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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1505	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Taphozous saccolaimus	Saccolaimus saccolaimus	Taphozous saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus	Saccolaimus saccolaimus		[MSW2] Corbet and Hill (1980:45) listed nudicluniatus as a distinct species without comment.; [MSW3] Corbet and Hill (1980) listed nudicluniatus as a distinct species without comment. Includes pulcher; see Medway (1977) and Goodwin (1979). Includes pluto; see Corbet and Hill (1992). Reviewed in part by Chimimba and Kitchener (1991) and Bates and Harrison (1997); also see Flannery (1995b) and Bonaccorso (1998).; [HMW] Taphozous saccolaimus Temminck, 1838 , Java , Indonesia . There are differing opinions regarding taxonomic status of S . saccolaimus in Australia, and clarification is required.. W. De Vis in 1905 first described Australian forms as a full species Taphozous nudicluniatus . Subsequently, it was placed in the genus Saccolaimus based on cranial, dental, and external morphology. R. E. Goodwin in 1979 and. T. Chimimba and D. J. Kitchener in 1991 synonymized it with Taphozous (= Saccolaimus ) saccolaimus . Some authors have considered nudicluniatus to be subspecies of widespread S . saccolaimus , but validity of subspecific status of nudicluniatus has been questioned. Melanesian and Australian populations are referred to subspecies nudicluniatus , and it is not clear if the Northern Territory population is part of the nominate form, although mtDNA studies showed limited differences between Northern Territory and Queensland groups. Five subspecies recognized.; [batnames2022] Corbet and Hill (1980) listed nudicluniatus as a distinct species without comment. Includes pulcher ; see Medway (1977) and Goodwin (1979). Includes pluto ; see Corbet and Hill (1992). Reviewed in part by Chimimba and Kitchener (1991) and Bates and Harrison (1997); also see Flannery (1995 b ) and Bonaccorso (1998).; [IUCN] Five valid subspecies are recognised currently within Saccolaimus saccolaimus : affinis (Borneo), crassus (South Asia and mainland South East Asia), nudicluniatus (Melanesia, Australia, possibly Timor Island), pluto (Phillippines), plus the nominate (Java, Sumatra) (Simmons 2005). The Melanesian and Australian populations are referred to the subspecies S. s. nudicluniatus , and it is not clear whether the Northern Territory population is part of the nominate race, though mitochondrial DNA studies showed limited differences between Northern Territory and Queensland groups (Milne et al . 2009). Taxonomic studies are ongoing (K.N. Armstrong unpublished).; [batnames2023] Corbet and Hill (1980) listed nudicluniatus as a distinct species without comment. Includes pulcher ; see Medway (1977) and Goodwin (1979). Includes pluto ; see Corbet and Hill (1992). Reviewed in part by Chimimba and Kitchener (1991) and Bates and Harrison (1997); also see Flannery (1995 b ) and Bonaccorso (1998). Synonymies remain unclear for several names; see Jackson and Groves (2015) for a summary.; [batnames2025_1.7] Corbet and Hill (1980) listed nudicluniatus as a distinct species without comment. Includes pulcher; see Medway (1977) and Goodwin (1979). Includes pluto; see Corbet and Hill (1992). Reviewed in part by Chimimba and Kitchener (1991) and Bates and Harrison (1997); also see Flannery (1995b) and Bonaccorso (1998). Synonymies remain unclear for several names; see Jackson and Groves (2015) for a summary.				affinis, flavomaculatus, nudicluniatus	(nudicluniatus)	affinis, crassus, flavomaculatus, granti, nudicluniatus, pulcher (see Medway, 1977:45, and Goodwin, 1979:102).	crassus, saccolaimus, affinis, nudicluniatus	saccolaimus, affinis, crassus, nudicluniatus, pluto	affinis - flavimaculatus; crassus - pulcher; nudicluniatus - granti; pluto - capito	sacco, affinis, crassus, nudicluniatus, pluto		saccolaimus, affinis, crassus, nudicluniatus	affinis - flavimaculatus; crassus - pulcher; nudicluniatus - granti, pluto, capito	saccolaimus, crassus, pulcher, affinis, nudicluniatus, pluto, granti, capito, flavimaculatus	Five valid subspecies are recognised currently within Saccolaimus saccolaimus : affinis (Borneo), crassus (South Asia and mainland South East Asia), nudicluniatus (Melanesia, Australia, possibly Timor Island), pluto (Phillippines), plus the nominate (Java, Sumatra) (Simmons 2005). The Melanesian and Australian populations are referred to the subspecies S. s. nudicluniatus , and it is not clear whether the Northern Territory population is part of the nominate race, though mitochondrial DNA studies showed limited differences between Northern Territory and Queensland groups (Milne et al . 