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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1510	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata	Saccopteryx bilineata		[MSW3] Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998). See Yancey et al. (1998a).; [HMW] Urocryptus bilineatus Temminck, 1838 , “ Surinam [= Suriname ].” Saccopteryx bilineata was originally described in the genus Urocryptus but later recognized as monophyletic in the existing genus Saccopteryx . Bats from Mexico and Central America previously were referred to centralis and those from Trinidad to perspicillifer, but they were considered to be synonymous with S . bilineata byJ. K. Jones, Jr . and. S. Hood in 1993. Monotypic.; [batnames2022] Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998). See Yancey et al. (1998 a ).; [IUCN] Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998).; [batnames2023] Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998). See Yancey et al. (1998 a ).; [batnames2025_1.7] Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998). See Yancey et al. (1998a).						centralis, insignis, perspicillifer.	bilineata, perspicillifer		centralis, insignis, perspicillifer		centralis, perspicillifer	bilineata	bilineata - centralis, insignis, perspicillifer	bilineata, insignis, perspicillifer, centralis	Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998).	bilineata	bilineata - centralis, insignis, perspicillifer	bilineata, insignis, perspicillifer, centralis	bilineata, insignis, perspicillifer, centralis	bilineata 	bilineata - centralis, insignis, perspicillifer	bilineata (Temminck, 1838)|insignis (J. A. Wagner, 1855)|perspicillifer G. S. Miller, 1899|centralis O. Thomas, 1904		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Greater white-lined bat	W, E Mexico – Peru, Brazil	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Saccopteryx bilineata	Surinam.	Temminck	1838	Tijdschr. Nat. Gesch. Physiol., 5:33.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Greater white-lined bat (Sac-winged bat)	W, E Mexico – Bolivia, Brazil; Trinidad	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1838	Tijdschr. Nat. Gesch. Physiol., 5:33.		Jalisco and Veracruz (Mexico) to Bolivia, Guianas, and E Brazil south to Rio de Janiero; Trinidad and Tobago.	Surinam.		TEMMINCK	1838	Size relatively large (forearm length, 41-51 mm). Free edge of plagiopatagium attached to tibia. Two longitudi nal whitish lines on blackish back.	Distribution: Same as for genus.	Two subspecies are currently recognized.	S. b. bilineata (distribution except for northern Venezuela, Trinidad, and Tobago). S. b. perspicillifer (northern Venezuela, Trinidad and Tobago).	46	species	S. bilineata	TEMMINCK	1838	Saccopteryx	genus	Saccopteryx bilineata				Size relatively large (forearm length, 41-51 mm). Free edge of plagiopatagium attached to tibia. Two longitudi nal whitish lines on blackish back.	Two subspecies are currently recognized.		1. S. bilineata (TEMMINCK 1838).	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Emballonurinae		Saccopteryx bilineata	Saccopteryx		bilineata	Temminck	y	1838		Tijdschr. Nat. Gesch. Physiol.	5		33		Greater Sac-winged Bat	Surinam.	Jalisco and Veracruz (Mexico) to Bolivia, Guianas, and E Brazil south to Rio de Janiero; Trinidad and Tobago.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	centralis Thomas, 1904; insignis Wagner, 1855; perspicillifer Miller, 1899.	Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998). See Yancey et al. (1998a).	03D587F2FFD84C13FF7F3AA7F45AF854	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	371	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFD94C13FF153987FE0DFD29.xml	Saccopteryx bilineata	Emballonuridae	Saccopteryx	bilineata		1838	Greater Sac-winged Bat @en | Saccoptère à deux bandes @fr | Grosse Sackflügelfledermaus @de | Sacóptero mayor @es	Urocryptus bilineatus Temminck, 1838 , “ Surinam [= Suriname ].” Saccopteryx bilineata was originally described in the genus Urocryptus but later recognized as monophyletic in the existing genus Saccopteryx . Bats from Mexico and Central America previously were referred to centralis and those from Trinidad to perspicillifer, but they were considered to be synonymous with S . bilineata byJ. K. Jones, Jr . and. S. Hood in 1993. Monotypic.	From C Mexico (Jalisco and Veracruz) S throughout both Pacific and Atlantic slopes of Central America to Colombia, E to E & SE Brazil (S to Rio de Janeiro State), and S to Ecuador, E Peru, W Brazil, and N & NE Bolivia; also on Trinidad and Tobago Is.	