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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1520	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo	Scotoecus hirundo		[MSW2] Includes hindei; see Hayman and Hill (1971:36) and Robbins (1980:84); but see also). E. Hill (1974b).; [MSW3] Does not include albigula or hindei; see Happold et al. (1987), Happold and Happold (1989), and Taylor and Van der Merwe (1998); also see Happold and Happold (1997b).; [HMW] of dark-winged Scotoecus has been much debated. In one of the first revisions of the genus in 1953, Ellerman and colleagues placed all dark-winged Scotoecus in S. hirundo . In 1974, Hill divided them into S. hirundo and S. hinder, the second including a subspecies albigula. After evidence of sympatry between hindei and albigula, the latter was raised to species level. However, differences previously recognized as useful for species distinction were subsequently attributed to secondary sexual dimorphism. Against this background, in 1994 K. F. Koopman placed all dark-winged forms in S. hirundo , recognizing four subspecies: hirundo ( Senegal to Benin , possibly to Ethiopia ); falabae ( Nigeria and Cameroon ); hindei ( South Sudan and Somalia to south-eastern DR Congo ); and albigula ( Kenya to Angola ). In 1998, P. J. Taylor and M. van der Merwe tested Koopman’s arrangement using principal component analysis, which revealed that these forms were morphometrically distinguishable, and this was the basis for recognizing albigula and hinder as full species again. Although N. B. Simmons in 2005 treated albigula and hinde: as full species, in the absence of further evidence, M. Happold’s arrangement from 2013 for African Scotoecus is followed here, which considers albigula and hindei to be synonyms of S. hirundo . Thus, synonyms include albigula, artinu, falabae, and hindei. According to Happold in 2013, external measurements and those of teeth, measured by different authors indicate geographic variation, with increase in size from Ghana / Benin to Sudan / Ethiopia , to Kenya / Tanzania , and on to Malawi . Pelage color also varies, with bicolored hairs in individuals from West Africa, and unicolored hairs in individuals from Malawi . Although no subspecies are recognized here, the taxonomic status of names currently included under S. hirundo requires further clarification. Monotypic.; [batnames2022] Does not include albigula or hindei ; see Happold et al. (1987), Happold and Happold (1989), and Taylor and Van der Merwe(1998); also see Happold and Happold (1997 b ).; [MDD2022] sometimes considered to included S. albigula and S. hindei, although recent publications have retained them and they are tentatively retained here; [IUCN] We follow Happold (in press) by including Scotoecus albigula and S. hindei within S. hirundo .; [batnames2023] Does not include albigula or hindei ; see Happold et al. (1987), Happold and Happold (1989), and Taylor and Van der Merwe(1998); also see Happold and Happold (1997 b ).; [MDD2023] sometimes considered to included S. albigula and S. hindei, although recent publications have retained them and they are tentatively retained here; [MDD2025_2.0] sometimes considered to included S. albigula and S. hindei, although recent publications have retained them and they are tentatively retained here; [batnames2025_1.7] Does not include albigula or hindei; see Happold et al. (1987), Happold and Happold (1989), and Taylor and Van der Merwe(1998); also see Happold and Happold (1997b).; [MDD2025_2.2] sometimes considered to included S. albigula and S. hindei, although recent publications have retained them and they are tentatively retained here				hindei		albigula, artinii, falabae, hindei.	hirundo, falabae, hindei, albigula		artinii		albigula, artinii, falabae, and hindei	hirundo	hirundo - artinii	hirundo, artinii	We follow Happold (in press) by including Scotoecus albigula and S. hindei within S. hirundo .	