2009). Taxonomic studies are ongoing (K.N. Armstrong unpublished).	saccolaimus, affinis, crassus, nudicluniatus, pluto	affinis - flavimaculatus; crassus - pulcher; nudicluniatus - granti; pluto - capito	saccolaimus, crassus, pulcher, affinis, nudicluniatus, pluto, granti, capito, flavimaculatus	saccolaimus, crassus, pulcher, affinis, nudicluniatus, pluto, granti, capito, flavimaculatus		affinis, flavimaculatus, crassus, pulcher, nudicluniatus, granti, pluto, capito	saccolaimus (Temminck, 1838)|crassus (E. Blyth, 1844)|pulcher (W. Elliot, 1844)|affinis (Dobson, 1875)|saccolaemus (de Seabra, 1900) [incorrect subsequent spelling]|nudicluniatus (De Vis, 1905)|pluto (G. S. Miller, 1910)|granti (O. Thomas, 1911)|capito (Hollister, 1913)|flavimaculatus Sody, 1931		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		India – Java, Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Saccolaimus saccolaimus	Indonesia, Java.	Temminck	1838	Tijdschr. Nat. Gesch. Physiol., 5:14.	Distribution: From India and Ceylon through southeastern Asia and the Malay archipelago (but not the Phi lippines) to Guadalcanal in the Solomons and to northeastern Queensland and northern Northern Territory in Australia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		India – Sulawesi, Timor, New Guinea, Solomon Is, N, NE Australia	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1838	Tijdschr. Nat. Gesch. Physiol., 5:14.	Corbet and Hill (1980:45) listed nudicluniatus as a distinct species without comment.	India and Sri Lanka through SE Asia to Borneo, Sumatra, Java, and Timor (Indonesia); New Guinea; NE Queensland (Australia); Guadalcanal Isl (Solomon Isis).	Indonesia, Java.		TEMMINCK	1841	Posterior floor of mesopterygoid fossa not deeply grooved. Basisphenoid pits separated by a relatively low septum. Size medium (forearm length, 67-76 mm). Radio-metacarpal pouch virtually absent. Sagittal crest relatively high with occipital "helmet" usually well developed.	Distribution: From India and Ceylon through southeastern Asia and the Malay archipelago (but not the Phi lippines) to Guadalcanal in the Solomons and to northeastern Queensland and northern Northern Territory in Australia.	Four currently recog nized subspecies:	S. s. crassus (Asian mainland and Sumatran range), S. s. saccolaimus (Java), S. s. affinis (= flavomaculatus) (Borneo), S. s. nudicluniatus (New Guinea, Guadalcanal, northeastern Queensland). The Celebes, Timor, and North ern Territory populations have not been allocated to sub species.	43	species	S. saccolaimus	TEMMINCK	1841	Saccolaimus	genus	Saccolaimus saccolaimus				Posterior floor of mesopterygoid fossa not deeply grooved. Basisphenoid pits separated by a relatively low septum. Size medium (forearm length, 67-76 mm). Radio-metacarpal pouch virtually absent. Sagittal crest relatively high with occipital "helmet" usually well developed.	Four currently recog nized subspecies:		3. S. saccolaimus (TEMMINCK 1841).	3	_S. s. affinis_ (Dobson, 1875) (synonyms: _flavimaculatus_ Sody, 1931); _S. s. crassus_ (Blyth, 1844) (synonyms: _pulcher_ (Elliot, 1844)); _S. s. nudicluniatus_ (De Vis, 1905) (synonyms: _granti_ (Thomas, 1911)); _S. s. pluto_ (Miller, 1910) (synonyms: _capito_ (Hollister, 1913)); _S. s. saccolaimus_ (Temminck, 1838)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Taphozoinae		Saccolaimus saccolaimus	Saccolaimus		saccolaimus	Temminck	y	1838		Tijdschr. Nat. Gesch. Physiol.	5		14		Naked-rumped Pouched Bat	Indonesia, Java.	Bangladesh, India, and Sri Lanka through SE Asia (including Burma, Cambodia, Thailand, and the Nicobar Isls) to the Philippines, Sulawesi, and Borneo, Sumatra, Java, Bali, and Timor (Indonesia); New Guinea; New Britain and Bougainville Isls (Papua New Guinea); NE Queensland (Australia); Guadalcanal Isl (Solomon Isls).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	affinis Dobson, 1875; flavimaculatus Sody, 1931; crassus Blyth, 1844; pulcher Blyth, 1844; nudicluniatus De Vis, 1905; granti Thomas, 1911; pluto Miller, 1910; capito Hollister, 1913.	Corbet and Hill (1980) listed nudicluniatus as a distinct species without comment. Includes pulcher; see Medway (1977) and Goodwin (1979). Includes pluto; see Corbet and Hill (1992). Reviewed in part by Chimimba and Kitchener (1991) and Bates and Harrison (1997); also see Flannery (1995b) and Bonaccorso (1998).	