Head-body 47-56 mm, tail 16-23 mm, ear 13-17 mm, hindfoot 10-12 mm, forearm 44-48 mm; weight 6-9-3 g. Female Greater Sac-winged Bats are larger than males. Dorsal fur is black, with two prominent, wavy bully stripes extending from neck to rump; venter is dark gray. Dorsal fur is black when pelage is “fresh,” but it grades to deep brown when it becomes worn. Although it is not known if multiple molts occur annually, five of nine individuals observed in Trinidad were molting in August. Ears extend slightly above crown of head. Eyes are large and conspicuous, with brown irises. Radio-metacarpal sacs open along forward edge of propatagium and are especially prominent in males. Flight membranes are black. Uropatagium is slightly hairy at its base and around tail. When fully extended, uropatagium is support by stiff ", long calcars and reaches almost to full extremity of claws on hindfeet. Dental formula of all species of Saccopteryx is 11/3, C 1/1, P 2/2, M 3/3 (x2) = 32.	Lowland evergreen, semideciduous, dry forests, forest edges, villages, and urban areas generally below elevations of 500 m, with a few records up to 1000 m. Greater Sac-winged Bats often forage near streams and in moist areas. They prefer multi-strata evergreen forests and forages in natural and man-made forest clearings, edges, and corridors. In Mexico, it has also been reported in secondary forests, croplands, and grasslands.	The Greater Sac-winged Bat is an aerial insectivore that forages in cluttered spaces. Its diet includes flies, beetles, moths, and even butterflies that are active in late afternoon. Foraging habitats shift seasonally when Greater Sac-winged Bats cue on variation in insect phenology and abundance. They forage alone; individuals usually move from one feeding area to another after c.25-30 minutes.	Gestation of Greater Sac-winged Bats across Mexico and Central America begins near end of dry season, and one young is bom annually at onset of rainy season. Timing of parturition varies with climate across the distribution. Females carry young or leave them at hidden night roosts when they forage. Young are able to fly at two weeks old but continue to suckle for 10-12 weeks. Yearling females disperse to other roosting groups, but young males remain with parental roost groups, waiting their chance to succeed older males.	Greater Sac-winged Bats are crepuscular and can be seen in foraging areas in late afternoon, well before sunset, within shaded forest understories. This primary foraging activity occurs for c .3 hours into the night. A secondary foraging period occurs in early morning, and individuals return to day roosts soon after dawn. Mean flight speed is 5-8 m/s. Roost shelters include dimly lit cave entrances, trees hollows, boles and between plank buttresses, buildings, window screens of inhabited buildings, and undersides of bridges. The Greater Sac-winged Bat capably maintains appreciable temperature differentials with its environment . It usually remains normothermic and is alert and active in daytime roosts. It maintains a resting rate of metabolism that is high compared with small insectivorous bats in other families. It is likely that its diet of small, abundant insects provides enough energy year-round to permit a roosting strategy of alertness and vigilance in dimly lit shelters potentially vulnerable to predators. By maintaining individual distances from roost mates, groups of Greater Sac-winged Bats are able to simultaneously fly as an escape response, which would not be possible for bats that cluster tighdy because individuals on outer edges of a cluster must fly off first and those in the center of a cluster must wait for critical seconds before taking flight. Search-phase echolocation pulses of Greater Sac-winged Bats are relatively long (5-6 milliseconds) and paired. They are characterized by slight rise in frequency at onset of pulse, followed by a long F part, and terminated with slight decrease in frequency. Maximum energy in prey searching echolocation calls varies with geography at 42-50 kHz from populations in Belize, Costa Rica, and Panama. During approach or pursuit phase, calls become shorter, with more pronounced rise in frequency than in search phase. Terminal stage emissions, or feeding buzzes, are even shorter ( c .1-1.7 milliseconds), with less pronounced rise in frequency.	