hirundo	hirundo - artinii	hirundo, artinii	hirundo, artinii	hirundo 	hirundo - artinii	hirundo (de Winton, 1899)|artinii De Beaux, 1923		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Senegal – Ethiopia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Scotoecus hirundo	Ghana, Gambaga.	De Winton	1899	Ann. Mag. Nat. Hist., ser. 7, 4:355.	Distribution: Ranging from Senegal to Ethiopia and south to Malawi and Angola, but largely avoiding forested regions.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Senegal – Ethiopia; refs. 4.74, 84	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	de Winton	1899	Ann. Mag. Nat. Hist., ser. 7, 4:355.	Includes hindei; see Hayman and Hill (1971:36) and Robbins (1980:84); but see also). E. Hill (1974b).	Senegal to Ethiopia, south to Angola, Zambia and Malawi.	Ghana, Gambaga.		DE WINTON	1899	Tragus relatively short and broad, bent forward distally. Anterior lower molar not reduced, but compressed in toothrow. Chiropatagium darkly pigmented. Ventral hair grayish. Braincase slightly elevated. Anterior upper premolar present or absent. Size varying from relatively small to relatively large (forearm length, 29-38 mm). Two species may be represented in this complex.	Distribution: Ranging from Senegal to Ethiopia and south to Malawi and Angola, but largely avoiding forested regions.	Four subspecies are here recognized:	S. h. hirundo (Senegal to Benin, and possibly to Ethiopia), S. h. falabae (Nigeria and Cameroon), S. h. hindei (southern Sudan and Somalia to southeastern Zaire), S. h. albigula (Kenya to Angola).	127	species	S. hirundo	DE WINTON	1899	Scotoecus	genus	Scotoecus hirundo				Tragus relatively short and broad, bent forward distally. Anterior lower molar not reduced, but compressed in toothrow. Chiropatagium darkly pigmented. Ventral hair grayish. Braincase slightly elevated. Anterior upper premolar present or absent. Size varying from relatively small to relatively large (forearm length, 29-38 mm). Two species may be represented in this complex.	Four subspecies are here recognized:		3. S. hirundo (DE WINTON 1899).	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nycticeiini	Scotoecus hirundo	Scotoecus		hirundo	de Winton	y	1899		Ann. Mag. Nat. Hist., ser. 7	4		355		Dark-winged Lesser House Bat	Ghana, Gambaga.	Senegal to Ethiopia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	artinii de Winton, 1899.	Does not include albigula or hindei; see Happold et al. (1987), Happold and Happold (1989), and Taylor and Van der Merwe (1998); also see Happold and Happold (1997b).	4C3D87E8FFF86A47FF7D941D1D73B206	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	761	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFF86A40FF809F241ACAB9A6.xml	Scotoecus hirundo	Vespertilionidae	Scotoecus	hirundo		1899	Scotoéque a ailes noires @fr | Schwarzflligel-Hausfledermaus @de | Scotoecode alas negras @es | \\White-bellied Lesser House Bat @en | Hinde's Lesser House Bat @en	of dark-winged Scotoecus has been much debated. In one of the first revisions of the genus in 1953, Ellerman and colleagues placed all dark-winged Scotoecus in S. hirundo . In 1974, Hill divided them into S. hirundo and S. hinder, the second including a subspecies albigula. After evidence of sympatry between hindei and albigula, the latter was raised to species level. However, differences previously recognized as useful for species distinction were subsequently attributed to secondary sexual dimorphism. Against this background, in 1994 K. F. Koopman placed all dark-winged forms in S. hirundo , recognizing four subspecies: hirundo ( Senegal to Benin , possibly to Ethiopia ); falabae ( Nigeria and Cameroon ); hindei ( South Sudan and Somalia to south-eastern DR Congo ); and albigula ( Kenya to Angola ). In 1998, P. J. Taylor and M. van der Merwe tested Koopman’s arrangement using principal component analysis, which revealed that these forms were morphometrically distinguishable, and