03D587F2FFD84C13F82C3D5AF55FEEEE	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	356	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFC84C0CF82C33C3F98CF8C6.xml	Saccolaimus saccolaimus	Emballonuridae	Saccolaimus	saccolaimus	Temminck	1838	16 @en | Bare-rumped Sheath-tailed Bat @en | Taphien de Temminck @fr | Nacktbauch-Glattnasenfreischwanz @de | Tafozo de Temminck @es | Naked-rumped ree-tail Bat @en | Naked-rumped Pouched Bat @en | Naked-rumped Sheath-tailed Bat @en	Taphozous saccolaimus Temminck, 1838 , Java , Indonesia . There are differing opinions regarding taxonomic status of S . saccolaimus in Australia, and clarification is required.. W. De Vis in 1905 first described Australian forms as a full species Taphozous nudicluniatus . Subsequently, it was placed in the genus Saccolaimus based on cranial, dental, and external morphology. R. E. Goodwin in 1979 and. T. Chimimba and D. J. Kitchener in 1991 synonymized it with Taphozous (= Saccolaimus ) saccolaimus . Some authors have considered nudicluniatus to be subspecies of widespread S . saccolaimus , but validity of subspecific status of nudicluniatus has been questioned. Melanesian and Australian populations are referred to subspecies nudicluniatus , and it is not clear if the Northern Territory population is part of the nominate form, although mtDNA studies showed limited differences between Northern Territory and Queensland groups. Five subspecies recognized.	S . s. sacco / awnus Temminck, 1838 — Sumatra andJava . S . s. affinis Dobson, 1875 — Borneo and offshore islands including Labuan I. S . s. crassus Blyth, 1844 — S & SE Asia from India (including Great Nicobar I) and Sri Lanka to Indochina and Peninsular Malaysia. S . s. nudicluniatus De Vis, 1905 - New Guinea (including Yapen and Trobriand Is), Bismarck Archipelago (New Britain I), Solomon Is (Bougainville and Guadalcanal), and N & NE Australia (Top End and NE Queensland); possibly also on Timor I. S . s. pluto G. S. Miller, 1910 — Philippines (Luzon, Catanduanes, Mindoro, Negros, and Mindanao Is). Present also in Sulawesi, Moluccas (Halmahera I), Talaud Is, and Ternate I, but subspecies to which these populations belong are unknown. Distribution and boundaries of different subspecies are poorly known.	Head—body 88—100 mm, tail 22—35 mm, ear 16-21 mm, hindfoot 15-17 mm, forearm 68-78 mm; weight 40-50 g. Dorsal fur of the Bare-rumped Sheathtailed Bat is reddish brown to dark brown and irregularly flecked with white patches. Venter is white or less frequently dark brown. Rump, legs, and feet are naked. Males have well-developed glandular gular pouch, which is rudimentary in females. Radialmetacarpal wing pouches are poorly developed. Short, broadly rounded ear is heavily ribbed on interior of pinna. Tragus is short, and its margin is semicircular. Wings are long and narrow. There might be white translucent parts on black-pigmented wings. Other exposed body skin including naked legs and feet is reddish, brown, or black. Strong sagittal crest on skull diverges in postorbital region. Upper incisors are minute.	Dry woodlands to rainforests from sea level to elevations of c .1200 m. In Australia, the Bare-rumped Sheath-tailed Bat is restricted to forested habitats and frequently occurs in hollow limbs of Eucalyptus alba ( Myrtaceae ).	The Bare-rumped Sheath-tailed Bat preys on insects high over gardens, woodlands, and rainforests. Audible low-frequency echolocation clicks are produced while foraging.	In Australia, parturition of Bare-rumped Sheath-tailed Bats occurs at beginning of wet season, and lactation occurs later in wet season ( November—April in Northern Territory). Females give birth to one young. Three to four individuals are often found in tree hollows in Australia. Individual spacing is maintained in roosts.	The Bare-rumped Sheath-tailed Bat roosts in tree hollows, rock crevices , caves, and tombs; between large boulders; and under eaves of houses. Flight path is high and fest during foraging. Bare-rumped Sheath-tailed Bats are alert in roosts, clinging to substrates while supported by thumbs, wrists, and feet. They scurry over roost substrate if disturbed. Echolocation calls are multiharmonic. There is sharp curvilinear downward sweep in frequency at end of long duration calls. Individuals from northern Australia have characteristic frequencies of 19-4-23-4 kHz (mean 20-8 kHz); whereas, in Brunei, frequency bands are 24-5-26-1 kHz. Calls have long durations of 18-25 milliseconds.	Colonies of Bare-rumped Sheathtailed Bats range from 3-4 individuals to hundreds of individuals. Contextual use of secretions from throat pouches of males is unknown, but they likely are used in social attraction of breeding females.	