westernmost populations and description of a new species for the eastern populations ( S. bergmansi ). Populations from Nigeria might belong either to S. zenkeri or S. occidentalis , additional molecular data are needed. Identity of specimens from Bioko Island (currently assigned to bedfordi) needs to be confirmed. Existing karyological data belong to a bat from Gabon , which is currently assigned to S. bergmansi . Two subspecies recognized.; [batnames2022] Does not include occidentalis ; see Hassanin et al. (2015).; [MDD2022] previously included S. occidentalis; [IUCN] Scotonycteris zenkeri was considered to be widely distributed from Liberia in the west to the Democratic Republic of Congo in central Africa until the taxonomy of the tribe Scotonycterini was resolved (Hassanin et al. ; 2015) and S. zenkeri is now considered restricted to southwestern Cameroon and southeastern Nigeria.; [batnames2023] Does not include occidentalis ; see Hassanin et al. (2015).; [MDD2023] previously included S. occidentalis; [MDD2025_2.0] previously included S. occidentalis; [batnames2025_1.7] Does not include occidentalis; see Hassanin et al. (2015).; [MDD2025_2.2] previously included S. occidentalis						occidentalis.		luctus, foetidus, lanosus, morio, perniger, spurcus		zenkeri, bedfordi		zenkeri, bedfordi		zenkeri, bedfordi	Scotonycteris zenkeri was considered to be widely distributed from Liberia in the west to the Democratic Republic of Congo in central Africa until the taxonomy of the tribe Scotonycterini was resolved (Hassanin et al. ; 2015) and S. zenkeri is now considered restricted to southwestern Cameroon and southeastern Nigeria.	zenkeri, bedfordi		zenkeri, bedfordi	zenkeri, bedfordi	bedfordi, zenkeri 		zenkeri Matschie, 1894|bedfordi O. Thomas, 1904		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Zenker's fruit bat	Liberia – E Zaire	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Scotonycteris zenkeri	Cameroun, Yaunde.	Matschie	1894	Sitzb. Ges. Naturf. Fr. Berlin, p. 202.	Distribution: Same as for genus, including Fer nando Poo.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Zenker's fruit bat	Liberia – E Zaire	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Matschie	1894	Sitzb. Ges. Naturf. Fr. Berlin, p. 202.		Liberia to Congo Republic and E Zaire; Bioko.	Cameroon, Yaunde.		MATSCHIE	1894	Tufts at bases of ears absent. Size relatively small (forearm length, 45-55 mm; total length of skull, 24-27 mm).	Distribution: Same as for genus, including Fer nando Poo.	Two poorly marked subspecies.		32	species	S. zenkeri	MATSCHIE	1894	Scotonycteris	genus	Scotonycteris zenkeri				Tufts at bases of ears absent. Size relatively small (forearm length, 45-55 mm; total length of skull, 24-27 mm).	Two poorly marked subspecies.		1. S. zenkeri MATSCHIE 1894.	1	_S. z. bedfordi_ Thomas, 1904; _S. z. zenkeri_ Matschie, 1894			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Scotonycteris zenkeri	Scotonycteris		zenkeri	Matschie		1894		Sitzb. Ges. Naturf. Fr. Berlin			202		Zenker's Fruit Bat	Cameroon, Yaunde (Yaoundé).	Liberia to Republic of Congo and E Dem. Rep. Congo.	IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Lower Risk (lc).	bedfordi Thomas, 1904; occidentalis Hayman, 1947.	Current subspecific nomenclature does not adequately describe the known range of variation in this species; see Bergmans (1990).	4C3D87E8FF7A6AC5FF5397B81C10BD0F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	81	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFD3F63D89613651FEC1F305.xml	Scotonycteris zenkeri	Pteropodidae	Scotonycteris	zenkeri	Matschie	1894	Scotonyctére de Zenker @fr | Zenker Harlekinflughund @de | Scotonicterio de Zenker @es	Scotonycteris zenkeri Matschie, 1894 , “Yaunde [= Yaoundé] Station,” Centre Region, Cameroon . Until recently, S. zenkeri was considered to have a relatively wide but disjunct distribution; however, mtDNA analyses distinguished four clades that resulted in raising subspecies occidentalis to the species level for the westernmost populations and description of a new species for the eastern populations ( S. bergmansi ). Populations from Nigeria might belong either to S. zenkeri or S. occidentalis , additional molecular data are needed. Identity of specimens from Bioko Island (currently assigned to bedfordi) needs to be confirmed. Existing karyological data belong to a bat from Gabon , which is currently assigned to S. bergmansi . Two subspecies recognized.	