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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1516	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nycticeius rueppelli	Nycticeius rueppellii	Scoteanax rueppellii	Nycticeius rueppellii	Nycticeius rueppellii	Scoteanax rueppellii	Scoteanax rueppellii	Scoteanax rueppellii	Scoteanax rueppellii	Scoteanax rueppellii	Scoteanax rueppellii	Scoteanax rueppellii	Scoteanax rueppellii	Scoteanax rueppellii	Scoteanax rueppellii		[MSW2] Subgenus Scoteanax. Reviewed by Kitchener and Caputi (1985).; [MSW3] Reviewed by Kitchener and Caputi (1985).; [HMW] Nyticejus ruppelliz Peters, 1866 , “Sydney,” New South Wales , Australia . Phylogenetic position of S. rueppellii is currently unknown because it has not been included in any broad-scale phylogenetic studies; it is tentatively included under Eptesicini. Monotypic.; [batnames2022] Reviewed by Kitchener and Caputi (1985).; [batnames2023] Reviewed by Kitchener and Caputi (1985).; [batnames2025_1.7] Reviewed by Kitchener and Caputi (1985).									australis			rueppellii	rueppellii - australis	rueppellii, australis		rueppellii	rueppellii - australis	rueppellii, australis	rueppellii, australis, rueppelli	rueppellii 	rueppellii - australis	rueppellii (W. C. H. Peters, 1866)|rueppelli (J. D. Ogilby, 1892) [incorrect subsequent spelling]|australis (J. MacGillivray in Iredale, 1937)|ruppellii Troughton, 1944 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Rüppell's broad-nosed bat	E Queensland, E New South Wales	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Nycticeius rueppellii	Australia, New South Wales, Sydney.	Peters	1866	Monatsb. Preuss. Akad. Wiss. Berlin, p. 21.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Riippell's broad-nosed bat (Greater broad-nosed bat)	E Queensland, E New South Wales	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1866	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1866:21.	Subgenus Scoteanax. Reviewed by Kitchener and Caputi (1985).	E Queensland and E New South Wales (Australia).	Australia, New South Wales, Sydney.		PETERS	1866	Size relatively large (forearm length, 50-56 mm).	Distribution: Same as for genus.	No subspecies.		126	species	N. rueppellii	PETERS	1866	Scoteanax	subgenus	Nycticeius rueppellii				Size relatively large (forearm length, 50-56 mm).	No subspecies.		6. N. rueppellii PETERS 1866.	6	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nycticeiini	Scoteanax rueppellii	Scoteanax		rueppellii	Peters	y	1866		Monatsb. K. Preuss. Akad. Wiss. Berlin	1866		21		Rüppell's Broad-nosed Bat	Australia, New South Wales, Sydney.	E Queensland and E New South Wales (Australia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt) as Nycticeius ruppellii.	australis Iredale (ex MacGillivray), 1937.	Reviewed by Kitchener and Caputi (1985).	4C3D87E8FFFF6A40FF5294841DE4B0CE	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	837	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFB46A0BFF7D9B4C184EB2A2.xml	Scoteanax rueppellii	Vespertilionidae	Scoteanax	rueppellii		1866	Sérotine de Rippell @fr | Gro RRe Breitnasenfledermaus @de | Scoteno de Rippell @es | Greater Broad-nosed Bat @en	Nyticejus ruppelliz Peters, 1866 , “Sydney,” New South Wales , Australia . Phylogenetic position of S. rueppellii is currently unknown because it has not been included in any broad-scale phylogenetic studies; it is tentatively included under Eptesicini. Monotypic.	E & SE Queensland and E New South Wales , Australia .	