http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1500	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rousettus obliviosus	Rousettus obliviousus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus	Rousettus obliviosus		[MSW2] Subgenus Rousettus.; [MSW3] Subgenus Rousettus. Considered a subspecies of madagascarensis by Peterson et al. (1995), but see Bergmans (1994).; [HMW] Rousettus obliviosus Kock, 1978 , near Bomboni, 640 m , Grand Comore , Comoro Islands . Rousettus obliviosus is not a subspecies of R. madagascariensis as once thought; genetic studies have shown a clear separation between the two. Monotypic.; [batnames2022] Subgenus Rousettus . Considered a subspecies of madagascarensis by Peterson et al. (1995), but clearly distinct see Bergmans (1994) and Goodman et al. (2010).; [IUCN] <p>Rousettus obliviosus was previously considered to be a subspecies of R. madagascariensis (e.g., Peterson et al. 1995), but the two forms have been shown to be distinct species (Kock 1978, Goodman et al. 2010a).</p>; [batnames2023] Subgenus Rousettus . Considered a subspecies of madagascarensis by Peterson et al. (1995), but clearly distinct see Bergmans (1994) and Goodman et al. (2010).; [batnames2025_1.7] Subgenus Rousettus. Considered a subspecies of madagascarensis by Peterson et al. (1995), but clearly distinct see Bergmans (1994) and Goodman et al. (2010).														obliviosus	<p>Rousettus obliviosus was previously considered to be a subspecies of R. madagascariensis (e.g., Peterson et al. 1995), but the two forms have been shown to be distinct species (Kock 1978, Goodman et al. 2010a).</p>			obliviosus	obliviosus			obliviosus Kock, 1978|obliviousus Honacki, Kinman, & Koeppl, 1982 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Comoro Is; ref. 4.123	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rousettus obliviousus	Comoro Isis., Grand Comoro, near Boboni, 640 m.	Kock	1978	Proc. 4th Int. Bat Res. Conf. Nairobi, p. 208.	Distribution: This spe cies is confined to the Comoro islands (between eastern Africa and northern Madagascar).		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Comoro Is	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Kock	1978	Proc. 4th Int. Bat Res. Conf. Nairobi, p. 208.	Subgenus Rousettus.	Comoro Isis.	Comoro Isis, Grand Comoro, near Boboni, 640m.		KOCK	1978	A small species (forearm length, 70-75 mm; pollex, 22-2 5 mm). Molars relatively broad. Median edge of pla giopatagium attached to side of body. Uropatagium relatively naked. Body fur relative ly short. Basicranial deflection greater than in other species of R. (Rousettus), but not as great as in R. (Stenonycteris).	Distribution: This spe cies is confined to the Comoro islands (between eastern Africa and northern Madagascar).	No subspecies.		20	species	R. obliviosus	KOCK	1978	Rousettus	subgenus	Rousettus obliviosus				A small species (forearm length, 70-75 mm; pollex, 22-2 5 mm). Molars relatively broad. Median edge of pla giopatagium attached to side of body. Uropatagium relatively naked. Body fur relative ly short. Basicranial deflection greater than in other species of R. (Rousettus), but not as great as in R. (Stenonycteris).	No subspecies.		7. R. obliviosus KOCK 1978.	7	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Rousettus obliviosus	Rousettus	Rousettus	obliviosus	Kock		1978		Proc. 4th Int. Bat Res. Conf. Nairobi			208		Comoro Rousette	Comoro Isls, Grand Comoro, near Boboni, 640 m.	Comoro Isls.	IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Lower Risk (nt).		Subgenus Rousettus. Considered a subspecies of madagascarensis by Peterson et al. (1995), but see Bergmans (1994).	03AD87FAFFDFF6368C6B3925FDCBF3C1	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	88	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFDAF6358CBB3313FEF3FB7E.xml	Rousettus obliviosus	Pteropodidae	Rousettus	obliviosus	Kock	1978	Roussette des Comores @fr | Komoren-Flughund @de | Rosetus de las Comoras @es | Comoros Rousette @en	Rousettus obliviosus Kock, 1978 , near Bomboni, 640 m , Grand Comore , Comoro Islands . Rousettus obliviosus is not a subspecies of R. madagascariensis as once thought; genetic studies have shown a clear separation between the two. Monotypic.	Comoros Is ( Grande Comore , Mohéli , and Anjouan ).	