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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L16	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus crassulus	Pipistrellus crassulus	Pipistrellus crassulus	N/A	Pipistrellus crassulus	Hypsugo crassulus	Hypsugo crassulus	Nycticeinops crassulus	Nycticeinops crassulus	Pipistrellus crassulus	Nycticeinops crassulus	Nycticeinops crassulus	Nycticeinops crassulus	Afropipistrellus crassulus	Afropipistrellus crassulus		[MSW3] Includes bellieri; see Heller et al. (1994).; [HMW] Pipistrellus crassulus Thomas, 1904 , “Efulen, Cameroons [= Cameroon ].” Relationships are currently unresolved. Genetic data place race bellieri close to Hypsugo aisentrauti (but see H. eisentrauti ). Nominate crassulus has not yet been included in a phylogenetic analysis; bellieri might represent a distinct species. Two subspecies recognized.; [batnames2022] Does not includes bellieri ; see Hutterer et al (2019), but see Heller et al. (1994).; [MDD2022] previously included N. bellieri; moved from Hypsugo to the recently described Parahypsugo, and then to Nycticeinops; [IUCN] The West African population distributed in Guinea, Liberia and Côte d'Ivoire forms the subspecies P. c. bellieri , which may be valid at the species level.; [batnames2023] Does not includes bellieri ; see Hutterer et al (2019), but see Heller et al. (1994).; [MDD2023] previously included N. bellieri; moved from Hypsugo to the recently described Parahypsugo, and then to Nycticeinops; [MDD2025_2.0] previously included N. bellieri; moved from Hypsugo to the recently described Parahypsugo, and then to Nycticeinops; [batnames2025_1.7] Does not includes bellieri; see Hutterer et al (2019), but see Heller et al. (1994).; [MDD2025_2.2] previously included N. bellieri; moved from Hypsugo to the recently described Parahypsugo, and then to Nycticeinops before now being placed in Afropipistrellus									bellieri	crassulus, bellieri				crassulus	The West African population distributed in Guinea, Liberia and Côte d'Ivoire forms the subspecies P. c. bellieri , which may be valid at the species level.			crassulus	crassulus			crassulus (O. Thomas, 1904)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.			Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus crassulus	Cameroun, Efulen.	Thomas	1904	Ann. Mag. Nat. Hist., ser. 7, 13:206.	Distribution: Ranging in the forested regions of tropical Africa from southern Cameroon and southwestern Sudan to northern Angola.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-6											THOMAS	1904	Size relatively small (forearm length, 28-29 mm). Rostrum fairly broad. Braincase considerably inflated but flat on top.	Distribution: Ranging in the forested regions of tropical Africa from southern Cameroon and southwestern Sudan to northern Angola.	No subspecies.		117	species	P. crassulus	THOMAS	1905	Vansonia	subgenus	Pipistrellus crassulus				Size relatively small (forearm length, 28-29 mm). Rostrum fairly broad. Braincase considerably inflated but flat on top.	No subspecies.		45. P. crassulus THOMAS 1904.	46	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo crassulus	Hypsugo		crassulus	Thomas	y	1904		Ann. Mag. Nat. Hist., ser. 7	13		206		Broad-headed Pipistrelle	Cameroon, Efulen.	Liberia, Côte d’Ivoire, Cameroon, Dem. Rep. Congo, N Angola, S Sudan.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc) as Pipistrellus crassulus.	bellieri De Vree, 1972.	Includes bellieri; see Heller et al. (1994).	4C3D87E8FFC26A7EFA8E9D881FF8B3D6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	815	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFC26A7EFA8E9D881FF8B3D6.xml	Hypsugo crassulus	Vespertilionidae	Hypsugo	crassulus		1904	Vespere a grosse téte @fr | Breitkopfige Zwergfledermaus @de | Hypsugo cabezudo @es	Pipistrellus crassulus Thomas, 1904 , “Efulen, Cameroons [= Cameroon ].” Relationships are currently unresolved. Genetic data place race bellieri close to Hypsugo aisentrauti (but see H. eisentrauti ). Nominate crassulus has not yet been included in a phylogenetic analysis; bellieri might represent a distinct species. Two subspecies recognized.	