Head-body 63-3— 72-7 mm, tail 44-5-58-5> mm, ear 15-6-17-8 mm, forearm 50-5-56-2 mm; weight 20-39-8 g. Riippell’s Broad-nosed Bat is similar to species of Scotorepens but is much larger. Dorsal pelage is reddish brown to dark brown; ventral pelageis slightly paler. Ears are slender and triangular, with moderately rounded tips and notches on posterior edges. Tragus is triangular because of well-defined lobe midway along posterior edge, and anterior edge is straight or slightly concave. Muzzle is broad and short, with square inflated glands on each side. Ears and membranes are blackish brown, and bare face, forearms, and digits are dark pinkish brown. Uropatagium attaches at calcar and extends to tip of tail. Penis is mostly hairless. Baculum is long (8-5-8-8 mm) and vaguely T-shaped in lateral view, with flat tip, thickened upward turned base, and basal notch running up dorso-ventrally broadened base. Skull is relatively narrow but large; sagittal and lambdoidal crests are well developed, forming prominent occipital helmet; anterior palatal emargination extends to behind upper canines; P* is two-thirds the height of C'; and M” is greatly reduced. Rappell’s Broad-nosed Bat is often confused with the Eastern False Pipistrelle ( Falsistrellus tasmaniensis ), from which it can be identified by having two instead of four upper incisors and absence of gap between canines and incisors. Dental formulais11/3, C 1/1, Pl/2,M 3/3 (x2) =30.	Various habitats including mature coastalforests, rainforests, open woodlands, Melaleuca ( Myrtaceae ) swamp woodlands, wet and dry sclerophyll forests, cleared paddocks with remnanttrees, and tree-lined creeks in open regions from sea level up to ¢. 1200 m . Typically associated with mild winters and rainfall over 600 mm /year.	Riippell’s Broad-nosed Bats forage by aerial-hawking and perchhunting and fly at moderate speeds with limited maneuverability. They are primarily insectivorous and mostly eat beetles but also moths, ants, and large flies. Spiders were also found (20% occurrence) in one study, which was unexpected given morphology and general feeding habits. Although not recorded in the wild, they probably regularly eat other bats; they have been observed circling around researchers while they were handling bats and have been seen chasing Chocolate Wattled Bats ( Chalinolobus morio ) when they emerged from their roosts at dusk. In captivity, they eat other bats and break into adjacent cages to eat them. They have also been seen eating bats caught in mist nets with them, particularlyjuveniles.	Female Riippell’s Broad-nosed Bats give birth to one young in January and form maternity colonies a little before giving birth.	Ruppell’s Broad-nosed Bats are nocturnal and have been found roosting in tree hollows, crevices, roofs of old buildings, and under bark on trunks and dead branches. Call shapeis steep FM/QCF sweep, with characteristic frequencies of 32-36-5 kHz in New South Wales and south-eastern Queensland .	Riippell’s Broad-nosed Bats seem to roost in colonies, and maternity colonies do not have males. They might also roost alone and appear to often switch roosts.	Classified as Least Concern on The IUCN Red List.	Baverstock et al. (1987) | Calaby (1966) | Campbell (2001) | Churchill (2008) | Hoye & Richards (2008) | Kitchener & Caputi (1985) | Lunney & Pennay (2008) | Reinhold, Law et al. (2001) | Wilson (2006) | Wooodside & Long (1984)	https://zenodo.org/record/6398122/files/figure.png	164. Ruppell’s Broad-nosed Bat Scoteanax rueppellii French: Sérotine de Rippell / German: GroRRe Breitnasenfledermaus / Spanish: Scoteno de Rippell Other common names: Greater Broad-nosed Bat Taxonomy. Nyticejus ruppelliz Peters, 1866 , “Sydney,” New South Wales , Australia . Phylogenetic position of S. rueppellii is currently unknown because it has not been included in any broad-scale phylogenetic studies; it is tentatively included under Eptesicini. Monotypic. Distribution. E & SE Queensland and E New South Wales , Australia . Descriptive notes. Head-body 63-3— 72-7 mm, tail 44-5-58-5> mm, ear 15-6-17-8 mm, forearm 50-5-56-2 mm; weight 20-39-8 g. Riippell’s Broad-nosed Bat is similar to species of Scotorepens but is much larger. Dorsal pelage is reddish brown to dark brown; ventral pelageis slightly paler. Ears are slender and triangular, with moderately rounded tips and notches on posterior edges. Tragus is triangular because of well-defined lobe midway along posterior edge, and anterior edge is straight or slightly concave. Muzzle is broad and short, with square inflated glands on each side. Ears and membranes