Roosting colonies of Greater Sacwinged Bats average 12 individuals but can range from five to 50 individuals. Larger roosting groups are composed of several male-dominated harems with 1-9 breeding females. Mating system is polygynous; males typically defend harems of 3-5 breeding females and 1-3 m2 of vertical roosting space. Most adult males roost alone or on peripheries of breeding colonies. Males obtain harems by splitting an existing harem or replacing a harem male that has been lost. Males have well-developed wing sacs in their ante-brachial membranes and emit scent while flapping their wings in ritualized mating displays called “salting” that are accompanied by singing. Males spend considerable time during the day cleaning their wing sacs with saliva and urine, which is followed by applying genital secretion to wing sacs with their chins. Natural selection favors smaller male Greater Sac-winged Bats due to energetic demands associated with aerial courtship and defense of harem females, and small breeding males sire more offspring than large males. Greater Sac-winged Bats use a variety of multisyllabic social calls in various contexts including sex-specific signatures and individual signatures in their calls. Isolation calls are used by young to call mothers and by young males to signify subordination to dominant males. Mothers respond to isolation calls of young with directive calls, especially to heel young bats when they begin to fly about the roost. Young vocalize in long bouts of “babbling” to practice social calls. Males produce territorial and courtship songs including whistling components, and males that sing often have greater reproductive success. Screeches are used in aggressive encounters by both sexes. Distress calls are produced during encounters with predators. Fur and skin of Greater Sac-winged Bats are often infested by ectoparasites that might influence condition and health, which include the mites Eutrombicula gpldii, Trombiculida vesperuginis, and. saccopteryx. The endoparasitic tapeworm, Hymenolepis mazanensis , has also been found in Greater Sac-winged Bats. Roosts of Greater Sac-winged Bats can be shared with a great many other species of bats including other species of sympatric emballonurids.	Classified as Least Concern on The IUCN Red List. The Greater Sac-winged Bat has a large distribution and presumably large and stable overall population, and it occupies a large variety of habitats. It occurs in many protected areas throughout its distribution.	Barclay (1983) | Behr & von Helversen (2004) | Behr et al. (2009) | Bradbury & Emmons (1974) | Bradbury & Vehrencamp (1977a, 1977b) | Carter et al. (1981) | Davidson & Wilkinson (2002, 2004) | Genoud & Bonaccorso (1986) | Goodwin & Greenhall (1961) | Heckel & von Helversen (2002, 2003) | Heckel , Achmann & Mayer (2000) | Heckel, Voigt et al. (1999) | Jones & Hood (1993) | Knörnschild & von Helversen (2008) | Knörnschild , Behr & von Helversen (2006) | Knörnschild, Jung et al. (2012) | Pye (1973) | Simmons (2005) | Tannenbaum (1975) | Voigt & von Helversen (1999) | Voigt , Heckel & von Helversen (2006) | Voigt, Heckel & Mayer (2005) | Yancey et al. (1998)	https://zenodo.org/record/3747999/files/figure.png	50 . Greater Sac-winged Bat Saccopteryx bilineata French: Saccoptère à deux bandes / German: Grosse Sackflügelfledermaus / Spanish: Sacóptero mayor Other common names: Greater White-lined Bat Taxonomy . Urocryptus bilineatus Temminck, 1838 , “ Surinam [= Suriname ].” Saccopteryx bilineata was originally described in the genus Urocryptus but later recognized as monophyletic in the existing genus Saccopteryx . Bats from Mexico and Central America previously were referred to centralis and those from Trinidad to perspicillifer, but they were considered to be synonymous with S . bilineata byJ. K. Jones, Jr . and. S. Hood in 1993. Monotypic. Distribution. From C Mexico (Jalisco and Veracruz) S throughout both Pacific and Atlantic slopes of Central America to Colombia, E to E & SE Brazil (S to Rio de Janeiro State), and S to Ecuador, E Peru, W Brazil, and N & NE Bolivia; also on Trinidad and Tobago Is. Descriptive notes. Head-body 47-56 mm, tail 16-23 mm, ear 13-17 mm, hindfoot 10-12 mm, forearm 44-48 mm; weight 6-9-3 g. Female Greater Sac-winged Bats are larger than males. Dorsal fur is black, with two prominent, wavy bully stripes extending from neck to rump; venter is dark gray. Dorsal fur is black when pelage is “fresh,” but it grades to deep brown when it becomes worn. Although it is not known if multiple molts occur annually, five of nine individuals observed in Trinidad were molting in August. Ears extend slightly above crown of head. Eyes