Classified as Least Concern on The IUCN Red List. The Bare-rumped Sheath-tailed Bat has a large distribution and presumably large and stable overall population, uses multiple habitats, occurs in protected areas, faces no significant threats, and does not show evidence of declines affecting significant parts of the population. In Australia, it is listed as critically endangered under the Commonwealth Environment Protection and Biodiversity Conservation Act of 1999. It is listed as endangered under the Queensland Nature Conservation Act of 1992. In the Northern Territory , it is listed as data deficient under the Territory Parks and Wildlife Conservation Act of 2000. Surveys to find roosts and echolocation surveys to better understand distribution and foraging behavior would significantly help to increase knowledge of ecology , behavior, and regional population trends.	Bates, Harrison & Muni (1994a, 1994b, 1994c) | Bonaccorso (1998) | Brosset (1963) | Chimimba & Kitchener (1991) | Corbet & Hill (1992) | De Vis (1905) | lannery (1995a, (1995b) | Goodwin (1979) | Hall, Thomson & Milne (2008) | Heaney, Balete et al. (1998) | Heaney, Gonzales et al. (1991) | Koopman (1984b) | Lekagul & McNeely (1977) | Milne (2002) | Milne, Jackling et al. (2009) | Milne, Reardon &Watt (2003) | Molur et al. (2002) | Sarker & Sarker (2005) | Schulz & Thomson (2007) | Simmons (2005) | Srinivasulu & Srinivasulu (2005)	https://zenodo.org/record/3747934/files/figure.png	16 . Bare-rumped Sheath-tailed Bat Saccolaimus saccolaimus French:Taphien deTemminck /German: Nacktbauch-Glattnasenfreischwanz/ Spanish: Tafozo deTemminck Other common names: Naked-rumped ree-tail Bat, Naked-rumped Pouched Bat, Naked-rumped Sheath-tailed Bat, Naked-tailed Saccolaimus, Pouch-bearing Tomb Bat, PouchedTomb Bat Taxonomy . Taphozous saccolaimus Temminck, 1838 , Java , Indonesia . There are differing opinions regarding taxonomic status of S . saccolaimus in Australia, and clarification is required.. W. De Vis in 1905 first described Australian forms as a full species Taphozous nudicluniatus . Subsequently, it was placed in the genus Saccolaimus based on cranial, dental, and external morphology. R. E. Goodwin in 1979 and. T. Chimimba and D. J. Kitchener in 1991 synonymized it with Taphozous (= Saccolaimus ) saccolaimus . Some authors have considered nudicluniatus to be subspecies of widespread S . saccolaimus , but validity of subspecific status of nudicluniatus has been questioned. Melanesian and Australian populations are referred to subspecies nudicluniatus , and it is not clear if the Northern Territory population is part of the nominate form, although mtDNA studies showed limited differences between Northern Territory and Queensland groups. Five subspecies recognized. Subspecies and Distribution. S . s. sacco / awnus Temminck, 1838 — Sumatra andJava . S . s. affinis Dobson, 1875 — Borneo and offshore islands including Labuan I. S . s. crassus Blyth, 1844 — S & SE Asia from India (including Great Nicobar I) and Sri Lanka to Indochina and Peninsular Malaysia. S . s. nudicluniatus De Vis, 1905 - New Guinea (including Yapen and Trobriand Is), Bismarck Archipelago (New Britain I), Solomon Is (Bougainville and Guadalcanal), and N & NE Australia (Top End and NE Queensland); possibly also on Timor I. S . s. pluto G. S. Miller, 1910 — Philippines (Luzon, Catanduanes, Mindoro, Negros, and Mindanao Is). Present also in Sulawesi, Moluccas (Halmahera I), Talaud Is, and Ternate I, but subspecies to which these populations belong are unknown. Distribution and boundaries of different subspecies are poorly known. Descriptive notes. Head—body 88—100 mm, tail 22—35 mm, ear 16-21 mm, hindfoot 15-17 mm, forearm 68-78 mm; weight 40-50 g. Dorsal fur of the Bare-rumped Sheathtailed Bat is reddish brown to dark brown and irregularly flecked with white patches. Venter is white or less frequently dark brown. Rump, legs, and feet are naked. Males have well-developed glandular gular pouch, which is rudimentary in females. Radialmetacarpal wing pouches are poorly developed. Short, broadly rounded ear is heavily ribbed on interior of pinna. Tragus is short, and its margin is semicircular. Wings are long and narrow. There might be white translucent parts on black-pigmented wings. Other exposed body skin including naked legs and feet is reddish, brown, or black. Strong sagittal crest on skull diverges in postorbital region. Upper incisors are minute. Habitat . Dry woodlands to rainforests from sea level to elevations of c .1200 m. In Australia, the Bare-rumped Sheath-tailed Bat