this was the basis for recognizing albigula and hinder as full species again. Although N. B. Simmons in 2005 treated albigula and hinde: as full species, in the absence of further evidence, M. Happold’s arrangement from 2013 for African Scotoecus is followed here, which considers albigula and hindei to be synonyms of S. hirundo . Thus, synonyms include albigula, artinu, falabae, and hindei. According to Happold in 2013, external measurements and those of teeth, measured by different authors indicate geographic variation, with increase in size from Ghana / Benin to Sudan / Ethiopia , to Kenya / Tanzania , and on to Malawi . Pelage color also varies, with bicolored hairs in individuals from West Africa, and unicolored hairs in individuals from Malawi . Although no subspecies are recognized here, the taxonomic status of names currently included under S. hirundo requires further clarification. Monotypic.	Patchily from Senegal , Gambia , and Guinea-Bissau E to Ethiopia and Somalia and S to S Angola , S Zambia , and S Mozambique .	Head-body ¢.49-65 mm, tail 28-40 mm, ear 9-14 mm, hindfoot 8-10 mm, forearm 31-40 mm (males) and 28-37 mm (females); weight 8-15 g. Males average larger than females in some external measurements, have more massive teeth, greater upper tooth row length and molar width. Penis of the Dark-winged Lesser House Bat is extremely long (14-16 mm). Pelage is soft. Wing and interfemoral membranes are blackish brown, slightly translucent. Dorsal pelage is chocolate brown or medium to pale sepia-brown, unicolored ( Malawi ) or bicolored, with white bases and sepia-brown tips (West Africa). Ventral pelage varies from gray or grayish white to beige or pale beige, with chest sometimes paler than rest of venter. Head is fairly flattened; muzzle is broad, dark brown, and almost naked. Nostrils open to sides from small transverse pads. Eyes are small. Ears are well separated, oval, rounded, and comparatively short for a vespertilionid. Tragus is short (4-6 mm), with anterior margin strongly concave, posterior margin with smooth, obtuse angle above mid-height, and rounded tip. Skull is somewhat flattened but less flattened than in the Light-winged Lesser House Bat (S. albofuscus ). C!' has anterior surface flat and is lightly grooved. P? is usually present, displaced lingually, and minute. P* is pointed, and in contact with C'. M® has three ridges. P, is at least two-thirds the height of P,. Dental formula is as in congenersor 11/3, C1/1, P 2/2, M 3/3 (x2) = 32. Two distinct karyotypes have been recorded for specimens tentatively assigned to S. hirundo: D. Koubinova and colleagues in 2013 reported that a male from Senegal had 2n = 30, FN = 50, and FNa = 46, with a metacentric X-chromosome, and a probably acrocentric Y-chromosome, whereas a female from the Ivory Coast reported by M. Volleth and colleagues in 2006 had 2n = 30, FN = 54, and FNa = 50. The specimen tentatively identified as S. hirundo from Senegal differed 1:6% in the rag2 gene in comparison with a specimen from Tanzania , and may represent an undescribed cryptic species.	Open woodlands in West Africa, woodlands and deciduous thicket and bushland in the east of its range, and both wetter and drier miombo woodlands in the south of its range. In Malawi , the Dark-winged Lesser House Bat occurs in miombo woodland,relict rainforest, and riverine woodland, gardens,villages and towns. In Nigeria , it occurs in the Guinea savanna zone, especially on the Jos Plateau. Elevational range is 100-1500 m.	Dark-winged Lesser House Bats forage by moderately fast hawking, in open spaces, above the canopy, or over grassland, usually 5-25 m aboveground. There 1s no information about diet.	