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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1521	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus	Scotoecus pallidus		[MSW2] Included in Nycticeius by Ellerman and Morrison-Scott (1951:177); but see J. E. Hill (1974b).; [MSW3] Included in Nycticeius by Ellerman and Morrison-Scott (1951); but see J. E. Hill (1974c). Reviewed by Bates and Harrison (1997). S. noctulinus may be an earlier name for this species.; [HMW] Scotophilus pallidus Dobson, 1876 , Mian Mir, Lahore, Punjab , Pakistan . Previously placed in Nycticeius , S. pallidus was transferred to Scotoecusas its only Asian member by J. E. Hill in 1974. In 1942, G. H. H. Tate suggested that the name noctulinus, introduced by I. Geoffroy Saint-Hilaire in 1834, might be an earlier name for the species, but until this is validated the name pallidus is here retained. Monotypic.; [batnames2022] Included in Nycticeius by Ellerman and Morrison-Scott (1951); but see J. E. Hill (1974 c ). Reviewed by Bates and Harrison(1997). S . noctulinus may be an earlier name for this species.; [IUCN] Ellerman and Morrison-Scott (1951) treated this taxon as Nycticeius pallidus Dobson, 1876. However, Hill (1974) reviewed the status of this species and included this taxon under Scotoecus Thomas, 1901, a trend that was followed by all subsequent workers. Tate (1942) opined that if noctulinus I. Geoffroy, 1831 is identifiable then it would be first name for pallidus Dobson, 1876 (Srinivasulu and Srinivasulu 2012).; [batnames2023] Included in Nycticeius by Ellerman and Morrison-Scott (1951); but see J. E. Hill (1974 c ). Reviewed by Bates and Harrison(1997). S . noctulinus may be an earlier name for this species.; [batnames2025_1.7] Included in Nycticeius by Ellerman and Morrison-Scott (1951); but see J. E. Hill (1974c). Reviewed by Bates and Harrison(1997). S. noctulinus may be an earlier name for this species.									noctulinus			pallidus 	pallidus - noctulinus	noctulinus, pallidus	Ellerman and Morrison-Scott (1951) treated this taxon as Nycticeius pallidus Dobson, 1876. However, Hill (1974) reviewed the status of this species and included this taxon under Scotoecus Thomas, 1901, a trend that was followed by all subsequent workers. Tate (1942) opined that if noctulinus I. Geoffroy, 1831 is identifiable then it would be first name for pallidus Dobson, 1876 (Srinivasulu and Srinivasulu 2012).	pallidus 	pallidus - noctulinus	noctulinus, pallidus	noctulinus, pallidus	pallidus 	pallidus - noctulinus	noctulinus (I. Geoffroy Saint-Hilaire, 1831)|pallidus (Dobson, 1876)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Pakistan, N India	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Scotoecus pallidus	Pakistan, Punjab, Lahore, Mian Mir.	Dobson	1876	Monogr. Asiat. Chiroptera, App. D:186.	Distribution: Confined to Pakistan and northern India.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Pakistan, N India	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Dobson	1876	Monogr. Asiat. Chiroptera, App. D:186.	Included in Nycticeius by Ellerman and Morrison-Scott (1951:177); but see J. E. Hill (1974b).	Pakistan, N India.	Pakistan, Punjab, Lahore, Mian Mir.		DOBSON	1876	Tragus relatively long, narrow, and straight. Anterior lower premolar reduced but not compressed in toothrow. Chiropatagium darkly pigmented. Ventral hair grayish white. Braincase slightly flattened. Anterior upper premolar absent. Size relatively large (forearm length, 34-38 mm).	Distribution: Confined to Pakistan and northern India.	No subspecies.		127	species	S. pallidus	DOBSON	1876	Scotoecus	genus	Scotoecus pallidus				Tragus relatively long, narrow, and straight. Anterior lower premolar reduced but not compressed in toothrow. Chiropatagium darkly pigmented. Ventral hair grayish white. Braincase slightly flattened. Anterior upper premolar absent. Size relatively large (forearm length, 34-38 mm).	No subspecies.		1. S. pallidus (DOBSON 1876).	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nycticeiini	Scotoecus pallidus	Scotoecus		pallidus	Dobson	y	1876		Monogr. Asiat. Chiroptera, App. D			186		Desert Yellow Lesser House Bat	Pakistan, Punjab, Lahore, Mian Mir.	