S.z.zenkeriMatschie,1894—CameroonandpossiblyNigeria,butfurtheranalysisofmoleculardataisneededtoconfirmitspresenceinlattercountry. S. z. bedfordi Thomas, 1904 — Bioko I.	Head-body 65-85 mm (tailless), ear 12-17 mm , hindfoot 11- 14 mm , forearm 47-55 mm (on average females slightly longer); weight 16-24 g . Zenker’s Fruit Bat has white face markings on forehead and posterior corners of eyes; partially pale or white lips, especially around corners; short and slender muzzle; large dark brown eyes; naked ears with rounded tips, dark brown with lighter base, no basal ear patches; and moderately expansible lips. There are no epaulettes on adult males; dorsum is generally medium rusty brown to sepia-brown and speckled; hairs have dark brown basal one-third, pale gray middles, and rusty brown tips; pelage is dense, soft, woolly, and mid-dorsally 9-10 mm long. Central lower chest and belly are whitish to pale gray; flanks are medium to dark brown, not as sharply contrasting as in Hayman’s Fruit Bat ( S. occidentalis ); and pelage of underparts is shorter and sparser than dorsum, with stiff hairs on chest and belly. Wings have claw on second digit; membranes from sides of body are light brown to dark greenish brown and reticulated, attaching to first toes; finger joints are not yellowish; calcar is present. Skull is short and delicate; forehead region is almost straight; rostrum is relatively short; braincase is rounded; zygomatic width is relatively small, with relatively lightly built arches; palate is weakly arched; there are four thick, smooth palatal ridges, followed by 6-9 very thin, serrated, and irregular ridges; ridges 1-3 are not divided, and ridge 4 is occasionally divided in middle; bony palate extends clearly beyond posterior teeth; and post-dental palate has straight, converging lateral margins. Dental formula for all species of Scotonycterisis12/2,C1/1,P2/3, M 1/2 (x2) = 28. Upper incisors are relatively short and hardly curved, with blurred inner groove and without secondary cusps or serrated inner edges; C! is long and recurved; and premolars and molars are rounded, almost oval in transverse section, with relatively weak cusps.	[Lowland and montane forests, including untouched and disturbed rainforests, forest fringes, cultivated clearings over lowland forests (Bioko Island) from sea level up to elevations of ¢. 1100 m . Zenker’s Fruit Bat is never in large cleared areas or far from forests.	Zenker’s Fruit Bat is presumably frugivorous.	Pregnant Zenker’s Fruit Bats were captured in February in south-western Cameroon and October on Bioko Island.	No information.	Zenker’s Fruit Bat usually roosts alone.	Classified as Least Concern on The IUCN Red List. Zenker’s Fruit Bat has a wide distribution and apparently large population, and it is unlikely to be declining fast enough to be listed in a higher category. Nevertheless,itis rarely encountered, and current status needs to be reviewed because this assessment also includes populations currently assigned to Hayman’s Fruit Bat and Bergmans’s Fruit Bat ( S. bergmansi ). Habitat loss from logging and mining activities and conversion to agricultural use is a major threat.	