Head-body 121- 142 mm , tail 14-25 mm , ear 15-18 mm , hindfoot 13-17 mm , forearm 70-78 mm ; weight 42-73 g . The Comoro Rousette is smaller than the Egyptian Rousette ( R. aegyptiacus ) and similar in size to the Malagasy Rousette ( R. madagascariensis ), but it has a more robust skull and strongly deflected braincase, with alveolar line projected backward passing through occiput above foramen magnum. Fur color is dull gray-brown, and dorsal side oftibia is naked. Postorbital width is usually larger than interorbital width, supra-occipital crest is well developed, and premaxillae are not co-ossified. Palatal ridge pattern is normally 4 + 3 + 1. Cheekteeth have typical rousettine proportions, and P' is relatively large.	Widely distributed in now limited tropical moist forest and agricultural land from sea level up to elevations of ¢. 1750 m . Comoro Rousettes are very tolerant of human-dominated landscapes.	Fruits have been identified as food sources of the Comoro Rousette based on regurgitated pellets, including Ficus ( Moraceae ), Gambeya ( Sapotaceae ), Anthocleista grandiflora ( Gentianaceae ), and introduced species such as papaya ( Carica papaya, Caricaceae ), jackfruit ( Artocarpus integrifolia, Moraceae ), banana ( Musa spp. , Musaceae ), and Ceiba pentandra ( Malvaceae ). They also use flowers inferred by pollen on facial fur and visits to flowering plants. Use of leaves was inferred from small pellets. Comoro Rousettes forage in agricultural fields, degraded or intact forests, and understory plantations.	Lactating and pregnant Comoro Rousettes were captured in July, and nonreproductive young were captured in July-August on Anjouan . Pregnant and lactating females and reproductive males were captured in November on Grande Comore . Based on these observations, the Comoro Rousette probably is seasonally monoestrous, with reproduction in wet seasons (October—April).	Comoro Rousettes are strictly nocturnal, usually leaving roostsites around sunset and circling outside rock shelters before departing in large swarms. They return to caves gradually, swarming and circling entrances until sunrise. They have been seen flying above, at, or below canopy levels. They are very maneuverable and can hover briefly. They roost in caves formed by lava tubes and rock shelters (shallow caves), overhanging above watercourses and cascades. Caves are occupied beyond twilight zones, up to 150 m from entrances. Roost sites were found in densely forested, isolated areas on Mohéli and in large caves surrounded by agricultural fields on Grande Comore , although cave entrances were always covered by thick vegetation.	Comoro Rousettes cluster in small (50 individuals) to large colonies (3000-4000 individuals). A genetic study based on cytochrome-b and six microsatellite loci showed no interisland genetic structure, indicating that distances of 40-80 km between Comoros Islands are not enough to stop dispersal of Comoro Rousettes across islands. Nevertheless, they are absent from Mayotte probably because of lack of adequate roosting sites.	Classified as Vulnerable on The IUCN Red List. Conservation status of the Comoro Rousette is due to scarcity of roosts, physical instability of caves, and rapid deforestation in the Comoros Islands in recent years, despite its adaptability to disturbed habitat. The Comoro Rousette receives the highest level of legal protection available in the Union of the Comoros . It was listed as an “integrally-protected species” in 2001, which prohibits capture or detention of individuals without a permit. The national Conservation Action Plan for Livingstone’s Flying Fox ( Pteropus livingstonit), another threatened pteropodid from Comoros , includes an appendix with recommendations for conservation of the Comoro Rousette. These recommendations focus on protection of roost caves, discouraging hunting through environmental education, and population monitoring.	Bergmans (1994) | Goodman, Chan et al. (2010) | Goodman, Weyeneth et al. (2010) | Sewall (2016) | Sewall et al. (2003)		48. Comoro Rousette Rousettus obliviosus French: Roussette des Comores / German: Komoren-Flughund / Spanish: Rosetus de las Comoras Other common names: Comoros Rousette Taxonomy. Rousettus obliviosus Kock, 1978 , near Bomboni, 640 m , Grand Comore , Comoro Islands . Rousettus obliviosus is not a subspecies of R. madagascariensis as once thought; genetic studies have shown a clear separation between the two. Monotypic. Distribution. Comoros Is ( Grande Comore , Mohéli , and Anjouan ). Descriptive notes. Head-body 121- 142 mm , tail 14-25 mm , ear 15-18 mm , hindfoot 13-17 mm , forearm 70-78 mm ; weight 42-73 g . The Comoro Rousette is smaller than the Egyptian Rousette ( R. aegyptiacus ) and similar in size to the Malagasy Rousette ( R. madagascariensis ), but it has a more robust skull and strongly deflected braincase, with alveolar line projected backward passing through occiput above foramen magnum. Fur color is dull gray-brown, and dorsal side oftibia is naked. Postorbital width is usually larger than interorbital width, supra-occipital crest is well developed, and premaxillae are not co-ossified. Palatal ridge pattern is normally 4 + 3 + 1. Cheekteeth have typical rousettine proportions, and P' is relatively large. Habitat. Widely distributed in now limited tropical moist forest and agricultural land from sea level up to elevations of ¢. 