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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1503	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Taphozous mixtus	Saccolaimus mixtus	Taphozous mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus	Saccolaimus mixtus		[MSW3] Reviewed by Chimimba and Kitchener (1991). Also see Flannery (1995a) and Bonaccorso (1998).; [HMW] Saccolaimus mixtus Troughton, 1925 , Port Moresby , Papua New Guinea . This species is monotypic.; [batnames2022] Reviewed by Chimimba and Kitchener (1991). Also see Flannery (1995 a ) and Bonaccorso (1998).; [batnames2023] Reviewed by Chimimba and Kitchener (1991). Also see Flannery (1995 a ) and Bonaccorso (1998).; [batnames2025_1.7] Reviewed by Chimimba and Kitchener (1991). Also see Flannery (1995a) and Bonaccorso (1998).														mixtus				mixtus	mixtus			mixtus Troughton, 1925		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Troughton's pouched bat	S, E New Guinea, N Queensland	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Saccolaimus mixtus	Papua New Guinea, Central Prov., Port Moresby.	Troughton	1925	Rec. Aust. Mus., 14:322.	Distribution: Confined to southern and eastern New Guinea and the northern end of Cape York peninsula (Australia).		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Troughton's pouched bat	S, E New Guinea, N Queensland	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Troughton	1925	Rec. Aust. Mus., 14:322.		SE New Guinea, NE Queensland (Australia).	Papua New Guinea, Central Prov., Port Moresby.		TROUGHTON	1925	Posterior floor of mesopterygoid fossa deeply grooved. Basis phenoid pits separated by a high septum. Size rela tively small (forearm length, 61 -6 8 mm). Radio metacarpal pouch small but distinct. Sagittal crest relatively low and not forming an occipital "hel met".	Distribution: Confined to southern and eastern New Guinea and the northern end of Cape York peninsula (Australia).	No subspecies.		43	species	S. mixtus	TROUGHTON	1925	Saccolaimus	genus	Saccolaimus mixtus				Posterior floor of mesopterygoid fossa deeply grooved. Basis phenoid pits separated by a high septum. Size rela tively small (forearm length, 61 -6 8 mm). Radio metacarpal pouch small but distinct. Sagittal crest relatively low and not forming an occipital "hel met".	No subspecies.		1. S. mixtus TROUGHTON 1925.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Taphozoinae		Saccolaimus mixtus	Saccolaimus		mixtus	Troughton		1925		Rec. Aust. Mus.	14		322		Troughton's Pouched Bat	Papua New Guinea, Central Prov., Port Moresby.	SE New Guinea, NE Queensland (Australia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Reviewed by Chimimba and Kitchener (1991). Also see Flannery (1995a) and Bonaccorso (1998).	03D587F2FFC84C03FF793F73F38EF9AC	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	357	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFC74C0CFF1632ECFD40EE5E.xml	Saccolaimus mixtus	Emballonuridae	Saccolaimus	mixtus	Troughton	1925	17 @en | Papuan Sheath-tailed Bat @en | Taphien papou @fr | Papua-Glattnasenfreischwanz @de | Tafozode Papua @es | Cape York Sheath-tailed Bat @en	Saccolaimus mixtus Troughton, 1925 , Port Moresby , Papua New Guinea . This species is monotypic.	S Papua New Guinea (S coastal region in Western Province and National Capital District) and N Australia (N Cape York Peninsula); possibly on Torres Strait Is.	Head-body 72-79 mm, tail 22-29 mm, ear 17-20 mm, hindfoot 13—15 mm, forearm 61-68 mm; weight 21-27 g. Dorsum of the Papuan Sheath-tailed Bat is dark gray-brown, with some whitetipped hairs; venter is white or pale buff-brown. Gular pouch is well developed in males and poorly developed in females. Radio-metacarpal sacs are present and lined with white fur in both sexes. Rump is well haired. Flight membranes, face, ears, tail, forearm, and feet are black. Eyes large. Ears are broadly rounded and barely rise above crown of head. Strong, long calcars support trailing edges of uropatagium in flight.	Open eucalypt woodlands, sclerophyll forests, and along riparian systems. The Papuan Sheath-tailed Bat is common in larger tracts of Eucalyptus tetrodonta ( Myrt aceae) on Cape York .	The Papuan Sheath-tailed Bat forages for insects over and underneath tree canopies.	No information.	The Papuan Sheath-tailed Bat is crepuscular. It will roost in limestone caves in Papua New Guinea but more frequently shelters in tree hollows. In Australia, it often roosts in dead snags of E. tetrodonta . It emerges at dusk to forage on aerial insects above sclerophyll forest canopies or high over rivers and canyons. Low-frequency, long-duration echolocation calls of the Papuan Sheath-tailed Bat are very similar to those of the Bare-rumped Sheath-tailed Bat ( S . saccolaimus ), but they lack sharp downward frequency sweep near ends of calls of the latter. Multiharmonic call pulses of the Papuan Sheath-tailed Bat have characteristic frequency of 21 kHz.	