is restricted to forested habitats and frequently occurs in hollow limbs of Eucalyptus alba ( Myrtaceae ). Food and Feeding . The Bare-rumped Sheath-tailed Bat preys on insects high over gardens, woodlands, and rainforests. Audible low-frequency echolocation clicks are produced while foraging. Breeding . In Australia, parturition of Bare-rumped Sheath-tailed Bats occurs at beginning of wet season, and lactation occurs later in wet season ( November—April in Northern Territory). Females give birth to one young. Three to four individuals are often found in tree hollows in Australia. Individual spacing is maintained in roosts. Activity patterns. The Bare-rumped Sheath-tailed Bat roosts in tree hollows, rock crevices , caves, and tombs; between large boulders; and under eaves of houses. Flight path is high and fest during foraging. Bare-rumped Sheath-tailed Bats are alert in roosts, clinging to substrates while supported by thumbs, wrists, and feet. They scurry over roost substrate if disturbed. Echolocation calls are multiharmonic. There is sharp curvilinear downward sweep in frequency at end of long duration calls. Individuals from northern Australia have characteristic frequencies of 19-4-23-4 kHz (mean 20-8 kHz); whereas, in Brunei, frequency bands are 24-5-26-1 kHz. Calls have long durations of 18-25 milliseconds. Movements, Home range and Social organization. Colonies of Bare-rumped Sheathtailed Bats range from 3-4 individuals to hundreds of individuals. Contextual use of secretions from throat pouches of males is unknown, but they likely are used in social attraction of breeding females. Status and Conservation . Classified as Least Concern on The IUCN Red List. The Bare-rumped Sheath-tailed Bat has a large distribution and presumably large and stable overall population, uses multiple habitats, occurs in protected areas, faces no significant threats, and does not show evidence of declines affecting significant parts of the population. In Australia, it is listed as critically endangered under the Commonwealth Environment Protection and Biodiversity Conservation Act of 1999. It is listed as endangered under the Queensland Nature Conservation Act of 1992. In the Northern Territory , it is listed as data deficient under the Territory Parks and Wildlife Conservation Act of 2000. Surveys to find roosts and echolocation surveys to better understand distribution and foraging behavior would significantly help to increase knowledge of ecology , behavior, and regional population trends. Bibliography. Bates, Harrison & Muni (1994a, 1994b, 1994c), Bonaccorso (1998), Brosset (1963), Chimimba & Kitchener (1991), Corbet & Hill (1992), De Vis (1905), lannery (1995a, (1995b), Goodwin (1979), Hall, Thomson & Milne (2008), Heaney, Balete et al. (1998), Heaney, Gonzales et al. (1991), Koopman (1984b), Lekagul & McNeely (1977), Milne (2002), Milne, Jackling et al. (2009), Milne, Reardon &Watt (2003), Molur et al. (2002), Sarker & Sarker (2005), Schulz & Thomson (2007), Simmons (2005), Srinivasulu & Srinivasulu (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Saccolaimus saccolaimus	Saccolaimus		saccolaimus	Temminck	1838	1	Tijdschr. Nat. Gesch. Physiol.	5:14	Naked-rumped Pouched Bat	<b> affinis </b>Dobson, 1875; flavimaculatus Sody, 1931; <b> crassus </b>Blyth, 1844; pulcher Blyth, 1844; <b> nudicluniatus </b>De Vis, 1905; granti Thomas, 1911; <b> pluto </b>Miller, 1910; capito Hollister, 1913.	Indonesia, Java.	Bangladesh, India, and Sri Lanka through SE Asia (including Burma, Cambodia, Thailand, and the Nicobar Isls) to the Philippines, Sulawesi, and Borneo, Sumatra, Java, Bali, and Timor (Indonesia); New Guinea; New Britain and Bougainville Isls (Papua New Guinea); NE Queensland (Australia); Guadalcanal Isl (Solomon Isls).	Not listed.	Least Concern	Corbet and Hill (1980) listed nudicluniatus as a distinct species without comment. Includes pulcher ; see Medway (1977) and Goodwin (1979). Includes pluto ; see Corbet and Hill (1992). Reviewed in part by Chimimba and Kitchener (1991) and Bates and Harrison (1997); also see Flannery (1995 b ) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Saccolaimus saccolaimus	23	Bare-rumped Sheath-tailed Bat	Naked-rumped Free-tail Bat|Naked-rumped Pouched Bat|Naked-rumped Sheath-tailed Bat|Naked-tailed Saccolaimus|Pouch-bearing Tomb Bat|Pouched Tomb Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	TAPHOZOINAE	NA	Saccolaimus	NA	saccolaimus	Temminck	1838	1						Java, Indonesia.			