are blackish brown, and bare face, forearms, and digits are dark pinkish brown. Uropatagium attaches at calcar and extends to tip of tail. Penis is mostly hairless. Baculum is long (8-5-8-8 mm) and vaguely T-shaped in lateral view, with flat tip, thickened upward turned base, and basal notch running up dorso-ventrally broadened base. Skull is relatively narrow but large; sagittal and lambdoidal crests are well developed, forming prominent occipital helmet; anterior palatal emargination extends to behind upper canines; P* is two-thirds the height of C'; and M” is greatly reduced. Rappell’s Broad-nosed Bat is often confused with the Eastern False Pipistrelle ( Falsistrellus tasmaniensis ), from which it can be identified by having two instead of four upper incisors and absence of gap between canines and incisors. Dental formulais11/3, C 1/1, Pl/2,M 3/3 (x2) =30. Habitat. Various habitats including mature coastalforests, rainforests, open woodlands, Melaleuca ( Myrtaceae ) swamp woodlands, wet and dry sclerophyll forests, cleared paddocks with remnanttrees, and tree-lined creeks in open regions from sea level up to ¢. 1200 m . Typically associated with mild winters and rainfall over 600 mm /year. Food and Feeding. Riippell’s Broad-nosed Bats forage by aerial-hawking and perchhunting and fly at moderate speeds with limited maneuverability. They are primarily insectivorous and mostly eat beetles but also moths, ants, and large flies. Spiders were also found (20% occurrence) in one study, which was unexpected given morphology and general feeding habits. Although not recorded in the wild, they probably regularly eat other bats; they have been observed circling around researchers while they were handling bats and have been seen chasing Chocolate Wattled Bats ( Chalinolobus morio ) when they emerged from their roosts at dusk. In captivity, they eat other bats and break into adjacent cages to eat them. They have also been seen eating bats caught in mist nets with them, particularlyjuveniles. Breeding. Female Riippell’s Broad-nosed Bats give birth to one young in January and form maternity colonies a little before giving birth. Activity patterns. Ruppell’s Broad-nosed Bats are nocturnal and have been found roosting in tree hollows, crevices, roofs of old buildings, and under bark on trunks and dead branches. Call shapeis steep FM/QCF sweep, with characteristic frequencies of 32-36-5 kHz in New South Wales and south-eastern Queensland . Movements, Home range and Social organization. Riippell’s Broad-nosed Bats seem to roost in colonies, and maternity colonies do not have males. They might also roost alone and appear to often switch roosts. Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. Baverstock et al. (1987), Calaby (1966), Campbell (2001), Churchill (2008), Hoye & Richards (2008), Kitchener & Caputi (1985), Lunney & Pennay (2008), Reinhold, Law et al. (2001), Wilson (2006), Wooodside & Long (1984).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Scoteanax rueppellii	Scoteanax		rueppellii	Peters	1866	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	######	R&uuml;ppell's Broad-nosed Bat	 australis Iredale ( ex MacGillivray), 1937.	Australia, New South Wales, Sydney.	E Queensland and E New South Wales (Australia).	Not listed.	Least Concern	Reviewed by Kitchener and Caputi (1985).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Scoteanax rueppellii	23	Rüppell's Broad-nosed Bat	Greater Broad-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Scoteanax	NA	rueppellii	W. Peters	1866	1						"Sydney," New South Wales, Australia.			rueppellii (W. Peters, 1866)|australis (Iredale, 1937)	NA	NA	Australia	Oceania	Australasia/Oceania	LC	0	0	0	Scoteanax_rueppellii	0	sciname match	Scoteanax_rueppellii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14946	Scoteanax rueppellii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Scoteanax	rueppellii	(Peters, 1866)		20000000	Scoteanax rueppellii	Least Concern		2020	2019-05-30 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, presumed large population, and because it is not known to be declining fast enough to qualify for listing in a threatened category.	