1750 m . Comoro Rousettes are very tolerant of human-dominated landscapes. Food and Feeding. Fruits have been identified as food sources of the Comoro Rousette based on regurgitated pellets, including Ficus ( Moraceae ), Gambeya ( Sapotaceae ), Anthocleista grandiflora ( Gentianaceae ), and introduced species such as papaya ( Carica papaya, Caricaceae ), jackfruit ( Artocarpus integrifolia, Moraceae ), banana ( Musa spp. , Musaceae ), and Ceiba pentandra ( Malvaceae ). They also use flowers inferred by pollen on facial fur and visits to flowering plants. Use of leaves was inferred from small pellets. Comoro Rousettes forage in agricultural fields, degraded or intact forests, and understory plantations. Breeding. Lactating and pregnant Comoro Rousettes were captured in July, and nonreproductive young were captured in July-August on Anjouan . Pregnant and lactating females and reproductive males were captured in November on Grande Comore . Based on these observations, the Comoro Rousette probably is seasonally monoestrous, with reproduction in wet seasons (October—April). Activity patterns. Comoro Rousettes are strictly nocturnal, usually leaving roostsites around sunset and circling outside rock shelters before departing in large swarms. They return to caves gradually, swarming and circling entrances until sunrise. They have been seen flying above, at, or below canopy levels. They are very maneuverable and can hover briefly. They roost in caves formed by lava tubes and rock shelters (shallow caves), overhanging above watercourses and cascades. Caves are occupied beyond twilight zones, up to 150 m from entrances. Roost sites were found in densely forested, isolated areas on Mohéli and in large caves surrounded by agricultural fields on Grande Comore , although cave entrances were always covered by thick vegetation. Movements, Home range and Social organization. Comoro Rousettes cluster in small (50 individuals) to large colonies (3000-4000 individuals). A genetic study based on cytochrome-b and six microsatellite loci showed no interisland genetic structure, indicating that distances of 40-80 km between Comoros Islands are not enough to stop dispersal of Comoro Rousettes across islands. Nevertheless, they are absent from Mayotte probably because of lack of adequate roosting sites. Status and Conservation. Classified as Vulnerable on The IUCN Red List. Conservation status of the Comoro Rousette is due to scarcity of roosts, physical instability of caves, and rapid deforestation in the Comoros Islands in recent years, despite its adaptability to disturbed habitat. The Comoro Rousette receives the highest level of legal protection available in the Union of the Comoros . It was listed as an “integrally-protected species” in 2001, which prohibits capture or detention of individuals without a permit. The national Conservation Action Plan for Livingstone’s Flying Fox ( Pteropus livingstonit), another threatened pteropodid from Comoros , includes an appendix with recommendations for conservation of the Comoro Rousette. These recommendations focus on protection of roost caves, discouraging hunting through environmental education, and population monitoring. Bibliography. Bergmans (1994), Goodman, Chan et al. (2010), Goodman, Weyeneth et al. (2010), Sewall (2016), Sewall et al. (2003).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Rousettus obliviosus	Rousettus	Rousettus	obliviosus	Kock	1978	0	Proc. 4th Int. Bat Res. Conf. Nairobi	p. 208	Comoro Rousette	None.	Comoro Isls, Grand Comoro, near Boboni, 640 m.	Comoro Isls.	Not listed.	Vulnerable	Subgenus Rousettus . Considered a subspecies of madagascarensis by Peterson et al. (1995), but clearly distinct see Bergmans (1994) and Goodman et al. (2010).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rousettus obliviosus	23	Comoro Rousette	Comoros Rousette	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	ROUSETTINAE	ROUSETTINI	Rousettus	NA	obliviosus	Kock	1978	0	Rousettus_obliviosus	Kock, D. (1978) A new fruit bat of the genus Rousettus Gray, 1821, from the Comoro Islands, western Indian Ocean (Mammalia: Chiroptera). In Olembo, R.J., Castelino, J.B. & Mutere, F.A. (eds.). Proceedings of the 4th International Bat Research Conference Nairobi, 208.		