Pakistan, N India.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	noctulinus I. Geoffroy, 1831 [see discussion in Hill (1974c)].	Included in Nycticeius by Ellerman and Morrison-Scott (1951); but see J. E. Hill (1974c). Reviewed by Bates and Harrison (1997). S. noctulinus may be an earlier name for this species.	4C3D87E8FFB46A0BFF7997F21FD8B734	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	761	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFF86A47FF7D941D1D73B206.xml	Scotoecus pallidus	Vespertilionidae	Scotoecus	pallidus		1876	Scotoéque d’Asie @fr | Fahle Hausfledermaus @de | Scotoecode Asia @es | Desert Yellow Bat @en | Desert Yellow House Bat @en	Scotophilus pallidus Dobson, 1876 , Mian Mir, Lahore, Punjab , Pakistan . Previously placed in Nycticeius , S. pallidus was transferred to Scotoecusas its only Asian member by J. E. Hill in 1974. In 1942, G. H. H. Tate suggested that the name noctulinus, introduced by I. Geoffroy Saint-Hilaire in 1834, might be an earlier name for the species, but until this is validated the name pallidus is here retained. Monotypic.	Pakistan ( Punjab and Sindh ) and N India ( Himachal Pradesh , Uttar Pradesh , Bihar , Jharkhand , and West Bengal ); it may occur in Maharashtra , WC India .	Head-body 47-59 mm, tail 27-2-41 mm, ear 9-15 mm, hindfoot 6-11 mm, forearm 34-39-5 mm; weight 9-14-3 g. Dorsal pelage of the Desert Yellow Lesser House Bat is pale brown tinged with fawn (hairs with pale bases), ventral pelage is grayish white. Ears, wings, and uropatagium are brown. Nostrils are circular, muzzle is broad and flat. Ears are medium-sized with rounded tip; tragus is narrow and long. Uropatagium stretches from well-developed calcar more or less to tail tip (except extreme tip). Penis is extremely elongate (c. 7 mm ). Upper surface of skull is flat and broad as in other Scotoecus ; there is one pair of upper and two pairs of lower premolars; rostrum is very broad and gently slopes from incisors to lambdoid crests; zygomatic arches are slender and weak; sagittal crest and lambdoid crests are weakly developed but more pronounced in older individuals, and there is a slight helmetto skull. Dental formula for all members of the genusis11/3,C1/1,P1/2,M 3/3 (x2) = 30.	Tropical thorn forests and urban areas. Recorded in a monsoon rice-growing area in Punjab , Pakistan . Ranges from sea level to 2500 m elevation.	No information.	In Bihar , northern India , pregnant females were reported in April, each having two fetuses.	Desert Yellow Lesser House Bats seem to prefer to roost in crevices in old buildings and hollows in trees.	Up to ten individuals have been recorded together in a colony.	Classified as Least Concern on The IUCN Red List. The Desert Yellow Lesser House Bat is relatively uncommon throughout its distribution but seems to be tolerant to some habitat alteration. However, the species is locally threatened by habitat loss due to urbanization.	Bates & Harrison (1997) | Belanger (1834) | Das (2003) | Ellerman & Morrison-Scott (1951) | Hill (1974a) | Javid (2011) | Molur & Srinivasulu (2008a) | Molur et al. (2002) | Roberts (1997) | Rosevear (1965) | Salim (2018) | Shahbaz et al. (2015) | Srinivasulu, C. et al. (2010) | Tate (1942b)	https://zenodo.org/record/6397760/files/figure.png	1. Desert Yellow Lesser House Bat Scotoecus pallidus French: Scotoéque d’Asie / German: Fahle Hausfledermaus / Spanish: Scotoeco de Asia Other common names: Desert Yellow Bat , Desert Yellow House Bat Taxonomy. Scotophilus pallidus Dobson, 1876 , Mian Mir, Lahore, Punjab , Pakistan . Previously placed in Nycticeius , S. pallidus was transferred to Scotoecusas its only Asian member by J. E. Hill in 1974. In 1942, G. H. H. Tate suggested that the name noctulinus, introduced by I. Geoffroy Saint-Hilaire in 1834, might be an earlier name for the species, but until this is validated the name pallidus is here retained. Monotypic. Distribution. Pakistan ( Punjab and Sindh ) and N India ( Himachal Pradesh , Uttar Pradesh , Bihar , Jharkhand , and West Bengal ); it may occur in Maharashtra , WC India . Descriptive notes. Head-body 47-59 mm, tail 