In Malawi , restricted seasonal monoestry with early pregnancy in September, pregnancy in October, parturition in November, adult females lactating from November to January, volant young with unfused epiphyses from January to March, adult females post-lactating from February to March, and adults not palpably pregnant from April to June. From September to January, most adult males had abdominal testes; from February to May, most had large scrotal testes. Mating seems to occur around May, followed by sperm storage or reproductive delay until about late August. Litter size 1S two.	Natural day roosts are not known, but the species is known to roost under corrugated iron roofs of huts and houses. Search-phase call shape of seven Malawian individuals flying in open space after release was very shallow linear FM, occasionally alternated with a curvilinear steep/shallow FM. For very shallow linear FM calls, intensity was high, end frequencies were 30-34 kHz, bandwidths were 2—4 kHz, and call durations were ¢.5-6 milliseconds. For curvilinear steep/shallow FM calls, intensity was high, end frequencies were 32-34 kHz, bandwidths were ¢.6-7 kHz, and call durations were ¢.5—6 milliseconds. When calls were alternated, end frequency was often lower in very shallow linear FM than in curvilinear steep/shallow FM. Approach-phase calls were firstly curvilinear steep/shallow FM sweeps of bandwidth up to 15 kHz, and then steep FM/QCF of greater bandwidth. Calls emitted close to the ground in cluttered areas were steep FM/QCF sweeps, intensity was high, with start frequency up to 70 kHz, and end frequencies of 33-40 kHz. Reflecting the differences in skull size, males echolocate at lower frequencies than females.	In Masalani, southern Kenya , in 1980, T. J. O’Shea and T. A. Vaughan observed that the abundance of the Dark-winged Lesser House Bat was comparatively low during the long dry season (May-October), and this was preceded by a buildup of subcutaneous fat. These observations led the authors to speculate on emigration from Masalanito places that receive rainfall during May-October (e.g. highlands of western Kenya or the coast). The bedbug Cacodmus sparsilis ( Hemiptera ) and the flea Echidnophaga aethiops ( Siphonaptera ) have been recorded on this species. On following pages 3 ught-wınged Lesser House Bat (Sooroecus albofuscus ) 4 Rupee s Bat (Vansoma meppeln) 5 7 CommonThıd<thumbed Bat (G sdıropus tylopus ) B JavanThıd<thumbed Bat (G sdvropuspvanus) 9 Dormers Bat Noctule ( Nyctalus furvus ), 13 Mounts n Noctule (Nycralus montanus ).	Classified as Least Concern on The IUCN Red List. The Darkwinged Lesser House Bat is apparently rare in West Africa, but common in Malawi .	Ansell & Dowsett (1988) | Ellerman & Morrison-Scott (1951) | Ellerman et al. (1953) | Happold, M. (2013bh) | Hill (1974a) | Hill & Harrison (1987) | Koopman (1993, 1994) | Koubinové etal. (2013) | O'Shea & Vaughan (1980) | Simmons (2005) | Taylor & van der Merwe (1998) | Taylor et al. (2005) | Volleth et al. (2006)	https://zenodo.org/record/6397762/files/figure.png	2. Dark-winged Lesser House Bat Scotoecus hirundo French: Scotoéque a ailes noires / German: Schwarzflligel-Hausfledermaus / Spanish: Scotoeco de alas negras Other common names: \\White-bellied Lesser House Bat , Hinde's Lesser House Bat Taxonomy. Scotophilus hirundo de Winton, 1899 , “Gambaga [ Ghana ], 1300 feet [= 396 m ].” Scotoecus was considered a subgenus of Nycticetus by J. R. Ellerman and T. C. S. Morrison-Scott in 1951 but subsequently recognized as a distinct genus by D. R. Rosevear in 1965, a treatment followed by J. E. Hill in 1974. Taxonomy of dark-winged Scotoecus has been much debated. In one of the first revisions of the genus in 1953, Ellerman and colleagues placed all dark-winged Scotoecus in S. hirundo . In 1974, Hill divided them into S. hirundo and S. hinder, the second including a subspecies albigula. After evidence of sympatry between hindei and albigula, the latter was raised to species level. However, differences previously recognized as useful for species distinction were subsequently attributed to secondary sexual dimorphism. Against this background, in 1994 K. F. Koopman placed all