ZMB 58207		near Bomboni, 640 m, Grand Comore, Comoro Islands.			obliviosus Kock, 1978	NA	NA	Comoros	Africa	Afrotropic	VU	0	0	0	Rousettus_obliviosus	0	sciname match	Rousettus_obliviosus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19757	Rousettus obliviosus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Rousettus	obliviosus	Kock, 1978	<p>Rousettus obliviosus was previously considered to be a subspecies of R. madagascariensis (e.g., Peterson et al. 1995), but the two forms have been shown to be distinct species (Kock 1978, Goodman et al. 2010a).</p>	200000000	Rousettus obliviosus	Vulnerable	B1ab(iii)+2ab(iii)	2020	2016-05-30 00:00:00 UTC	3.1	English	The species has a limited range; both its extent of occurrence and area of occupancy are less than thresholds of 20,000 kmÂ² and ; 2,000 kmÂ², respectively (the minimum convex polygon including all three islands it inhabits is 9,085 kmÂ², and the total area of all three islands it inhabits is 1,783 kmÂ²). It is also known to exist at no more than ten locations (seven consistently-used roosting caves are known), and it is experiencing a continuing decline in the extent and quality of its habitat (due to increasing human pressures leading to substantial loss of native forests; ongoing loss of trees from plantations, orchards, and agroforestry areas; and increasing human disturbance at roost sites). Thus, the species is listed as Vulnerable under B1ab(iii)+2ab(iii).	The Comoros Islands harbour a distinct forest type of tropical moist broadleaf forest, part of the â€˜Comoros forestsâ€™ ecoregion (Olson et al.  2001), which formerly served as the foraging habitat for R. obliviosus . However, nearly all large patches of this forest type, except in some higher elevations, have been destroyed. In parts of the remaining forest, the understory has been replaced by banana plantations, taro, or clove trees. R. obliviosus has been widely recorded as occurring in native forest, underplanted forest, and heavily-degraded forest within agricultural zones (Clark et al.  1997, Sewall et al.  2003, Goodman et al.  2010b). These records, and the presence in its diet of fruit from cultivated, non-native trees (Sewall et al.  2003), suggest that this species has thus far been able to exploit human-dominated landscapes â€“ including agroforestry lands and orchards, as well as agricultural landscapes retaining remnant trees and forest patches â€“ as foraging habitat. R. obliviosus has not been observed in villages or towns (Sewall et al.  2003, Goodman et al.  2010b). Rousettus obliviosus is a cave-roosting species that is only known to roost in caves and shallow rock shelters in areas infrequently visited by people (Sewall et al.  2003). The species appears to be selective in roost sites, as it has been found in &lt;20% of surveyed caves (Middleton 1998, 1999; Sewall et al.  2003; Hume and Middleton 2011; Middleton, pers. comm.; BJS, pers. obs.). Roosting habitat is either large lava tubes that are difficult to access or cave localities or shallow rock overhangs associated with streams or waterfalls in remote areas with steep terrain or dense vegetation (Sewall et al.  2003). Availability of such roosting habitat appears critical for this species. The lack of suitable roosting habitat on Mayotte may explain the absence of this species there (Goodman et al.  2010b) even though the species is capable of traversing open water over distances equal to that between Mayotte and its nearest neighbour, Anjouan (Goodman et al.  2010a).	Rousettus obliviosus has a limited range; its area of occupancy is less than the total area of the three islands (Sewall et al.  2003), which is 1,783 kmÂ² (Louette et al.  2004), and its extent of occurrence (including unsuitable ocean areas between the islands) is less than 9,085 kmÂ². It is therefore susceptible to single threatening processes, like cyclones, that could simultaneously or rapidly affect its entire range. The major threat to the species within this range appears to be disturbance at roost sites. The bats appear highly susceptible to human disturbance, and infrequent hunting and rock falls in unstable lava tubes have been recorded at roost sites (Middleton 1998, 1999; Sewall et al.  2003). Additionally, cave surveys between 2000-2005 revealed the bats were absent from three caves where they apparently formerly occurred (Sewall et al.  2003, W. Masefield, pers. comm.), though it is unclear whether this suggests intermittent use of some roost sites or abandonment after disturbance. Over the last few decades, the Comoros have undergone a sustained, rapid