are large and conspicuous, with brown irises. Radio-metacarpal sacs open along forward edge of propatagium and are especially prominent in males. Flight membranes are black. Uropatagium is slightly hairy at its base and around tail. When fully extended, uropatagium is support by stiff ", long calcars and reaches almost to full extremity of claws on hindfeet. Dental formula of all species of Saccopteryx is 11/3, C 1/1, P 2/2, M 3/3 (x2) = 32. Habitat . Lowland evergreen, semideciduous, dry forests, forest edges, villages, and urban areas generally below elevations of 500 m, with a few records up to 1000 m. Greater Sac-winged Bats often forage near streams and in moist areas. They prefer multi-strata evergreen forests and forages in natural and man-made forest clearings, edges, and corridors. In Mexico, it has also been reported in secondary forests, croplands, and grasslands. Food and Feeding . The Greater Sac-winged Bat is an aerial insectivore that forages in cluttered spaces. Its diet includes flies, beetles, moths, and even butterflies that are active in late afternoon. Foraging habitats shift seasonally when Greater Sac-winged Bats cue on variation in insect phenology and abundance. They forage alone; individuals usually move from one feeding area to another after c.25-30 minutes. Breeding . Gestation of Greater Sac-winged Bats across Mexico and Central America begins near end of dry season, and one young is bom annually at onset of rainy season. Timing of parturition varies with climate across the distribution. Females carry young or leave them at hidden night roosts when they forage. Young are able to fly at two weeks old but continue to suckle for 10-12 weeks. Yearling females disperse to other roosting groups, but young males remain with parental roost groups, waiting their chance to succeed older males. Activity patterns. Greater Sac-winged Bats are crepuscular and can be seen in foraging areas in late afternoon, well before sunset, within shaded forest understories. This primary foraging activity occurs for c .3 hours into the night. A secondary foraging period occurs in early morning, and individuals return to day roosts soon after dawn. Mean flight speed is 5-8 m/s. Roost shelters include dimly lit cave entrances, trees hollows, boles and between plank buttresses, buildings, window screens of inhabited buildings, and undersides of bridges. The Greater Sac-winged Bat capably maintains appreciable temperature differentials with its environment . It usually remains normothermic and is alert and active in daytime roosts. It maintains a resting rate of metabolism that is high compared with small insectivorous bats in other families. It is likely that its diet of small, abundant insects provides enough energy year-round to permit a roosting strategy of alertness and vigilance in dimly lit shelters potentially vulnerable to predators. By maintaining individual distances from roost mates, groups of Greater Sac-winged Bats are able to simultaneously fly as an escape response, which would not be possible for bats that cluster tighdy because individuals on outer edges of a cluster must fly off first and those in the center of a cluster must wait for critical seconds before taking flight. Search-phase echolocation pulses of Greater Sac-winged Bats are relatively long (5-6 milliseconds) and paired. They are characterized by slight rise in frequency at onset of pulse, followed by a long F part, and terminated with slight decrease in frequency. Maximum energy in prey searching echolocation calls varies with geography at 42-50 kHz from populations in Belize, Costa Rica, and Panama. During approach or pursuit phase, calls become shorter, with more pronounced rise in frequency than in search phase. Terminal stage emissions, or feeding buzzes, are even shorter ( c .1-1.7 milliseconds), with less pronounced rise in frequency. Movements, Home range and Social organization. Roosting colonies of Greater Sacwinged Bats average 12 individuals but can range from five to 50 individuals. Larger roosting groups are composed of several male-dominated harems with 1-9 breeding females. Mating system is polygynous; males typically defend harems of 3-5 breeding females and 1-3 m2 of vertical roosting space. Most adult males roost alone or on peripheries of breeding colonies. Males obtain harems by splitting an existing harem or replacing a harem male that has been lost. Males have well-developed wing sacs in their ante-brachial membranes and emit scent while flapping their wings in