This species is found in tropical moist forest, dry and wet sclerophyll woodland, Melaleuca swamp and eucalypt forest. Its roosting habits are poorly known, but it has been found roosting in tree hollows, crevices, and under bark. Females give birth to a single young (Duncan et al. 1999, Hoye and Richards 2008).	It is suspected to be threatened by clearance and fragmentation of suitable forest habitat. In addition to the degradation of foraging habitat, the loss of roosting trees due to logging may be a threat (Duncan et al. 1999).	The species is uncommonly reported from surveys within its range. In New South Wales it has a long term average reporting rate of around 1 per 170 bats observed within its range (Pennay et al. 2011)	Stable	This species is endemic to Australia, where it has a disjunct distribution. It has been recorded around the Atherton tablelands area of north Queensland and in coastal areas from south-east Queensland along the east coast of New South Wales. In the south it typically ranges from sea level to around 500 m asl., along the coast and in coastal ranges. In North Queensland it is found at higher elevations on the Atherton Plateau. The range of this species is not well understood because of identification issues. The species bearing a superficial resemblance, and often being confused with Falsistrellus tasmaniensis in public datasets (D. Lunney pers. comm.).		Terrestrial	The species is present in many protected areas within its range. There is a need to identify and protect important populations of this species. Further studies are needed into the ecology, distribution, abundance, and reproduction of this species.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Scoteanax		rueppellii	Peters	1866	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	########	R&uuml;ppell's Broad-nosed Bat	 australis Iredale ( ex MacGillivray), 1937.	Australia, New South Wales, Sydney.	E Queensland and E New South Wales (Australia).	Not listed.	Least Concern	Reviewed by Kitchener and Caputi (1985).	Scoteanax rueppellii	1005563	23	Rüppell's Broad-nosed Bat	Greater Broad-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Scoteanax	NA	rueppellii	W. Peters	1866	1						"Sydney," New South Wales, Australia.			rueppellii (W. Peters, 1866)|australis (Iredale, 1937)	NA	NA				Australia	Oceania	Australasia/Oceania	LC	0	0	0	Scoteanax_rueppellii	0	sciname match	Scoteanax_rueppellii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Scoteanax_rueppellii	1005563	23	Rüppell's Broad-nosed Bat	Greater Broad-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Scoteanax	NA	rueppellii	W. C. H. Peters	1	Nycticejus Rüppellii	Peters, W.C.H. 1866. Ãœber einige neue oder weniger bekannte Flederthiere. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1866:16-25.	https://www.biodiversitylibrary.org/page/39052567	SMF:MAMM:4284	holotype		"Sydney," New South Wales, Australia.			NA	NA				Australia	Oceania (Continent)	Australasia	LC	0	0	0	Scoteanax_rueppellii	0	sciname match	Scoteanax_rueppellii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Scoteanax		rueppellii	Peters	1866	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	########	R&uuml;ppell's Broad-nosed Bat	australis Iredale (ex MacGillivray), 1937.	Australia, New South Wales, Sydney.	E Queensland and E New South Wales (Australia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14946/22009127/' target='_blank'>Least Concern</a>	Reviewed by Kitchener and Caputi (1985).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Scoteanax rueppellii; Scoteanax rueppellii; Scoteanax rueppellii; Scoteanax rueppellii; Scoteanax rueppellii; Scoteanax rueppellii; australis; australis; rueppellii; australis; Sérotine de Rippell; Gro RRe Breitnasenfledermaus; Scoteno de Rippell; Greater Broad-nosed Bat; Rüppell's Broad-nosed Bat; Greater Broad-nosed Bat; Rüppell's Broad-nosed Bat; R&uuml;ppell's Broad-nosed Bat; S. rueppellii