deforestation, resulting in the loss of nearly all its native forests. During the past 20 years alone, the country has lost 75% of its remaining forests, the fastest rate of any country in the world (UNDP 2013). Human pressures on native forests are expected to continue, as the growth rate of the human population remains high at 2.5% (UNDP 2013). Because R. obliviosus is regularly recorded in human-modified landscapes, loss of foraging habitat via deforestation is possibly not a major threat (Bergmans 1994). However, even human-modified landscapes containing non-native and remnant native trees are disappearing in the Comoros to make room for ground crops (FAO 2010, BJS pers. obs.); thus food resources for this species have probably been declining in recent years. Further, the loss of natural and underplanted forests may reduce its ability to cope with droughts and cyclones, and may limit its access to food resources when fruit from horticultural species is unavailable (Sewall et al.  2003). In addition, the replacement of native forests with agricultural land on Anjouan and MohÃ©li may render formerly inaccessible roost sites on Anjouan and MohÃ©li more easily reached, possibly exposing roosts to increased levels of human disturbance. Finally, the loss of native forests has resulted in impermanence or complete drying of nearly all rivers on Anjouan and most on MohÃ©li (Louette et al.  2004). R. obliviosus roosts on those islands are typically near waterfalls, and it is unclear whether that is simply where suitable caves and rock shelters occur or if the bats benefit from the adjacent water source. The species could benefit from an adjacent water source if, for instance, local humidity reduces evaporative water loss in the normally hot climate of the Comoros. If so, then it is possible the seasonal or year-round loss of water flow could affect this species on Anjouan and MohÃ©li as well.	Rousettus obliviosus is rarely observed outside of research studies due to its nocturnal foraging and cryptic roosting habits. It has nonetheless been captured frequently by bat researchers (Reason et al.   1994, Clark et al.   1997, Sewall et al.   2003, Goodman et al.   2010b), and thus the species is considered fairly common (Reason et al.   1994, Louette et al.   2004, Goodman et al.   2010b). The true population size is unknown. The only abundance estimates available are from roost counts, and very few roost sites are known. Sewall et al.   (2003) searched Grande Comore, Anjouan, and MohÃ©li and found or confirmed five permanent roosts plus a sixth that was intermittently used. They estimated that colony size at roosts ranges from around 100 to several thousand animals, and that the total population of these six sites was between 7,100 and 17,100 bats (Sewall et al.   2003). Since then, two additional roosts have been discovered, one small (~100 bats) and one mid-sized (~2,000 bats) (Hume and Middleton 2011, I. SaÃ¯d and W. Masefield pers. comm.). Thus, the current population at known roosts is estimated to number about 10,000 to 20,000 bats. Estimates of demographic parameters from genetic analyses suggest recent population stability in this species (Goodman et al.   2010a). Further, no genetic population structure is evident among the three islands, despite separation of at least 40 km of open water between adjacent islands; this indicates inter-island dispersal (Goodman et al.   2010a).	Unknown	Rousettus obliviosus has been recorded only from the three islands of Grande Comore, Anjouan, and MohÃ©li of the Union of the Comoros, western Indian Ocean (Sewall et al.   2003, Louette et al.   2004, Goodman et al.   2010b). The species has been recorded from 30-1,750 m on Grande Comore, from 200 to 1,350 m on Anjouan, and from 10-700 m on MohÃ©li (Meirte 1984, Reason et al.   1994, Sewall et al.   2003, Goodman et al.   2010b). Since the highest points on these islands are 2,361 m, 1,595 m, and 790 m, respectively (Battistini and VÃ©rin 1984), and since diverse canopy rainforest or cloud forest habitat is replaced by tree-fern, scrub, grassland, or heath habitat above 1,800 m on Mt. Karthala on Grande Comore (Louette et al.   2004), the bat species probably occurs from near sea level to about 1,800 m on Grande Comore, and across most of the elevational ranges of Anjouan and MohÃ©li. These records also suggest that the area of occupancy is less than the total area of the three islands (Sewall et al.   2003), which is 1,783 kmÂ² (Louette et al.   2004). Likewise, these records suggest that the extent of occurrence is less than the total area of the three islands and intervening unsuitable ocean habitat, which is 9,085 kmÂ². The species is apparently absent from Mayotte, the other large island in the Comoros Archipelago, as it has never been observed there despite extensive searching and trapping effort (Sewall et al.   2003, Louette et al.   2004, Goodman et al.   2010b).	