Bergmans (1979a, 1991) | Bergmans, Fahr et al. (2017) | Eisentraut (1960a, 1973) | Fahr (2013c) | Hassanin et al. (2015)		32. Zenker’s Fruit Bat Scotonycteris zenkeri French: Scotonyctére de Zenker / German: ZenkerHarlekinflughund / Spanish: Scotonicterio de Zenker Taxonomy. Scotonycteris zenkeri Matschie, 1894 , “Yaunde [= Yaoundé] Station,” Centre Region, Cameroon . Until recently, S. zenkeri was considered to have a relatively wide but disjunct distribution; however, mtDNA analyses distinguished four clades that resulted in raising subspecies occidentalis to the species level for the westernmost populations and description of a new species for the eastern populations ( S. bergmansi ). Populations from Nigeria might belong either to S. zenkeri or S. occidentalis , additional molecular data are needed. Identity of specimens from Bioko Island (currently assigned to bedfordi) needs to be confirmed. Existing karyological data belong to a bat from Gabon , which is currently assigned to S. bergmansi . Two subspecies recognized. Subspecies and Distribution. S.z.zenkeriMatschie,1894—CameroonandpossiblyNigeria,butfurtheranalysisofmoleculardataisneededtoconfirmitspresenceinlattercountry. S. z. bedfordi Thomas, 1904 — Bioko I. Descriptive notes. Head-body 65-85 mm (tailless), ear 12-17 mm , hindfoot 11- 14 mm , forearm 47-55 mm (on average females slightly longer); weight 16-24 g . Zenker’s Fruit Bat has white face markings on forehead and posterior corners of eyes; partially pale or white lips, especially around corners; short and slender muzzle; large dark brown eyes; naked ears with rounded tips, dark brown with lighter base, no basal ear patches; and moderately expansible lips. There are no epaulettes on adult males; dorsum is generally medium rusty brown to sepia-brown and speckled; hairs have dark brown basal one-third, pale gray middles, and rusty brown tips; pelage is dense, soft, woolly, and mid-dorsally 9-10 mm long. Central lower chest and belly are whitish to pale gray; flanks are medium to dark brown, not as sharply contrasting as in Hayman’s Fruit Bat ( S. occidentalis ); and pelage of underparts is shorter and sparser than dorsum, with stiff hairs on chest and belly. Wings have claw on second digit; membranes from sides of body are light brown to dark greenish brown and reticulated, attaching to first toes; finger joints are not yellowish; calcar is present. Skull is short and delicate; forehead region is almost straight; rostrum is relatively short; braincase is rounded; zygomatic width is relatively small, with relatively lightly built arches; palate is weakly arched; there are four thick, smooth palatal ridges, followed by 6-9 very thin, serrated, and irregular ridges; ridges 1-3 are not divided, and ridge 4 is occasionally divided in middle; bony palate extends clearly beyond posterior teeth; and post-dental palate has straight, converging lateral margins. Dental formula for all species of Scotonycterisis12/2,C1/1,P2/3, M 1/2 (x2) = 28. Upper incisors are relatively short and hardly curved, with blurred inner groove and without secondary cusps or serrated inner edges; C! is long and recurved; and premolars and molars are rounded, almost oval in transverse section, with relatively weak cusps. Habitat. [Lowland and montane forests, including untouched and disturbed rainforests, forest fringes, cultivated clearings over lowland forests (Bioko Island) from sea level up to elevations of ¢. 1100 m . Zenker’s Fruit Bat is never in large cleared areas or far from forests. Food and Feeding. Zenker’s Fruit Bat is presumably frugivorous. Breeding. Pregnant Zenker’s Fruit Bats were captured in February in south-western Cameroon and October on Bioko Island. Activity patterns. No information. Movements, Home range and Social organization. Zenker’s Fruit Bat usually roosts alone. Status and Conservation. Classified as Least Concern on The IUCN Red List. Zenker’s Fruit Bat has a wide distribution and apparently large population, and it is unlikely to be declining fast enough to be listed in a higher category. Nevertheless,itis rarely encountered, and current status needs to be reviewed because this assessment also includes populations currently assigned to Hayman’s Fruit Bat and Bergmans’s Fruit Bat ( S. bergmansi ). Habitat loss from logging and mining activities and conversion to agricultural use is a major threat. Bibliography. Bergmans (1979a, 1991), Bergmans, Fahr et al. (2017), Eisentraut (1960a, 1973), Fahr (2013c), Hassanin et al. (2015).