ritualized mating displays called “salting” that are accompanied by singing. Males spend considerable time during the day cleaning their wing sacs with saliva and urine, which is followed by applying genital secretion to wing sacs with their chins. Natural selection favors smaller male Greater Sac-winged Bats due to energetic demands associated with aerial courtship and defense of harem females, and small breeding males sire more offspring than large males. Greater Sac-winged Bats use a variety of multisyllabic social calls in various contexts including sex-specific signatures and individual signatures in their calls. Isolation calls are used by young to call mothers and by young males to signify subordination to dominant males. Mothers respond to isolation calls of young with directive calls, especially to heel young bats when they begin to fly about the roost. Young vocalize in long bouts of “babbling” to practice social calls. Males produce territorial and courtship songs including whistling components, and males that sing often have greater reproductive success. Screeches are used in aggressive encounters by both sexes. Distress calls are produced during encounters with predators. Fur and skin of Greater Sac-winged Bats are often infested by ectoparasites that might influence condition and health, which include the mites Eutrombicula gpldii, Trombiculida vesperuginis, and. saccopteryx. The endoparasitic tapeworm, Hymenolepis mazanensis , has also been found in Greater Sac-winged Bats. Roosts of Greater Sac-winged Bats can be shared with a great many other species of bats including other species of sympatric emballonurids. Status and Conservation . Classified as Least Concern on The IUCN Red List. The Greater Sac-winged Bat has a large distribution and presumably large and stable overall population, and it occupies a large variety of habitats. It occurs in many protected areas throughout its distribution. Bibliography. Barclay (1983), Behr & von Helversen (2004), Behr et al. (2009), Bradbury & Emmons (1974), Bradbury & Vehrencamp (1977a, 1977b), Carter et al. (1981), Davidson & Wilkinson (2002, 2004), Genoud & Bonaccorso (1986), Goodwin & Greenhall (1961), Heckel & von Helversen (2002, 2003), Heckel , Achmann & Mayer (2000), Heckel, Voigt et al. (1999), Jones & Hood (1993), Knörnschild & von Helversen (2008), Knörnschild , Behr & von Helversen (2006), Knörnschild, Jung et al. (2012), Pye (1973), Simmons (2005), Tannenbaum (1975), Voigt & von Helversen (1999), Voigt , Heckel & von Helversen (2006), Voigt, Heckel & Mayer (2005), Yancey et al. (1998).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Saccopteryx bilineata	Saccopteryx		bilineata	Temminck	1838	1	Tijdschr. Nat. Gesch. Physiol.	5:33	Greater Sac-winged Bat	 centralis Thomas, 1904; insignis Wagner, 1855; perspicillifer Miller, 1899.	Surinam.	Jalisco and Veracruz (Mexico) to Bolivia, Guianas, and E Brazil south to Rio de Janiero; Trinidad and Tobago.	Not listed.	Least Concern	Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998). See Yancey et al. (1998 a ).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Saccopteryx bilineata	23	Greater Sac-winged Bat	Greater White-lined Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	EMBALLONURINAE	DICLIDURINI	Saccopteryx	NA	bilineata	Temminck	1838	1	Urocryptus_bilineatus	Temminck, C. J. (1838). Over de geslachten Taphozous, Emballonura, Urocryptus en Diclidurus. Tijdschrift voor natuurlijke geschiedenis en physiologie, 5, 33.	https://www.biodiversitylibrary.org/item/48145#page/51/mode/1up	RMNH MAM.17461		"Surinam [= Suriname]."			bilineata (Temminck, 1838)|insignis (J. A. Wagner, 1855)|perspicillifer G. S. Miller, 1899|centralis O. Thomas, 1904	NA	NA	Mexico|Belize|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Saccopteryx_bilineata	0	sciname match	Saccopteryx_bilineata	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19804	Saccopteryx bilineata	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Saccopteryx	bilineata	(Temminck, 1838)	Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998).	20000000	Saccopteryx bilineata	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is Least Concern as it is common, widespread and tolerant to some habitat modification.	