dark-winged forms in S. hirundo , recognizing four subspecies: hirundo ( Senegal to Benin , possibly to Ethiopia ); falabae ( Nigeria and Cameroon ); hindei ( South Sudan and Somalia to south-eastern DR Congo ); and albigula ( Kenya to Angola ). In 1998, P. J. Taylor and M. van der Merwe tested Koopman’s arrangement using principal component analysis, which revealed that these forms were morphometrically distinguishable, and this was the basis for recognizing albigula and hinder as full species again. Although N. B. Simmons in 2005 treated albigula and hinde: as full species, in the absence of further evidence, M. Happold’s arrangement from 2013 for African Scotoecus is followed here, which considers albigula and hindei to be synonyms of S. hirundo . Thus, synonyms include albigula, artinu, falabae, and hindei. According to Happold in 2013, external measurements and those of teeth, measured by different authors indicate geographic variation, with increase in size from Ghana / Benin to Sudan / Ethiopia , to Kenya / Tanzania , and on to Malawi . Pelage color also varies, with bicolored hairs in individuals from West Africa, and unicolored hairs in individuals from Malawi . Although no subspecies are recognized here, the taxonomic status of names currently included under S. hirundo requires further clarification. Monotypic. Distribution. Patchily from Senegal , Gambia , and Guinea-Bissau E to Ethiopia and Somalia and S to S Angola , S Zambia , and S Mozambique . Descriptive notes. Head-body ¢.49-65 mm, tail 28-40 mm, ear 9-14 mm, hindfoot 8-10 mm, forearm 31-40 mm (males) and 28-37 mm (females); weight 8-15 g. Males average larger than females in some external measurements, have more massive teeth, greater upper tooth row length and molar width. Penis of the Dark-winged Lesser House Bat is extremely long (14-16 mm). Pelage is soft. Wing and interfemoral membranes are blackish brown, slightly translucent. Dorsal pelage is chocolate brown or medium to pale sepia-brown, unicolored ( Malawi ) or bicolored, with white bases and sepia-brown tips (West Africa). Ventral pelage varies from gray or grayish white to beige or pale beige, with chest sometimes paler than rest of venter. Head is fairly flattened; muzzle is broad, dark brown, and almost naked. Nostrils open to sides from small transverse pads. Eyes are small. Ears are well separated, oval, rounded, and comparatively short for a vespertilionid. Tragus is short (4-6 mm), with anterior margin strongly concave, posterior margin with smooth, obtuse angle above mid-height, and rounded tip. Skull is somewhat flattened but less flattened than in the Light-winged Lesser House Bat (S. albofuscus ). C!' has anterior surface flat and is lightly grooved. P? is usually present, displaced lingually, and minute. P* is pointed, and in contact with C'. M® has three ridges. P, is at least two-thirds the height of P,. Dental formula is as in congenersor 11/3, C1/1, P 2/2, M 3/3 (x2) = 32. Two distinct karyotypes have been recorded for specimens tentatively assigned to S. hirundo: D. Koubinova and colleagues in 2013 reported that a male from Senegal had 2n = 30, FN = 50, and FNa = 46, with a metacentric X-chromosome, and a probably acrocentric Y-chromosome, whereas a female from the Ivory Coast reported by M. Volleth and colleagues in 2006 had 2n = 30, FN = 54, and FNa = 50. The specimen tentatively identified as S. hirundo from Senegal differed 1:6% in the rag2 gene in comparison with a specimen from Tanzania , and may represent an undescribed cryptic species. Habitat. Open woodlands in West Africa, woodlands and deciduous thicket and bushland in the east of its range, and both wetter and drier miombo woodlands in the south of its range. In Malawi , the Dark-winged Lesser House Bat occurs in miombo woodland,relict rainforest, and riverine woodland, gardens,villages and towns. In Nigeria , it occurs in the Guinea savanna zone, especially on the Jos