saccolaimus (Temminck, 1838)|crassus (Blyth, 1844)|pulcher (Blyth, 1844)|affinis (Dobson, 1875)|nudicluniatus (De Vis, 1905)|pluto (G. S. Miller, 1910)|granti (O. Thomas, 1911)|capito (Hollister, 1913)|flavimaculatus Sody, 1931	NA	NA	India|Sri Lanka|Bangladesh|Myanmar|Thailand|Cambodia|Vietnam|Malaysia|Singapore|Nicobar Islands|Indonesia|Brunei|East Timor|Philippines|Papua New Guinea|Solomon Islands|Australia	Asia|Oceania	Indomalaya	LC	0	0	0	Saccolaimus_saccolaimus	0	sciname match	Saccolaimus_saccolaimus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19802	Saccolaimus saccolaimus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Saccolaimus	saccolaimus	(Temminck, 1838)	Five valid subspecies are recognised currently within Saccolaimus saccolaimus : affinis (Borneo), crassus (South Asia and mainland South East Asia), nudicluniatus (Melanesia, Australia, possibly Timor Island), pluto (Phillippines), plus the nominate (Java, Sumatra) (Simmons 2005). The Melanesian and Australian populations are referred to the subspecies S. s. nudicluniatus , and it is not clear whether the Northern Territory population is part of the nominate race, though mitochondrial DNA studies showed limited differences between Northern Territory and Queensland groups (Milne et al . 2009). Taxonomic studies are ongoing (K.N. Armstrong unpublished).	200000000	Saccolaimus saccolaimus	Least Concern		2017	2016-07-31 00:00:00 UTC	3.1	English	<p><span lang="EN-AU">This bat is listed as Least Concern given its wide distribution, use of a broad range of habitats, presumed large population size in at least some parts of its range, its occurrence in protected areas, and the absence of significant key threats or evidence for a decline that would affect a significant proportion of the population. ;</p>	<p><span lang="EN-AU">The ecology of S. saccolaimus varies somewhat according to subspecies and region. In the Nicobar Islands in India, it is found in dense forests near the coast. In Sri Lanka, it has been recorded from dense forests, swamp habitat and plantations. There, diurnal roosts include hollows of old and decaying trees including Kitul Palm and Arecanut Palm, old buildings and rocky crevices in small colonies of five or six. No sexual segregation is observed in roosts. This species is known to feed on termites, beetles and other insects and sometimes forages close to the ground in open areas. It is also known to fly fast and high, up to 300 – 400 m from ground, and over forest canopy. It is recorded to emerge very early in the evening from its roost to forage (Bates and Harrison 1997). In South East Asia, the species is commonly observed over modified habitats, including agricultural areas. In some parts of Asia, roosts have been observed in buildings and shallow caves, sometimes in large groups. In the Philippines, roosts have mainly been observed in hollow coconut trees (L. Heaney pers. comm. 2006). In Australia and Melanesia S. saccolaimus has been recorded in dry open sclerophyll woodland, tall open eucalypt forests, dense tropical moist forest, open Pandanus woodland, and coastal lowlands. It roosts mainly in tree hollows, with colonies ranging in size from a few individuals to several hundred animals. Females gives birth to a single young (Flannery 1995a,b; Bonaccorso 1998; Hall et al. 2008;). ;</p>	<p><span lang="EN-AU">There are no major threats to this widespread and adaptable species as a whole. In South Asia, it is locally threatened by deforestation, generally resulting from logging operations and from conversion of land to agricultural and other uses (Molur <span lang="EN-AU">et al .<span lang="EN-AU"> 2002). In Australia, it is threatened in coastal Queensland by clearance of coastal tropical woodland and inappropriate burning regimes, which might affect roost availability (Woinarski et al . 2014). ;</span></p>	<p><span lang="EN-AU">In general, this is a common species. In South Asia, the population status is considered to be poorly known, but according to Bates and Harrison (1997) it is abundant. In the Philippines, it is poorly known, but might be moderately common in agricultural areas (Heaney <span lang="EN-AU">et al .<span lang="EN-AU"> 1998), and is probably common in populated areas, but this require confirmation as there has been little direct survey work undertaken in these modified habitats (L. Heaney pers. comm.). It is common locally in other parts of its Southeast Asian range outside of the Philippines. In Australia, it is present at relatively low density, and while the number of records has increased in the last two decades, the overall verifiable observations are rela<a name="_GoBack"></a>tively low and many museum specimens are the result of collections derived from felled trees. Echolocation calls that might be attributable to S. saccolaimus have been recorded on several occasions in Papua New Guinea (Sandaun, Gulf, Western, Morobe and Southern Highlands provinces; K.N. Armstrong and K.P. Aplin unpublished data), suggesting it might be widespread, but capture is required for verification.