This bat usually forages near streams and in moist areas. It prefers multistratal evergreen forest and forage in clearings and forest edges and forest corridors (Eisenberg 1989). Clearings and corridors are both natural and man made. ;In Mexico it has also been reported for secondary forests, croplands and grasslands (de Grammont pers. comm.). ;Roosting colonies average about 12 individuals. It may roost with other species in hollow trees or caves. Males defend harems, they have well-developed wing sacs in their ante-brachial membranes and emit scent while flapping their wings in ritualized combat (Bradbury and Emmons 1974). This species is an aerial insectivore that forages in background cluttered space.	In general, deforestation is a potential threat in the region but this is probably not specific to any species of New World emballonurid bat because none of them have a restricted area of endemism, other than perhaps Balantioperyx infusca and Saccopteryx antioquensis .	This species ;is common in lowland evergreen and semi-deciduous forest and forest edge. It is rare in dry deciduous forest (Reid 1997).	Unknown	This species is found in Central and South America. This species ranges from Jalisco and Veracruz (Mexico) to Bolivia, Guianas, and eastern Brazil south to Rio de Janeiro, Trinidad and Tobago (Simmons 2005). It is widely distributed at low elevations, generally below 500 m asl (Eisenberg 1989).	This species is not used.	Terrestrial	Retention of primary forest is the recommended conservation action. This species occurs in some protected areas, as with most New World emballonurid bats because they are usually widely distributed.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Saccopteryx		bilineata	Temminck	1838	1	Tijdschr. Nat. Gesch. Physiol.	5:33	Greater Sac-winged Bat	 centralis Thomas, 1904; insignis Wagner, 1855; perspicillifer Miller, 1899.	Surinam.	Jalisco and Veracruz (Mexico) to Bolivia, Guianas, and E Brazil south to Rio de Janiero; Trinidad and Tobago.	Not listed.	Least Concern	Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998). See Yancey et al. (1998 a ).	Saccopteryx bilineata	1004794	23	Greater Sac-winged Bat	Greater White-lined Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	EMBALLONURINAE	DICLIDURINI	Saccopteryx	NA	bilineata	Temminck	1838	1	Urocryptus_bilineatus	Temminck, C. J. (1838). Over de geslachten Taphozous, Emballonura, Urocryptus en Diclidurus. Tijdschrift voor natuurlijke geschiedenis en physiologie, 5, 33.	https://www.biodiversitylibrary.org/item/48145#page/51/mode/1up	RMNH MAM.17461		"Surinam [= Suriname]."			bilineata (Temminck, 1838)|insignis (J. A. Wagner, 1855)|perspicillifer G. S. Miller, 1899|centralis O. Thomas, 1904	NA	NA				Mexico|Belize|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Saccopteryx_bilineata	0	sciname match	Saccopteryx_bilineata	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Saccopteryx_bilineata	1004794	23	Greater Sac-winged Bat	Greater White-lined Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Emballonurinae	Diclidurini	Saccopteryx	NA	bilineata	Temminck	1	Urocryptus bilineatus	Temminck, C.J. 1838. Over de geslachten _Taphozous_, _Emballonura_, _Urocryptus_ en _Diclidurus_. Tijdschrift voor Natuurlijke Geschiedenis en Physiologie 5(1-2):1-34.	https://www.biodiversitylibrary.org/page/13474807	RMNH.MAM.17461	holotype	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.17461.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.17461.b	"Surinam [= Suriname]."			NA	NA				Mexico|Belize|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad and Tobago|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Saccopteryx_bilineata	0	sciname match	Saccopteryx_bilineata	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Saccopteryx		bilineata	Temminck	1838	1	Tijdschr. Nat. Gesch. Physiol.	5:33	Greater Sac-winged Bat	centralis Thomas, 1904; insignis Wagner, 1855; perspicillifer Miller, 1899.	Surinam.	Jalisco and Veracruz (Mexico) to Bolivia, Guianas, and E Brazil south to Rio de Janiero; Trinidad and Tobago.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19804/22004716/' target='_blank'>Least Concern</a>	Several subspecies have been recognized, but these do not appear justified; see Simmons and Voss (1998). See Yancey et al. (1998a).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Saccopteryx bilineata; Saccopteryx bilineata; Saccopteryx bilineata; Saccopteryx bilineata; Saccopteryx bilineata; Saccopteryx bilineata; centralis; insignis; perspicillifer; centralis; perspicillifer; centralis; insignis; perspicillifer; bilineata; insignis; perspicillifer; centralis; Greater Sac-winged Bat; Saccoptère à deux bandes; Grosse Sackflügelfledermaus; Sacóptero mayor; Greater Sac-winged Bat; Greater White-lined Bat; Greater Sac-winged Bat; Greater Sac-winged Bat; S. bilineata