Plateau. Elevational range is 100-1500 m. Food and Feeding. Dark-winged Lesser House Bats forage by moderately fast hawking, in open spaces, above the canopy, or over grassland, usually 5-25 m aboveground. There 1s no information about diet. Breeding. In Malawi , restricted seasonal monoestry with early pregnancy in September, pregnancy in October, parturition in November, adult females lactating from November to January, volant young with unfused epiphyses from January to March, adult females post-lactating from February to March, and adults not palpably pregnant from April to June. From September to January, most adult males had abdominal testes; from February to May, most had large scrotal testes. Mating seems to occur around May, followed by sperm storage or reproductive delay until about late August. Litter size 1S two. Activity patterns. Natural day roosts are not known, but the species is known to roost under corrugated iron roofs of huts and houses. Search-phase call shape of seven Malawian individuals flying in open space after release was very shallow linear FM, occasionally alternated with a curvilinear steep/shallow FM. For very shallow linear FM calls, intensity was high, end frequencies were 30-34 kHz, bandwidths were 2—4 kHz, and call durations were ¢.5-6 milliseconds. For curvilinear steep/shallow FM calls, intensity was high, end frequencies were 32-34 kHz, bandwidths were ¢.6-7 kHz, and call durations were ¢.5—6 milliseconds. When calls were alternated, end frequency was often lower in very shallow linear FM than in curvilinear steep/shallow FM. Approach-phase calls were firstly curvilinear steep/shallow FM sweeps of bandwidth up to 15 kHz, and then steep FM/QCF of greater bandwidth. Calls emitted close to the ground in cluttered areas were steep FM/QCF sweeps, intensity was high, with start frequency up to 70 kHz, and end frequencies of 33-40 kHz. Reflecting the differences in skull size, males echolocate at lower frequencies than females. Movements, Home range and Social organization. In Masalani, southern Kenya , in 1980, T. J. O’Shea and T. A. Vaughan observed that the abundance of the Dark-winged Lesser House Bat was comparatively low during the long dry season (May-October), and this was preceded by a buildup of subcutaneous fat. These observations led the authors to speculate on emigration from Masalanito places that receive rainfall during May-October (e.g. highlands of western Kenya or the coast). The bedbug Cacodmus sparsilis ( Hemiptera ) and the flea Echidnophaga aethiops ( Siphonaptera ) have been recorded on this species. On following pages 3 ught-wınged Lesser House Bat (Sooroecus albofuscus ) 4 Rupee s Bat (Vansoma meppeln) 5 7 CommonThıd<thumbed Bat (G sdıropus tylopus ) B JavanThıd<thumbed Bat (G sdvropuspvanus) 9 Dormers Bat Noctule ( Nyctalus furvus ), 13 Mounts n Noctule (Nycralus montanus ). 14 Greater Noctule (Nycta us lasıopteıuá '15 Common Indodı neseThıdı thumbed Bat (G sdvropus buoeplıs us) 6 DarkThıd<thumbed Bat (G sdrmpus equ us) (Scorozvus dormen) 10 Ch nese Noctu e (Nycra us plancyr) B rd-I ke Noctule (Nycta us avıaron 12 Japanese Noctule (Nycraus noctula ) 16 Leıslers Noctule (Nyctaus lens/on) 17 Azores Noctule (Nwtaus azoreunl) Status and Conservation. Classified as Least Concern on The IUCN Red List. The Darkwinged Lesser House Bat is apparently rare in West Africa, but common in Malawi . Bibliography. Ansell & Dowsett (1988), Ellerman & Morrison-Scott (1951), Ellerman et al. (1953), Happold, M. (2013bh), Hill (1974a), Hill & Harrison (1987), Koopman (1993, 1994), Koubinové etal. (2013), O'Shea & Vaughan (1980), Simmons (2005), Taylor & van der Merwe (1998), Taylor et al. (2005), Volleth et al. (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scotoecus hirundo	Scotoecus		hirundo	de Winton	1899	1	Ann. Mag. Nat. Hist.	ser. 7, 4: 355	Dark-winged Lesser House Bat	 artinii de Winton, 1899.	Ghana, Gambaga.	Senegal to Ethiopia.	Not listed.	