</span></p>	Stable	<span lang="EN-AU">This widespread species ranges from South Asia, through parts of continental and Island South East Asia, Melanesia and Australia. In South Asia, it is presently known from Bangladesh (Sylhet division) (Sarker and Sarker 2005, Srinivasulu and Srinivasulu 2005), India (Andaman and Nicobar Islands, Gujarat, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Meghalaya, Orissa, Uttar Pradesh and West Bengal) and Sri Lanka (North Central, Uva and Western provinces) (Molur <span lang="EN-AU">et al <span lang="EN-AU">. 2002). In South Asia, it has been recorded from sea level to 1,200 m asl (Molur <span lang="EN-AU">et al .<span lang="EN-AU"> 2002). It has been recorded from continental South East Asia in Myanmar, southern Thailand, Cambodia (known only from a collection in Phnom Phen; G. Csorba pers. comm.), south-western Viet Nam, Peninsular Malaysia and possibly Singapore. Within Island South East Asia, S. saccolaimus has been recorded from the islands of Sumatra (Indonesia) and Java (type locality: Indonesia), Borneo (Indonesia and Malaysia only), Sulawesi (Indonesia), the island of Timor (Indonesia and Timor-Leste), Halmahera (Indonesia), the Talaud Islands (Indonesia) and Ternate Island (Indonesia), and the Philippines. In the Philippines it has been recorded from sea level to 800 m asl (Heaney <span lang="EN-AU">et al. <span lang="EN-AU"> 1998) from Catanduanes, Luzon and Mindoro (Corbet and Hill 1992), Mindanao (Misamis Oriental, Zamboanga del Sur provinces), and Negros (Heaney <span lang="EN-AU">et al .<span lang="EN-AU"> 1998) although it is likely to occur throughout the country except for the Batanes/Babuyan region (L. Heaney pers. comm. 2006). It has been recorded from scattered localities on the island of New Guinea (Indonesia and Papua New Guinea), on Yapen Island (Papua Province, Indonesia), on the Bismarck Archipelago and the Trobriand Islands (Papua New Guinea), on Bougainville Island (Papua New Guinea). It has also been recorded from the island of Guadalcanal in the Solomon Islands and from the Top End of the Northern Territory and the eastern coast of northern Queensland in Australia (Corbet and Hill 1992; Flannery 1995a,b; Bonaccorso 1998; Hall et al . 2008).</span></span></span></span>		Terrestrial	<p><span lang="EN-AU">Saccolaimus saccolaimus <span lang="EN-AU"> has been recorded from many protected areas, and over much of its range no direct conservation actions are needed. In South Asia, the species has been recorded from protected areas such as Campbell Bay National Park (Andaman and Nicobar Islands) and Kanha National Park (Madhya Pradesh). Further studies are needed on distribution, abundance, breeding biology and general ecology of this species. In South Asia, populations of this species should be monitored to record changes in abundance and distribution. There is also a need to identify populations that are being threatened by human induced habitat alterations in order to develop mitigation measures (C. Srinivasulu pers. comm.). It has been recorded from several protected areas in Australia, however, there is a need to identify and protect important roosting and foraging sites for the species. Further studies are needed in Australia into the distribution, abundance, natural history and threats to this species. Work to better characterise its echolocation call in the context of other low-frequency emitting bats would help environmental assessments. Taxonomic studies focussing on the Australasian populations need to be completed. ;</span></p>	Indomalayan|Oceanian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Saccolaimus		saccolaimus	Temminck	1838	1	Tijdschr. Nat. Gesch. Physiol.	5:14	Naked-rumped Pouched Bat	<b> affinis </b>Dobson, 1875; flavimaculatus Sody, 1931; <b> crassus </b>Blyth, 1844; pulcher Blyth, 1844; <b> nudicluniatus </b>De Vis, 1905; granti Thomas, 1911; <b> pluto </b>Miller, 1910; capito Hollister, 1913.	Indonesia, Java.	Bangladesh, India, and Sri Lanka through SE Asia (including Burma, Cambodia, Thailand, and the Nicobar Isls) to the Philippines, Sulawesi, and Borneo, Sumatra, Java, Bali, and Timor (Indonesia); New Guinea; New Britain and Bougainville Isls (Papua New Guinea); NE Queensland (Australia); Guadalcanal Isl (Solomon Isls).	