H.c. crassulus Thomas,1904—scatteredrecordsinSCameroon,SGabon,SRepublicoftheCongo,NWAngola,NW&NEDRCongo,SWSouthSudan,W&SUganda,andWKenya. H. c. bellieri De Vree, 1972 — SE Guinea , Liberia , S Ivory Coast , and S Ghana .	Head-body c¢.43-52 mm, tail 25-32 mm, ear 9-13 mm, hindfoot 5-3-7-5 mm, forearm 28-33 mm; weight 4-8 g. Pelage of the Broad-headed Pipistrelle is soft and dense; dorsally dark brown or reddish brown in crassulus , medium to dark chocolate-brown in bellier: (hairs are unicolored); ventrally either the same as dorsum (hairs unicolored; crassulus ) or various shades of pale brown such as pale fawn and pale grayish fawn (hairs bicolored with medium to dark brown bases and pale tips; bellier:). Throat ofbellier: is usually paler, contrasting with ventral pelage but is the same as venter in crassulus . Ears are blackish ( crassulus ) or dark brown (bellieri) and broadly rounded; tragus is slightly less than one-half the ear height, being broadest at midpoint in bellieri and broadest at base of anterior margin in crassulus , and is smoothly convex on posterior margin with small basal lobe and rounded tip. Tail of crassulus is longer than in bellieri. Membranes are blackish brown ( crassulus ) or dark brown (bellieri). Penis is very long compared to other pipistrelle-like bats. Baculum ( crassulus ) is very long and slender with rounded tip and very small basal lobes in line with extended shaft in vertical plane. Skull is moderately robust; braincase is broad and rounded, low in crassulus and high in belliers; interorbital region is relatively broad; rostrum is moderately long and relatively broad; forehead is almost straight to weakly concave; sagittal and lambdoidal crests are weakly developed and form weak occipital helmet in bellieri but not crassulus . I is deeply bicuspid and very broad and robust; I’ is unicuspid; P* is minute and displaced lingually from tooth row, and barely visible to invisible above gum (larger and more visible in bellieri); P* and C' are usually in contact in crassulus but separated in bellieri; lower incisors are tricuspid; and lower molars are myotodont. Chromosomal complement has 2n = 30 and FNa = 56 ( crassulus ).	Lowland rainforest, swamp forest, coastal forest, and montane forest at elevations of 70-1600 m. Nominate crassulus seems to prefer higher elevations of 800-1600 m, but can be found down to 300 m , whereas race bellier : appears to prefer rainforest at 70-600 m.	The Broad-headed Pipistrelle forages by slow hawking in cluttered habitats.	In Ivory Coast , six pregnant females and one lactating female were found between late February and late March, four nulliparous females were captured between late August and late September, and a pregnant female was found in October. Scrotal males were found from late February to late March and in late August. Based on these data, the species may have more than one breeding season. Females produce single young.	Broad-headed Pipistrelles are nocturnal.	No information.	Classified as Least Concern on The IUCN Red Lust (as Pipistrellus crassulus ). Although rarely captured, the species appears to be widespread and locally common; e.g. in Tai National Park, Ivory Coast , where it was the most commonly captured pipistrelle-like bat.	ACR (2018) | Bates et al. (2013) | Decher et al. (2016) | Fahr (2013z) | Hayman (1954) | Hayman et al. (1966) | Heller et al. (1994) | Monadjem & Fahr (2007, 2017b) | Monadjem, Richards et al. (2013) | Volleth et al. (2001) | Weber & Fahr (2007a)	https://zenodo.org/record/6398012/files/figure.png	114. Broad-headed Pipistrelle Hypsugo crassulus French: Vespere a grosse téte / German: Breitkopfige Zwergfledermaus / Spanish: Hypsugo cabezudo Taxonomy. Pipistrellus crassulus Thomas, 1904 , “Efulen, Cameroons [= Cameroon ].” Relationships are currently unresolved. Genetic data place race bellieri close to Hypsugo aisentrauti (but see H. eisentrauti ). Nominate crassulus has not yet been included in a phylogenetic analysis; bellieri might represent a distinct species. Two subspecies recognized. Subspecies and