Least Concern	Does not include albigula or hindei ; see Happold et al. (1987), Happold and Happold (1989), and Taylor and Van der Merwe(1998); also see Happold and Happold (1997 b ).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scotoecus hirundo	23	Dark-winged Lesser House Bat	White-bellied Lesser House Bat|Hinde's Lesser House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Scotoecus	NA	hirundo	de Winton	1899	1	Scotophilus_hirundo	de Winton, W. E. (1899). On mammals collected by Lieut.-Colonel W. Giffard in the Northern Territory of the Gold Coast. Annals and Magazine of Natural History, ser. 7, 4, 355.	https://www.biodiversitylibrary.org/item/81048#page/371/mode/1up	BM 1899.6.15.8		"Gambaga [Ghana], 1300 feet [= 396 m]."			hirundo (de Winton, 1899)|artinii De Beaux, 1923	sometimes considered to included S. albigula and S. hindei, although recent publications have retained them and they are tentatively retained here	Happold, M. 2012. Scotoecus hirundo. In: D. C. D. Happold and M. Happold (eds), The Mammals of Africa, Academic Press, Amsterdam, The Netherlands.|Monadjem, A., Demos, T. C., Dalton, D. L., Webala, P. W., Musila, S., Kerbis Peterhans, J. C., & Patterson, B. D. (2020). A revision of pipistrelle-like bats (Mammalia: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society, zlaa087.	Senegal|Mali|Sierra Leone|Ghana|Benin|Nigeria|Cameroon|Central African Republic|South Sudan|Sudan|Ethiopia|Uganda|Kenya|Somalia|Tanzania	Africa	Afrotropic	LC	0	0	0	Scotoecus_hirundo	0	sciname match	Scotoecus_hirundo	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	20055	Scotoecus hirundo	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scotoecus	hirundo	de Winton, 1899	We follow Happold (in press) by including Scotoecus albigula and S. hindei within S. hirundo .	20000000	Scotoecus hirundo	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Although this species is known mainly from isolated records from a large area, it is listed as Least Concern in view of its wide distribution, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	There is little information available on the natural history of this poorly known bat. Rosevear (1965), indicates that the species is present in Sudan woodland and also high forest. Happold (1987), reports that in Nigeria the species is found in the Guinea savanna zone, especially on the Jos plateau, and is found roosting in houses. Bergmans (1977) recorded the species flying along a stream.	While the threats to this species are not well known, it is presumed not to have any major threats in view of the species wide distribution in habitats that are not rapidly declining throughout much of its recorded range.	It is very rarely recorded, and there is little information is available on the population abundance or size of this species.	Unknown	This widespread African species has been reported as scattered records from West Africa (The Gambia, Guinea, Sierra Leone, Côte d'Ivoire, Ghana and Nigeria), Central Africa (northern Cameroon, Central African Republic, the Democratic Republic of the Congo and Angola) and East Africa (Sudan, Ethiopia, Uganda, Kenya, Tanzania, Malawi, Mozambique and eastern Zambia). It is possibly present in Benin, Chad and Somalia but this requires confirmation.		Terrestrial	There appear to be no direct conservation measures in place. It has been recorded from the Zinave National Park in Mozambique (Cotterill 2001), and from the Mole National Park in Ghana (Grubb et al. 1998), and in view of its wide range, is presumably present in additional protected areas. Further studies are needed into the natural history and possible threats to this poorly known species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scotoecus		hirundo	de Winton	1899	1	Ann. Mag. Nat. Hist.	ser. 7, 4: 355	Dark-winged Lesser House Bat	 artinii de Winton, 1899.	Ghana, Gambaga.	Senegal to Ethiopia.	Not listed.	