The Papuan Sheath-tailed Bat roosts in groups of 2—7 individuals.	Classified as Near Threatened on 77 ze IUCNRed List. Numbers ofreproductively mature Papuan Sheath-tailed Bats have declined in some regional populations. Threats include large-scale removal of tall forests for bauxite mining on Cape York , habitat modification in southern New Guinea, and frequent hot burning regimes in parts of Cape York affecting succession and longevity of large, old trees required for roost hollows. The Papuan Sheath-tailed Bat occurs in protected areas on Cape York Peninsula and Tonda Wildlife Management Area in Papua New Guinea. Research on basic biology, in particular habitat and roosting preferences, is needed. Acoustic surveys should be conducted to better define distribution especially in Papua New Guinea.	Armstrong & Konishi (2013) | Armstrong, Broken-Brow et at. (2017) | Bonaccorso (1998) | Flannery (1995a) | Hall & Richards (1979) | Reardon et al. (2010) | Strahan (1995) | Tate (1941) | Thomson et al. (2008) | Troughton (1925) | Waithman (1979)	https://zenodo.org/record/3747936/files/figure.png	17 . Papuan Sheath-tailed Bat Saccolaimus mixtus French: Taphien papou / German: Papua-Glattnasenfreischwanz / Spanish: Tafozo de Papua Other common names: Cape York Sheath-tailed Bat Taxonomy . Saccolaimus mixtus Troughton, 1925 , Port Moresby , Papua New Guinea . This species is monotypic. Distribution. S Papua New Guinea (S coastal region in Western Province and National Capital District) and N Australia (N Cape York Peninsula); possibly on Torres Strait Is. Descriptive notes. Head-body 72-79 mm, tail 22-29 mm, ear 17-20 mm, hindfoot 13—15 mm, forearm 61-68 mm; weight 21-27 g. Dorsum of the Papuan Sheath-tailed Bat is dark gray-brown, with some whitetipped hairs; venter is white or pale buff-brown. Gular pouch is well developed in males and poorly developed in females. Radio-metacarpal sacs are present and lined with white fur in both sexes. Rump is well haired. Flight membranes, face, ears, tail, forearm, and feet are black. Eyes large. Ears are broadly rounded and barely rise above crown of head. Strong, long calcars support trailing edges of uropatagium in flight. Habitat . Open eucalypt woodlands, sclerophyll forests, and along riparian systems. The Papuan Sheath-tailed Bat is common in larger tracts of Eucalyptus tetrodonta ( Myrt aceae) on Cape York . Food and Feeding . The Papuan Sheath-tailed Bat forages for insects over and underneath tree canopies. Breeding . No information. Activity patterns. The Papuan Sheath-tailed Bat is crepuscular. It will roost in limestone caves in Papua New Guinea but more frequently shelters in tree hollows. In Australia, it often roosts in dead snags of E. tetrodonta . It emerges at dusk to forage on aerial insects above sclerophyll forest canopies or high over rivers and canyons. Low-frequency, long-duration echolocation calls of the Papuan Sheath-tailed Bat are very similar to those of the Bare-rumped Sheath-tailed Bat ( S . saccolaimus ), but they lack sharp downward frequency sweep near ends of calls of the latter. Multiharmonic call pulses of the Papuan Sheath-tailed Bat have characteristic frequency of 21 kHz. Movements, Home range and Social organization. The Papuan Sheath-tailed Bat roosts in groups of 2—7 individuals. Status and Conservation . Classified as Near Threatened on 77 ze IUCNRed List. Numbers ofreproductively mature Papuan Sheath-tailed Bats have declined in some regional populations. Threats include large-scale removal of tall forests for bauxite mining on Cape York , habitat modification in southern New Guinea, and frequent hot burning regimes in parts of Cape York affecting succession and longevity of large, old trees required for roost hollows. The Papuan Sheath-tailed Bat occurs in protected areas on Cape York Peninsula and Tonda Wildlife Management Area in Papua New Guinea. Research on basic biology, in particular habitat and roosting preferences, is needed. Acoustic surveys should be conducted to better define distribution especially in Papua New Guinea. Bibliography. Armstrong & Konishi (2013), Armstrong, Broken-Brow et at. (2017), Bonaccorso (1998), Flannery (1995a), Hall & Richards (1979), Reardon eta /. (2010), Strahan (1995),Tate (1941),Thomson eta /. (2008), Troughton (1925), Waithman (1979).