Distribution. H.c. crassulus Thomas,1904—scatteredrecordsinSCameroon,SGabon,SRepublicoftheCongo,NWAngola,NW&NEDRCongo,SWSouthSudan,W&SUganda,andWKenya. H. c. bellieri De Vree, 1972 — SE Guinea , Liberia , S Ivory Coast , and S Ghana . Descriptive notes. Head-body c¢.43-52 mm, tail 25-32 mm, ear 9-13 mm, hindfoot 5-3-7-5 mm, forearm 28-33 mm; weight 4-8 g. Pelage of the Broad-headed Pipistrelle is soft and dense; dorsally dark brown or reddish brown in crassulus , medium to dark chocolate-brown in bellier: (hairs are unicolored); ventrally either the same as dorsum (hairs unicolored; crassulus ) or various shades of pale brown such as pale fawn and pale grayish fawn (hairs bicolored with medium to dark brown bases and pale tips; bellier:). Throat ofbellier: is usually paler, contrasting with ventral pelage but is the same as venter in crassulus . Ears are blackish ( crassulus ) or dark brown (bellieri) and broadly rounded; tragus is slightly less than one-half the ear height, being broadest at midpoint in bellieri and broadest at base of anterior margin in crassulus , and is smoothly convex on posterior margin with small basal lobe and rounded tip. Tail of crassulus is longer than in bellieri. Membranes are blackish brown ( crassulus ) or dark brown (bellieri). Penis is very long compared to other pipistrelle-like bats. Baculum ( crassulus ) is very long and slender with rounded tip and very small basal lobes in line with extended shaft in vertical plane. Skull is moderately robust; braincase is broad and rounded, low in crassulus and high in belliers; interorbital region is relatively broad; rostrum is moderately long and relatively broad; forehead is almost straight to weakly concave; sagittal and lambdoidal crests are weakly developed and form weak occipital helmet in bellieri but not crassulus . I is deeply bicuspid and very broad and robust; I’ is unicuspid; P* is minute and displaced lingually from tooth row, and barely visible to invisible above gum (larger and more visible in bellieri); P* and C' are usually in contact in crassulus but separated in bellieri; lower incisors are tricuspid; and lower molars are myotodont. Chromosomal complement has 2n = 30 and FNa = 56 ( crassulus ). Habitat. Lowland rainforest, swamp forest, coastal forest, and montane forest at elevations of 70-1600 m. Nominate crassulus seems to prefer higher elevations of 800-1600 m, but can be found down to 300 m , whereas race bellier : appears to prefer rainforest at 70-600 m. Food and Feeding. The Broad-headed Pipistrelle forages by slow hawking in cluttered habitats. Breeding. In Ivory Coast , six pregnant females and one lactating female were found between late February and late March, four nulliparous females were captured between late August and late September, and a pregnant female was found in October. Scrotal males were found from late February to late March and in late August. Based on these data, the species may have more than one breeding season. Females produce single young. Activity patterns. Broad-headed Pipistrelles are nocturnal. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red Lust (as Pipistrellus crassulus ). Although rarely captured, the species appears to be widespread and locally common; e.g. in Tai National Park, Ivory Coast , where it was the most commonly captured pipistrelle-like bat. Bibliography. ACR (2018), Bates et al. (2013), Decher et al. (2016), Fahr (2013z), Hayman (1954), Hayman et al. (1966), Heller et al. (1994), Monadjem & Fahr (2007, 2017b), Monadjem, Richards et al. (2013), Volleth et al. (2001), Weber & Fahr (2007a).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Nycticeinops crassulus	Nycticeinops		crassulus	Thomas	1904	1	Ann. Mag. Nat. Hist.	ser. 7, 13: 206	Broad-headed Pipistrelle	None.	Cameroon, Efulen	Cameroon, Congo, DR Congo, Sudan, Uganda, Kenya, Angola	Not listed.	Least Concern as Pipistrellus crassulus 	Does not includes bellieri ; see Hutterer et al (2019), but see Heller et al. (1994).