Least Concern	Does not include albigula or hindei ; see Happold et al. (1987), Happold and Happold (1989), and Taylor and Van der Merwe(1998); also see Happold and Happold (1997 b ).	Scotoecus hirundo	1005645	23	Dark-winged Lesser House Bat	White-bellied Lesser House Bat|Hinde's Lesser House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Scotoecus	NA	hirundo	de Winton	1899	1	Scotophilus_hirundo	de Winton, W. E. (1899). On mammals collected by Lieut.-Colonel W. Giffard in the Northern Territory of the Gold Coast. Annals and Magazine of Natural History, ser. 7, 4, 355.	https://www.biodiversitylibrary.org/item/81048#page/371/mode/1up	BM 1899.6.15.8		"Gambaga [Ghana], 1300 feet [= 396 m]."			hirundo (de Winton, 1899)|artinii De Beaux, 1923	sometimes considered to included S. albigula and S. hindei, although recent publications have retained them and they are tentatively retained here	Happold, M. 2012. Scotoecus hirundo. In: D. C. D. Happold and M. Happold (eds), The Mammals of Africa, Academic Press, Amsterdam, The Netherlands.|Monadjem, A., Demos, T. C., Dalton, D. L., Webala, P. W., Musila, S., Kerbis Peterhans, J. C., & Patterson, B. D. (2020). A revision of pipistrelle-like bats (Mammalia: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society, zlaa087.				Senegal|Mali|Sierra Leone|Ghana|Benin|Nigeria|Cameroon|Central African Republic|South Sudan|Sudan|Ethiopia|Uganda|Kenya|Somalia|Tanzania	Africa	Afrotropic	LC	0	0	0	Scotoecus_hirundo	0	sciname match	Scotoecus_hirundo	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scotoecus_hirundo	1005645	23	Dark-winged Lesser House Bat	White-bellied Lesser House Bat|Hinde's Lesser House Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Scotoecus	NA	hirundo	de Winton	1	Scotophilus hirundo	de Winton, W.E. 1899-11-01. On mammals collected by Lieut.-Colonel W. Giffard in the northern territory of the Gold Coast. Annals and Magazine of Natural History (7)4(23):353-359.	https://www.biodiversitylibrary.org/page/19209098	BMNH:Mamm:1899.6.15.8	holotype	https://data.nhm.ac.uk/object/5b14bf84-225b-4786-8028-c97c028c108f | https://data.nhm.ac.uk/object/b20ed957-909b-40fe-bd10-574a3d366ca2	"Gambaga [Ghana], 1300 feet [= 396 m]."			sometimes considered to included S. albigula and S. hindei, although recent publications have retained them and they are tentatively retained here	Happold, M. 2012. Scotoecus hirundo. In: D. C. D. Happold and M. Happold (eds), The Mammals of Africa, Academic Press, Amsterdam, The Netherlands.|Monadjem, A., T. C. Demos, D. L. Dalton, P. W. Webala, S. Musila, J. C. K. Peterhans and B. D. Patterson (2021). A revision of pipistrelle-like bats (Mammalia: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society 191(4): 1114–1146.				Senegal|Mali|Sierra Leone|Ghana|Benin|Nigeria|Cameroon|Central African Republic|South Sudan|Sudan|Ethiopia|Uganda|Kenya|Somalia|Tanzania	Africa	Afrotropic	LC	0	0	0	Scotoecus_hirundo	0	sciname match	Scotoecus_hirundo	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scotoecus		hirundo	de Winton	1899	1	Ann. Mag. Nat. Hist.	ser. 7, 4: 355	Dark-winged Lesser House Bat	artinii de Winton, 1899.	Ghana, Gambaga.	Senegal to Ethiopia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/20055/22025420/' target='_blank'>Least Concern</a>	Does not include albigula or hindei; see Happold et al. (1987), Happold and Happold (1989), and Taylor and Van der Merwe(1998); also see Happold and Happold (1997b).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scotoecus hirundo; Scotoecus hirundo; Scotoecus hirundo; Scotoecus hirundo; Scotoecus hirundo; Scotoecus hirundo; artinii; albigula; artinii; falabae; and hindei; artinii; hirundo; artinii; Scotoéque a ailes noires; Schwarzflligel-Hausfledermaus; Scotoecode alas negras; \\White-bellied Lesser House Bat; Hinde's Lesser House Bat; Dark-winged Lesser House Bat; White-bellied Lesser House Bat; Hinde's Lesser House Bat; Dark-winged Lesser House Bat; Dark-winged Lesser House Bat; S. hirundo