Not listed.	Least Concern	Corbet and Hill (1980) listed nudicluniatus as a distinct species without comment. Includes pulcher ; see Medway (1977) and Goodwin (1979). Includes pluto ; see Corbet and Hill (1992). Reviewed in part by Chimimba and Kitchener (1991) and Bates and Harrison (1997); also see Flannery (1995 b ) and Bonaccorso (1998). Synonymies remain unclear for several names; see Jackson and Groves (2015) for a summary.	Saccolaimus saccolaimus	1004816	23	Bare-rumped Sheath-tailed Bat	Naked-rumped Free-tail Bat|Naked-rumped Pouched Bat|Naked-rumped Sheath-tailed Bat|Naked-tailed Saccolaimus|Pouch-bearing Tomb Bat|Pouched Tomb Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	TAPHOZOINAE	NA	Saccolaimus	NA	saccolaimus	Temminck	1838	1						Java, Indonesia.			saccolaimus (Temminck, 1838)|crassus (Blyth, 1844)|pulcher (Blyth, 1844)|affinis (Dobson, 1875)|nudicluniatus (De Vis, 1905)|pluto (G. S. Miller, 1910)|granti (O. Thomas, 1911)|capito (Hollister, 1913)|flavimaculatus Sody, 1931	NA	NA				India|Sri Lanka|Bangladesh|Myanmar|Thailand|Cambodia|Vietnam|Malaysia|Singapore|Nicobar Islands|Indonesia|Brunei|East Timor|Philippines|Papua New Guinea|Solomon Islands|Australia	Asia|Oceania	Indomalaya	LC	0	0	0	Saccolaimus_saccolaimus	0	sciname match	Saccolaimus_saccolaimus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Saccolaimus_saccolaimus	1004816	23	Bare-rumped Sheath-tailed Bat	Naked-rumped Free-tail Bat|Naked-rumped Pouched Bat|Naked-rumped Sheath-tailed Bat|Naked-tailed Saccolaimus|Pouch-bearing Tomb Bat|Pouched Tomb Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Taphozoinae	NA	Saccolaimus	NA	saccolaimus	Temminck	1	Taphozous Saccolaimus	Temminck, C.J. 1838. Over de geslachten _Taphozous_, _Emballonura_, _Urocryptus_ en _Diclidurus_. Tijdschrift voor Natuurlijke Geschiedenis en Physiologie 5(1-2):1-34.	https://www.biodiversitylibrary.org/page/13474788	RMNH.MAM.33630	lectotype	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.33630	Java, Indonesia.			NA	NA				India|Sri Lanka|Bangladesh|Myanmar|Thailand|Cambodia|Vietnam|Malaysia|Singapore|Andaman and Nicobar Islands|Indonesia|Brunei|East Timor|Philippines|Papua New Guinea|Solomon Islands|Australia	Asia|Oceania (Continent)	Australasia|Indomalaya	LC	0	0	0	Saccolaimus_saccolaimus	0	sciname match	Saccolaimus_saccolaimus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Saccolaimus		saccolaimus	Temminck	1838	1	Tijdschr. Nat. Gesch. Physiol.	5:14	Naked-rumped Pouched Bat	affinis Dobson, 1875; flavimaculatus Sody, 1931; crassus Blyth, 1844; pulcher Blyth, 1844; nudicluniatus De Vis, 1905; granti Thomas, 1911; pluto Miller, 1910; capito Hollister, 1913.	Indonesia, Java.	Bangladesh, India, and Sri Lanka through SE Asia (including Burma, Cambodia, Thailand, and the Nicobar Isls) to the Philippines, Sulawesi, and Borneo, Sumatra, Java, Bali, and Timor (Indonesia); New Guinea; New Britain and Bougainville Isls (Papua New Guinea); NE Queensland (Australia); Guadalcanal Isl (Solomon Isls).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19802/209550074/' target='_blank'>Least Concern</a>	Corbet and Hill (1980) listed nudicluniatus as a distinct species without comment. Includes pulcher; see Medway (1977) and Goodwin (1979). Includes pluto; see Corbet and Hill (1992). Reviewed in part by Chimimba and Kitchener (1991) and Bates and Harrison (1997); also see Flannery (1995b) and Bonaccorso (1998). Synonymies remain unclear for several names; see Jackson and Groves (2015) for a summary.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Saccolaimus saccolaimus; Saccolaimus saccolaimus; Saccolaimus saccolaimus; Saccolaimus saccolaimus; Saccolaimus saccolaimus; Saccolaimus saccolaimus; saccolaimus; affinis; crassus; nudicluniatus; pluto; affinis - flavimaculatus; crassus - pulcher; nudicluniatus - granti; pluto - capito; sacco; affinis; crassus; nudicluniatus; pluto; affinis; crassus; nudicluniatus; affinis - flavimaculatus; crassus - pulcher; nudicluniatus - granti; pluto; capito; saccolaimus; crassus; pulcher; affinis; nudicluniatus; pluto; granti; capito; flavimaculatus; 16; Bare-rumped Sheath-tailed Bat; Taphien de Temminck; Nacktbauch-Glattnasenfreischwanz; Tafozo de Temminck; Naked-rumped ree-tail Bat; Naked-rumped Pouched Bat; Naked-rumped Sheath-tailed Bat; Bare-rumped Sheath-tailed Bat; Naked-rumped Free-tail Bat; Naked-rumped Pouched Bat; Naked-rumped Sheath-tailed Bat; Naked-tailed Saccolaimus; Pouch-bearing Tomb Bat; Pouched Tomb Bat; Naked-rumped Pouched Bat; Naked-rumped Pouched Bat; S. saccolaimus