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nycticeinops crassulus	23	Broad-headed Serotine	Broad-headed Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Nycticeinops	NA	crassulus	O. Thomas	1904	1	Pipistrellus_crassulus	Thomas, O. (1904). New bats from British East Africa collected by Mrs. Hinde, and from the Cameroons by Mr. G. L. Bates. Annals and Magazine of Natural History, Ser. 7, 13, 206.	https://www.biodiversitylibrary.org/item/61725#page/230/mode/1up	BM 1904.2.8.1		"Efulen, Cameroons [= Cameroon]."			crassulus (O. Thomas, 1904)	previously included N. bellieri; moved from Hypsugo to the recently described Parahypsugo, and then to Nycticeinops	Roehrs, Z. P., Lack, J. B., & Van Den Bussche, R. A. (2010). Tribal phylogenetic relationships within Vespertilioninae (Chiroptera: Vespertilionidae) based on mitochondrial and nuclear sequence data. Journal of Mammalogy, 91(5), 1073-1092.|Hutterer, R., Decher, J., Monadjem, A., & Astrin, J. (2019). A new genus and species of vesper bat from West Africa, with notes on Hypsugo, Neoromicia, and Pipistrellus (Chiroptera: Vespertilionidae). Acta Chiropterologica, 21(1), 1-22.|Monadjem, A., Richards, L. R., Decher, J., Hutterer, R., Mamba, M. L., Guyton, J., Naskrecki, P., Markotter, W., Wipfler, B., Kropff, A. S., & Dalton, D. L (2020) A phylogeny for African Pipistrellus species with the description of a new species from West Africa (Mammalia: Chiroptera). Zoological Journal of the Linneaen Society, zlaa068.|Monadjem, A., Demos, T. C., Dalton, D. L., Webala, P. W., Musila, S., Kerbis Peterhans, J. C., & Patterson, B. D. (2020). A revision of the pipistrelle-like bats (Mammali: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society, zlaa087.	Cameroon|Gabon|Republic of the Congo|Angola|Democratic Republic of the Congo|South Sudan|Uganda|Kenya	Africa	Afrotropic	LC	0	0	0	Parahypsugo_crassulus	1	oldname match	Hypsugo_crassulus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	44853	Pipistrellus crassulus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	crassulus	Thomas, 1904	The West African population distributed in Guinea, Liberia and Côte d'Ivoire forms the subspecies P. c. bellieri , which may be valid at the species level.	20000000	Pipistrellus crassulus	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	It has been recorded from both montane and lowland tropical moist forest, and from lowland tropical dry forest.	There appear to be no major threats to this widespread species as a whole.	This species has rarely been recorded.	Unknown	This widespread African bat has been recorded from Guinea, Liberia and Côte d'Ivoire in the west, from Cameroon, Gabon, Congo, Democratic Republic of the Congo, northern Angola, southern Sudan, Uganda and western Kenya (Kakamega Forest).		Terrestrial	There appear to be no direct conservation measures in place, and it is not known if the species is present within any protected areas. Further studies are needed into the taxonomy, distribution and natural history of this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Nycticeinops		crassulus	Thomas	1904	1	Ann. Mag. Nat. Hist.	ser. 7, 13: 206	Broad-headed Pipistrelle	None.	Cameroon, Efulen	Cameroon, Congo, DR Congo, Sudan, Uganda, Kenya, Angola	Not listed.	Least Concern as Pipistrellus crassulus 	Does not includes bellieri ; see Hutterer et al (2019), but see Heller et al. (1994).	Nycticeinops crassulus	1005747	23	Broad-headed Serotine	Broad-headed Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Nycticeinops	NA	crassulus	O. Thomas	1904	1	Pipistrellus_crassulus	Thomas, O. (1904). New bats from British East Africa collected by Mrs. Hinde, and from the Cameroons by Mr. G. L. Bates. Annals and Magazine of Natural History, Ser. 7, 13, 206.	https://www.biodiversitylibrary.org/item/61725#page/230/mode/1up	BM 1904.2.8.1		"Efulen, Cameroons [= Cameroon]."			crassulus (O. Thomas, 1904)	previously included N. bellieri; moved from Hypsugo to the recently described Parahypsugo, and then to Nycticeinops	Roehrs, Z. P., Lack, J. B., & Van Den Bussche, R. A. (2010). Tribal phylogenetic relationships within Vespertilioninae (Chiroptera: Vespertilionidae) based on mitochondrial and nuclear sequence data. Journal of Mammalogy, 91(5), 1073-1092.