27-2-41 mm, ear 9-15 mm, hindfoot 6-11 mm, forearm 34-39-5 mm; weight 9-14-3 g. Dorsal pelage of the Desert Yellow Lesser House Bat is pale brown tinged with fawn (hairs with pale bases), ventral pelage is grayish white. Ears, wings, and uropatagium are brown. Nostrils are circular, muzzle is broad and flat. Ears are medium-sized with rounded tip; tragus is narrow and long. Uropatagium stretches from well-developed calcar more or less to tail tip (except extreme tip). Penis is extremely elongate (c. 7 mm ). Upper surface of skull is flat and broad as in other Scotoecus ; there is one pair of upper and two pairs of lower premolars; rostrum is very broad and gently slopes from incisors to lambdoid crests; zygomatic arches are slender and weak; sagittal crest and lambdoid crests are weakly developed but more pronounced in older individuals, and there is a slight helmetto skull. Dental formula for all members of the genusis11/3,C1/1,P1/2,M 3/3 (x2) = 30. Habitat. Tropical thorn forests and urban areas. Recorded in a monsoon rice-growing area in Punjab , Pakistan . Ranges from sea level to 2500 m elevation. Food and Feeding. No information. Breeding. In Bihar , northern India , pregnant females were reported in April, each having two fetuses. Activity patterns. Desert Yellow Lesser House Bats seem to prefer to roost in crevices in old buildings and hollows in trees. Movements, Home range and Social organization. Up to ten individuals have been recorded together in a colony. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Desert Yellow Lesser House Bat is relatively uncommon throughout its distribution but seems to be tolerant to some habitat alteration. However, the species is locally threatened by habitat loss due to urbanization. Bibliography. Bates & Harrison (1997), Belanger (1834), Das (2003), Ellerman & Morrison-Scott (1951), Hill (1974a), Javid (2011), Molur & Srinivasulu (2008a), Molur et al. (2002), Roberts (1997), Rosevear (1965), Salim (2018), Shahbaz et al. (2015), Srinivasulu, C. et al. (2010), Tate (1942b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scotoecus pallidus	Scotoecus		pallidus	Dobson	1876	1	Monogr. Asiat. Chiroptera, App. D	p.186	Desert Yellow Lesser House Bat	 noctulinus I. Geoffroy, 1831 [see discussion in Hill (1974c)].	Pakistan, Punjab, Lahore, Mian Mir.	Pakistan, N India.	Not listed.	Least Concern	Included in Nycticeius by Ellerman and Morrison-Scott (1951); but see J. E. Hill (1974 c ). Reviewed by Bates and Harrison(1997). S . noctulinus may be an earlier name for this species.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scotoecus pallidus	23	Desert Yellow Lesser House Bat	Desert Yellow Bat|Desert Yellow House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Scotoecus	NA	pallidus	Dobson	1876	1	Scotophilus_pallidus	Dobson, G. E. (1876). Monograph of the Asiatic Chiroptera and Catalogue of the Species of Bats in the Collection of the Indian Museum, Calcutta. London, 186.	https://wellcomecollection.org/works/wznx4y6x/items?canvas=204&langCode=eng&sierraId=b28107925&source=viewer%2Fpaginator	ZSI 15615		Mian Mir, Lahore, Punjab, Pakistan.			noctulinus (I. Geoffroy Saint-Hilaire, 1831)|pallidus (Dobson, 1876)	NA	NA	Pakistan|India	Asia	Indomalaya	LC	0	0	0	Scotoecus_pallidus	0	sciname match	Scotoecus_pallidus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	20056	Scotoecus pallidus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scotoecus	pallidus	(Dobson, 1876)	Ellerman and Morrison-Scott (1951) treated this taxon as Nycticeius pallidus Dobson, 1876. However, Hill (1974) reviewed the status of this species and included this taxon under Scotoecus Thomas, 1901, a trend that was followed by all subsequent workers. Tate (1942) opined that if noctulinus I. Geoffroy, 1831 is identifiable then it would be first name for pallidus Dobson, 1876 (Srinivasulu and Srinivasulu 2012).	20000000	Scotoecus pallidus	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	An uncommon and rarely collected species. However, it exhibits tolerance of a degree of habitat modification, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category. Hence, listed as Least Concern.	