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Saccolaimus mixtus	Saccolaimus		mixtus	Troughton	1925	0	Rec. Aust. Mus.	0.8069	Troughton's Pouched Bat	None.	Papua New Guinea, Central Prov., Port Moresby.	SE New Guinea, NE Queensland (Australia).	Not listed.	Near Threatened	Reviewed by Chimimba and Kitchener (1991). Also see Flannery (1995 a ) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Saccolaimus mixtus	23	Papuan Sheath-tailed Bat	Cape York Sheath-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	TAPHOZOINAE	NA	Saccolaimus	NA	mixtus	Troughton	1925	0	Saccolaimus_mixtus	Troughton, E. L. G. (1925). A revision of the genus Taphozous and Saccolaimus (Chiroptera) in Australia and New Guinea, including a new species and a note on two Malayan forms. Records of the Australian Museum, 14(4), 322.	https://journals.australian.museum/troughton-1925-rec-aust-mus-144-313341/	AM A.3257		Port Moresby, Papua New Guinea.			mixtus Troughton, 1925	NA	NA	Papua New Guinea|Australia	Oceania	Australasia/Oceania	NT	0	0	0	Saccolaimus_mixtus	0	sciname match	Saccolaimus_mixtus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19800	Saccolaimus mixtus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Saccolaimus	mixtus	Troughton, 1925		200000000	Saccolaimus mixtus	Near Threatened	C1	2021	2016-07-31 00:00:00 UTC	3.1	English	<p><span lang="EN-AU">The most recent assessment of Saccolaimus mixtus in its Australian range is Near Threatened based on total population size and evidence for past and anticipated future decline (approaches C1; Reardon et al . 2010, Woinarski et al . 2014). While there is recent and ongoing habitat loss from mainly bauxite mining, especially in core tall forest habitat on the west side of Cape York Peninsula, Queensland, the rate of population loss is expected to be less than 10% in three generations (around 20 years), and although population size has not been estimated, it is anticipated to be over 10,000 individuals. In the context of their global range, there is continuing habitat modification in areas around Port Moresby in Papua New Guinea, and an absence of records from that country for several decades, but the lack of information on occurrence in the lowland areas of southern Papua New Guinea limits the assessment. This species should be reassessed following habitat area modelling on Cape York Peninsula in the context of mining leases, and targeted surveys in southern New Guinea.</p>	<p><span lang="EN-AU">Saccolaimus mixtus <span lang="EN-AU"> forages over open eucalypt woodland and forest. It appears to be relatively common in the larger tracts of Eucalyptus tetrodonta forest on the western side of Cape York, but could feasibly roost in any area with larger trees. It is also associated with large river systems. While it has been collected once from a limestone cave in Papua New Guinea (Tate 1941), it usually roosts in tree hollows. It has been recorded roosting in dead E. tetrodonta stags in small groups of two to seven individuals (J. Broken-Brow unpublished data). It forages over canopy, but will also fly underneath and amongst the canopy. Its echolocation call is very similar to that of S. saccolaimus , which limi<a name="_GoBack"></a>ts the ability to distinguish these two species acoustically, but pulses are often a different shape and of higher frequency compared to S. flaviventris . ;</span></p>	<p><span lang="EN-AU">It is threatened by broad-scale removal of tall forests for bauxite mining on Cape York, and possibly ongoing habitat modification in areas of southern New Guinea. Inappropriate burning regimes (frequent, hot fires), which are becoming commonplace in some areas of Cape York, might affect the succession and longevity of roosting trees.</p>	<p><span lang="EN-AU">The tall eucalypt forests between Aurukun and Weipa on the western side of Cape York Peninsula appear to be core habitat, with recent surveys recording the species as common based on the capture of more individuals than current worldwide museum holdings (K.N. Armstrong 2012 unpub. data). The survey of Reardon et al . (2010) also presented numerous new records of occurrence from the northern half of Cape York, suggesting that it was not rare and that it utilises a range of habitat types, though subsequent surveys at the same sites have recorded it rarely, suggesting that local presence is not static (J. Broken-Brow unpublished data). On the basis of surveys in the past decade, it would be reasonable to suggest that total population size is greater than 10,000 individuals, but research is needed to clarify this.