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Scotonycteris zenkeri	Scotonycteris		zenkeri	Matschie	1894	0	Sitzb. Ges. Naturf. Fr. Berlin	p. 202	Zenker's Fruit Bat	<b> bedfordi </b> Thomas, 1904.	Cameroon, Yaunde (Yaoundé).	Liberia to Republic of Congo and E Dem. Rep. Congo.	Not listed.	Near Threatened	Does not include occidentalis ; see Hassanin et al. (2015).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scotonycteris zenkeri	23	Zenker's Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	ROUSETTINAE	SCOTONYCTERINI	Scotonycteris	NA	zenkeri	Matschie	1894	0	Scotonycteris_zenkeri	Matschie, P. (1894). Neue Säugetiere aus den Sammlungen der Herren Zenker, Neumann, Stuhlmann und Emin. Sitzungsberichte der Gesellschaft naturforschender Freunde Berlin, 1894, 202.	https://www.biodiversitylibrary.org/item/148076#page/210/mode/1up	ZMB 66533		"Yaunde [= Yaoundé] Station," Centre Region, Cameroon.			zenkeri Matschie, 1894|bedfordi O. Thomas, 1904	previously included S. occidentalis	Hassanin, A., Khouider, S., Gembu, G. C., Goodman, S. M., Kadjo, B., Nesi, N., ... & Bonillo, C. (2015). The comparative phylogeography of fruit bats of the tribe Scotonycterini (Chiroptera, Pteropodidae) reveals cryptic species diversity related to African Pleistocene forest refugia. Comptes Rendus Biologies, 338(3), 197-211.	Nigeria|Cameroon|Equatorial Guinea	Africa	Afrotropic	NT	0	0	0	Scotonycteris_zenkeri	0	sciname match	Scotonycteris_zenkeri	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Scotonycteris zenkeri	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Scotonycteris	zenkeri	Matschie, 1894	Scotonycteris zenkeri was considered to be widely distributed from Liberia in the west to the Democratic Republic of Congo in central Africa until the taxonomy of the tribe Scotonycterini was resolved (Hassanin et al. ; 2015) and S. zenkeri is now considered restricted to southwestern Cameroon and southeastern Nigeria.	200000000	Scotonycteris zenkeri	Near Threatened	A2c	2021	2020-11-28 00:00:00 UTC	3.1	English	Scotonycteris zenkeri is assessed Near Threatened as it nearly qualifies as Vulnerable under criterion A2c because the species global population is suspected to have declined by 25–29% in the number of mature individuals over the last three generations (ca 12.9 years; GL = 4.3 yrs; Pacifici et al. 2013). This suspected decline is associated with rapid rates of deforestation (Nigeria has lost an estimated 90% of its moist forest; Ite 1995). There are reports of the conversion of 90% of an 11,000 hectares of forest reserve to small holder cassava farms in less than a decade (Enuoh and Bisong 2015). The recent taxonomic resolution of the tribe Scotonycterini now restricts S. zenkeri to the forests of southwestern Cameroon and southeastern Nigeria.	Scotonycteris zenkeri is a foliage-roosting species found in lowland pristine and moderately disturbed forests and has been recorded up to 1,100 m on Mt Kupe and Rumbi Hills in Cameroon (Eisentraut 1973). It forages in forest understory as most captures are with ground-level mist-nets (Eisentraut 1973, Mongombe et al. 2020, Obitte Unpubl. data) and two individuals in harp traps in southeastern Nigeria (Tanshi Unpubl. data).	The main threats to this species are forest fragmentation, degradation, and loss through logging, small holder farms, and wildfires.	The population status and trend for S. zenkeri is unknown. Among fruit bats, it is one of the less common species to capture during forest surveys. A recent intensive fruit bat survey in Cross River National Park and Afi Mountains Wildlife Sanctuary in southeastern Nigeria between 2017–2020 recorded only two individuals (Obitte unpubl. data). Another survey in the same protected areas of eastern Nigeria between 2016-2018 recorded six individuals (Tanshi unpubl. data).	Unknown	This species is restricted to southwestern Cameroon and southeastern Nigeria. Populations east of the River Niger are small fitting with Scotonycteris zenkeri . On the other hand, morphology of records from western Nigeria and the upper Guinean forest fits S. occidentalis (Happold and Happold 1978, Tanshi et al. 2019). The western limit of the species range in the Niger Delta remains unknown.		