Little is known about the habitat or ecology of this species except that this species is found in tropical thorn forests and urban areas preferring to roost in crevices and holes in old buildings and hollows in trees (Molur et al. 2002).	This species is locally threatened by habitat loss, largely due to increase in urbanization leading to disturbance and loss of roosting sites, and competition from invasive alien species (Molur et al. 2002).	The abundance, population size and trends for this species are not known (Molur et al. 2002).	Unknown	This species is endemic to South Asia and is presently known from India (Bihar, Himachal Pradesh, Jharkhand, Uttar Pradesh and West Bengal) and Pakistan (Punjab and Sind) (Srinivasulu and Srinivasulu 2012, Shahbaz et al. 2015). It has been recorded from 46 m up to an elevation of 2,500 m asl (Molur et al. 2002).		Terrestrial	There are no direct conservation measures in place for this species. The species has not been recorded from any protected areas. Further studies are needed on distribution, abundance, reproduction, ecology, population monitoring, habitat maintenance, limiting factor research, public awareness and conservation lobbying are actions recommended for the species (Molur et al. 2002).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scotoecus		pallidus	Dobson	1876	1	Monogr. Asiat. Chiroptera, App. D	p.186	Desert Yellow Lesser House Bat	 noctulinus I. Geoffroy, 1831 [see discussion in Hill (1974c)].	Pakistan, Punjab, Lahore, Mian Mir.	Pakistan, N India.	Not listed.	Least Concern	Included in Nycticeius by Ellerman and Morrison-Scott (1951); but see J. E. Hill (1974 c ). Reviewed by Bates and Harrison(1997). S . noctulinus may be an earlier name for this species.	Scotoecus pallidus	1005646	23	Desert Yellow Lesser House Bat	Desert Yellow Bat|Desert Yellow House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Scotoecus	NA	pallidus	Dobson	1876	1	Scotophilus_pallidus	Dobson, G. E. (1876). Monograph of the Asiatic Chiroptera and Catalogue of the Species of Bats in the Collection of the Indian Museum, Calcutta. London, 186.	https://wellcomecollection.org/works/wznx4y6x/items?canvas=204&langCode=eng&sierraId=b28107925&source=viewer%2Fpaginator	ZSI 15615		Mian Mir, Lahore, Punjab, Pakistan.			noctulinus (I. Geoffroy Saint-Hilaire, 1831)|pallidus (Dobson, 1876)	NA	NA				Pakistan|India	Asia	Indomalaya	LC	0	0	0	Scotoecus_pallidus	0	sciname match	Scotoecus_pallidus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotoecus_pallidus	1005646	23	Desert Yellow Lesser House Bat	Desert Yellow Bat|Desert Yellow House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Scotoecus	NA	pallidus	Dobson	1	Scotophilus pallidus	Dobson, G.E. 1876. Monograph of the Asiatic Chiroptera, and Catalogue of the Species of Bats in the Collection of the Indian Museum, Calcutta. Printed by Order of the Trustees of the Indian Museum, London, 228 pp.	https://wellcomecollection.org/works/wznx4y6x	ZSI 15615	holotype		Mian Mir, Lahore, Punjab, Pakistan.			NA	NA				Pakistan|India	Asia	Indomalaya	LC	0	0	0	Scotoecus_pallidus	0	sciname match	Scotoecus_pallidus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scotoecus		pallidus	Dobson	1876	1	Monogr. Asiat. Chiroptera, App. D	p.186	Desert Yellow Lesser House Bat	noctulinus I. Geoffroy, 1831 [see discussion in Hill (1974c)].	Pakistan, Punjab, Lahore, Mian Mir.	Pakistan, N India.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/20056/22025293/' target='_blank'>Least Concern</a>	Included in Nycticeius by Ellerman and Morrison-Scott (1951); but see J. E. Hill (1974c). Reviewed by Bates and Harrison(1997). S. noctulinus may be an earlier name for this species.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotoecus pallidus; Scotoecus pallidus; Scotoecus pallidus; Scotoecus pallidus; Scotoecus pallidus; Scotoecus pallidus; noctulinus; noctulinus; noctulinus; pallidus; Scotoéque d’Asie; Fahle Hausfledermaus; Scotoecode Asia; Desert Yellow Bat; Desert Yellow House Bat; Desert Yellow Lesser House Bat; Desert Yellow Bat; Desert Yellow House Bat; Desert Yellow Lesser House Bat; Desert Yellow Lesser House Bat; S. pallidus