</p>	Decreasing	<p><span lang="EN-AU">Saccolaimus mixtus <span lang="EN-AU"> was first described from three specimens collected near Port Moresby, Papua New Guinea in 1878 (Troughton 1925; curated by the Australian Museum, Sydney). Only two other specimens from near-coastal lowland areas (0-100 m in elevation) have been collected from Papua New Guinea—in the National Capital District (one individual; Queensland Museum, Brisbane) and Western Province (one individual; American Museum of Natural History, New York)—with possibly two others (specimens at the University of Papua New Guinea and Natural History Museum, London) (records on GBIF <a href="http://www.gbif.org/species">http://www.gbif.org/species</a> and OZCAM <a href="http://ozcam.org.au/">http://ozcam.org.au/</a>; Bonaccorso 1998). It has not been seen in Papua New Guinea since at least 1973 (record from Natural History Museum, London). Most of the remaining 26 or so specimens have been collected from the northern part of Cape York Peninsula, Queensland, Australia. A survey by Reardon et al. (2010) found evidence of its presence via capture, spotlight observations and putatively attributed echolocation calls at several localities in the northern half of Cape York Peninsula and speculated that it could be present further south as well. Its presence on islands in the Torres Strait (Thomson et al . 2008) needs confirmation. Other capture records are available from the western side of Cape York Peninsula (K.N. Armstrong unpub. data).</span></p>		Terrestrial	<p><span lang="EN-AU">Saccolaimus mixtus <span lang="EN-AU"> occurs in several protected areas on the Cape York Peninsula, as well as the Tonda Wildlife Management Area in Papua New Guinea. Further research is required on basic biology, in particular habitat and roosting preference, plus some quantification on how much habitat of various vegetation communities is covered by mining tenements, and an estimate of population size. In Papua New Guinea, it should be targeted during research and environmental assessment surveys—with high mist nets and acoustic lures likely to increase the chance of capture.</span></p>	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Saccolaimus		mixtus	Troughton	1925	0	Rec. Aust. Mus.	0.806944	Troughton's Pouched Bat	None.	Papua New Guinea, Central Prov., Port Moresby.	SE New Guinea, NE Queensland (Australia).	Not listed.	Near Threatened	Reviewed by Chimimba and Kitchener (1991). Also see Flannery (1995 a ) and Bonaccorso (1998).	Saccolaimus mixtus	1004814	23	Papuan Sheath-tailed Bat	Cape York Sheath-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	TAPHOZOINAE	NA	Saccolaimus	NA	mixtus	Troughton	1925	0	Saccolaimus_mixtus	Troughton, E. L. G. (1925). A revision of the genus Taphozous and Saccolaimus (Chiroptera) in Australia and New Guinea, including a new species and a note on two Malayan forms. Records of the Australian Museum, 14(4), 322.	https://journals.australian.museum/troughton-1925-rec-aust-mus-144-313341/	AM A.3257		Port Moresby, Papua New Guinea.			mixtus Troughton, 1925	NA	NA				Papua New Guinea|Australia	Oceania	Australasia/Oceania	NT	0	0	0	Saccolaimus_mixtus	0	sciname match	Saccolaimus_mixtus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Saccolaimus_mixtus	1004814	23	Papuan Sheath-tailed Bat	Cape York Sheath-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Taphozoinae	NA	Saccolaimus	NA	mixtus	Troughton	0	Saccolaimus mixtus	Troughton, E.L.G. 1925-04-09. A revision of the genera _Taphozous_ and _Saccolaimus_ (Chiroptera) in Australia and New Guinea, including a new species and a note on two Malayan forms. Records of the Australian Museum 14(4):313-341.	https://doi.org/10.3853/j.0067-1975.14.1925.850	AM A.3257	holotype		Port Moresby, Papua New Guinea.			NA	NA				Papua New Guinea|Australia	Oceania (Continent)	Australasia	NT	0	0	0	Saccolaimus_mixtus	0	sciname match	Saccolaimus_mixtus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Saccolaimus		mixtus	Troughton	1925	0	Rec. Aust. Mus.	0.806944	Troughton's Pouched Bat	None.	Papua New Guinea, Central Prov., Port Moresby.	SE New Guinea, NE Queensland (Australia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19800/209535232/' target='_blank'>Near Threatened</a>	Reviewed by Chimimba and Kitchener (1991). Also see Flannery (1995a) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Saccolaimus mixtus; Saccolaimus mixtus; Saccolaimus mixtus; Saccolaimus mixtus; Saccolaimus mixtus; Saccolaimus mixtus; mixtus; 17; Papuan Sheath-tailed Bat; Taphien papou; Papua-Glattnasenfreischwanz; Tafozode Papua; Cape York Sheath-tailed Bat; Papuan Sheath-tailed Bat; Cape York Sheath-tailed Bat; Troughton's Pouched Bat; Troughton's Pouched Bat; S. mixtus