|Hutterer, R., Decher, J., Monadjem, A., & Astrin, J. (2019). A new genus and species of vesper bat from West Africa, with notes on Hypsugo, Neoromicia, and Pipistrellus (Chiroptera: Vespertilionidae). Acta Chiropterologica, 21(1), 1-22.|Monadjem, A., Richards, L. R., Decher, J., Hutterer, R., Mamba, M. L., Guyton, J., Naskrecki, P., Markotter, W., Wipfler, B., Kropff, A. S., & Dalton, D. L (2020) A phylogeny for African Pipistrellus species with the description of a new species from West Africa (Mammalia: Chiroptera). Zoological Journal of the Linneaen Society, zlaa068.|Monadjem, A., Demos, T. C., Dalton, D. L., Webala, P. W., Musila, S., Kerbis Peterhans, J. C., & Patterson, B. D. (2020). A revision of the pipistrelle-like bats (Mammali: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society, zlaa087.				Cameroon|Gabon|Republic of the Congo|Angola|Democratic Republic of the Congo|South Sudan|Uganda|Kenya	Africa	Afrotropic	LC	0	0	0	Parahypsugo_crassulus	1	oldname match	Hypsugo_crassulus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Afropipistrellus_crassulus	1005747	23	Broad-headed Serotine	Broad-headed Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Afropipistrellus	NA	crassulus	O. Thomas	1	Pipistrellus crassulus	Thomas, O. 1904-03-01. New bats from British East Africa collected by Mrs. Hinde, and from the Cameroons by Mr. G. L. Bates. Annals and Magazine of Natural History (7)13(75):206-210.	https://www.biodiversitylibrary.org/page/16108292	BMNH:Mamm:1904.2.8.1	holotype	https://data.nhm.ac.uk/object/4ad6cfeb-933e-48e3-a59b-eae3358f0517	"Efulen, Cameroons [= Cameroon]."			previously included N. bellieri; moved from Hypsugo to the recently described Parahypsugo, and then to Nycticeinops before now being placed in Afropipistrellus	Roehrs, Z. P., Lack, J. B., & Van Den Bussche, R. A. (2010). Tribal phylogenetic relationships within Vespertilioninae (Chiroptera: Vespertilionidae) based on mitochondrial and nuclear sequence data. Journal of Mammalogy, 91(5), 1073-1092.|Hutterer, R., Decher, J., Monadjem, A., & Astrin, J. (2019). A new genus and species of vesper bat from West Africa, with notes on Hypsugo, Neoromicia, and Pipistrellus (Chiroptera: Vespertilionidae). Acta Chiropterologica, 21(1), 1-22.|Monadjem, A., Richards, L. R., Decher, J., Hutterer, R., Mamba, M. L., Guyton, J., Naskrecki, P., Markotter, W., Wipfler, B., Kropff, A. S., & Dalton, D. L (2020) A phylogeny for African Pipistrellus species with the description of a new species from West Africa (Mammalia: Chiroptera). Zoological Journal of the Linneaen Society, zlaa068.|Monadjem, A., T. C. Demos, D. L. Dalton, P. W. Webala, S. Musila, J. C. K. Peterhans and B. D. Patterson (2021). A revision of pipistrelle-like bats (Mammalia: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society 191(4): 1114–1146.|Demos, T. C., Webala, P. W., Monadjem, A., & Patterson, B. D. (2024). Nuclear introns support the subtribe Laephotina and recently proposed genera of African Vespertilionidae. Acta Chiropterologica, 26(2), 143-152.				Cameroon|Gabon|Republic of the Congo|Angola|Democratic Republic of the Congo|South Sudan|Uganda|Kenya	Africa	Afrotropic	LC (as Pipistrellus crassulus)	0	0	0	Parahypsugo_crassulus	1	oldname match	Hypsugo_crassulus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Afropipistrellus		crassulus	Thomas	1904	1	Ann. Mag. Nat. Hist.	ser. 7, 13: 206	Broad-headed Pipistrelle	Parahypsugo	Cameroon, Efulen	Cameroon, Congo, DR Congo, Sudan, Uganda, Kenya,     Angola	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/44853/22072238/' target='_blank'>Least Concern as Pipistrellus crassulus</a>	Does not includes bellieri; see Hutterer et al (2019), but see Heller et al. (1994).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hypsugo crassulus; Hypsugo crassulus; Nycticeinops crassulus; Nycticeinops crassulus; Pipistrellus crassulus; Nycticeinops crassulus; bellieri; crassulus; bellieri; crassulus; Vespere a grosse téte; Breitkopfige Zwergfledermaus; Hypsugo cabezudo; Broad-headed Serotine; Broad-headed Pipistrelle; Broad-headed Pipistrelle; Broad-headed Pipistrelle; N. crassulus