Terrestrial	Most of the population are in protected areas, but enforcement is necessary to reduce the high rate of logging in these PAs. In addition, awareness is required to educate farmers in boundary communities on best practices in Agriculture. Intensive surveys are needed to quantify this species population. Molecular analysis is needed to resolve taxonomy of the Nigerian population, especially to establish the western limits of the species range within the Niger Delta. In addition, satellite tracking is critical for clear understanding of its roosting ecology and home range.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Scotonycteris		zenkeri	Matschie	1894	0	Sitzb. Ges. Naturf. Fr. Berlin	p. 202	Zenker's Fruit Bat	<b> bedfordi </b> Thomas, 1904.	Cameroon, Yaunde (Yaoundé).	Liberia to Republic of Congo and E Dem. Rep. Congo.	Not listed.	Near Threatened	Does not include occidentalis ; see Hassanin et al. (2015).	Scotonycteris zenkeri	1004552	23	Zenker's Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	ROUSETTINAE	SCOTONYCTERINI	Scotonycteris	NA	zenkeri	Matschie	1894	0	Scotonycteris_zenkeri	Matschie, P. (1894). Neue Säugetiere aus den Sammlungen der Herren Zenker, Neumann, Stuhlmann und Emin. Sitzungsberichte der Gesellschaft naturforschender Freunde Berlin, 1894, 202.	https://www.biodiversitylibrary.org/item/148076#page/210/mode/1up	ZMB 66533		"Yaunde [= Yaoundé] Station," Centre Region, Cameroon.			zenkeri Matschie, 1894|bedfordi O. Thomas, 1904	previously included S. occidentalis	Hassanin, A., Khouider, S., Gembu, G. C., Goodman, S. M., Kadjo, B., Nesi, N., ... & Bonillo, C. (2015). The comparative phylogeography of fruit bats of the tribe Scotonycterini (Chiroptera, Pteropodidae) reveals cryptic species diversity related to African Pleistocene forest refugia. Comptes Rendus Biologies, 338(3), 197-211.				Nigeria|Cameroon|Equatorial Guinea	Africa	Afrotropic	NT	0	0	0	Scotonycteris_zenkeri	0	sciname match	Scotonycteris_zenkeri	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotonycteris_zenkeri	1004552	23	Zenker's Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Epomophorinae	Scotonycterini	Scotonycteris	NA	zenkeri	Matschie	0	Scotonycteris zenkeri	Matschie, P. 1894. Derselbe legte hierauf neue Säugethiere aus den Sammlungen der Herren Zenker, Neumann, Stuhlmann und Emin vor. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1894:194-206.	https://www.biodiversitylibrary.org/page/43581312	ZMB 66533	holotype		"Yaunde [= Yaoundé] Station," Centre Region, Cameroon.	3.87	11.52	previously included S. occidentalis	Hassanin, A., Khouider, S., Gembu, G. C., Goodman, S. M., Kadjo, B., Nesi, N., ... & Bonillo, C. (2015). The comparative phylogeography of fruit bats of the tribe Scotonycterini (Chiroptera, Pteropodidae) reveals cryptic species diversity related to African Pleistocene forest refugia. Comptes Rendus Biologies, 338(3), 197-211.				Nigeria|Cameroon|Equatorial Guinea	Africa	Afrotropic	NT	0	0	0	Scotonycteris_zenkeri	0	sciname match	Scotonycteris_zenkeri	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Scotonycteris		zenkeri	Matschie	1894	0	Sitzb. Ges. Naturf. Fr. Berlin	p. 202	Zenker's Fruit Bat	bedfordi Thomas, 1904.	Cameroon, Yaunde (Yaoundé).	Liberia to Republic of Congo and E Dem. Rep. Congo.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84464403/192236400/' target='_blank'>Near Threatened</a>	Does not include occidentalis; see Hassanin et al. (2015).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotonycteris zenkeri; Scotonycteris zenkeri; Scotonycteris zenkeri; Scotonycteris zenkeri; Scotonycteris zenkeri; Scotonycteris zenkeri; zenkeri; bedfordi; occidentalis; zenkeri; bedfordi; bedfordi; zenkeri; bedfordi; Scotonyctére de Zenker; Zenker Harlekinflughund; Scotonicterio de Zenker; Zenker's Fruit Bat; Zenker's Fruit Bat; Zenker's Fruit Bat; S. zenkeri