Infrequent hunting has been recorded.	Terrestrial	Rousettus obliviosus receives the highest level of legal protection available in the Union of the Comoros. It is listed as an â€˜integrally-protected speciesâ€™ (list 1 of RFIC 2001), which prohibits the capture or detention of R. obliviosus individuals without a permit. This law also expressly prohibits the killing of members of the Family Pteropodidae (flying foxes); transport, purchase, sale, export or re-export of living or dead individuals or their body parts; all disruption during the period of reproduction and raising of young; and the destruction of roosts (RFIC 2001). The Union of the Comoros also ratified the Convention on Biological Diversity in 1994, and in response has developed a National Biodiversity Conservation Strategy (Roby and Dossar 2000). This strategy highlights the importance of, threats to, and conservation recommendations for fruit bats of the Union of the Comoros (Sewall and Granek 2000). In practice, however, there are currently no direct conservation actions in place for R. obliviosus , and no formal terrestrial protected areas in the country. The national Conservation Action Plan for Pteropus livingstonii (Sewall et al. 2007), another threatened pteropodid from the Comoros, includes an appendix for the conservation of Rousettus obliviosu s. This appendix addresses current and potential emerging threats to R. obliviosus , and recommends five targeted conservation actions for R. obliviosus . Specifically, it recommends (1) protection of roost caves, especially the caves with the largest known colony on each island (these are now thought to be Panga Chilamouinani near Fassi on Grande Comore, Bakomdrundru near Ndrondroni on MohÃ©li, and Mangua Mitsano near Limbi on Anjouan); (2) discouraging, through environmental education, the hunting of all fruit bats; (3) conducting a comprehensive field search for more roost sites; (4) providing incentives to individuals owning land near roost sites to conserve caves and bats; and (5) devising a suitable population monitoring protocol and conducting regular visits to all known roost sites to track population changes (Sewall et al. 2007). Other key actions for this species include efforts to prevent further native forest habitat loss and additional studies designed to track the speciesâ€™ population and inform its management (Sewall et al. 2003).	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Rousettus	Rousettus	obliviosus	Kock	1978	0	Proc. 4th Int. Bat Res. Conf. Nairobi	p. 208	Comoro Rousette	None.	Comoro Isls, Grand Comoro, near Boboni, 640 m.	Comoro Isls.	Not listed.	Vulnerable	Subgenus Rousettus . Considered a subspecies of madagascarensis by Peterson et al. (1995), but clearly distinct see Bergmans (1994) and Goodman et al. (2010).	Rousettus obliviosus	1004545	23	Comoro Rousette	Comoros Rousette	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	ROUSETTINAE	ROUSETTINI	Rousettus	NA	obliviosus	Kock	1978	0	Rousettus_obliviosus	Kock, D. (1978) A new fruit bat of the genus Rousettus Gray, 1821, from the Comoro Islands, western Indian Ocean (Mammalia: Chiroptera). In Olembo, R.J., Castelino, J.B. & Mutere, F.A. (eds.). Proceedings of the 4th International Bat Research Conference Nairobi, 208.		ZMB 58207		near Bomboni, 640 m, Grand Comore, Comoro Islands.			obliviosus Kock, 1978	NA	NA				Comoros	Africa	Afrotropic	VU	0	0	0	Rousettus_obliviosus	0	sciname match	Rousettus_obliviosus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rousettus_obliviosus	1004545	23	Comoro Rousette	Comoros Rousette	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Epomophorinae	Rousettini	Rousettus	NA	obliviosus	Kock	0	Rousettus (Rousettus) obliviosus	Kock, D. 1978. A new fruit bat of the genus _Rousettus_ Gray 1821, from the Comoro Islands, western Indian Ocean (Mammalia: Chiroptera). Pp. 205â€“216 in Olembo, R.J., Castelino, J.B. and Mutere, F.A. (eds.). Proceedings of the Fourth International Bat Research Conference. Kenya Literature Bureau, Nairobi.		ZMB 58207	holotype		near Bomboni, 640 m, Grand Comore, Comoro Islands.	-11.6	43.31667	NA	NA				Comoros	Africa	Afrotropic	VU	0	0	0	Rousettus_obliviosus	0	sciname match	Rousettus_obliviosus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Rousettus	Rousettus	obliviosus	Kock	1978	0	Proc. 4th Int. Bat Res. Conf. Nairobi	p. 208	Comoro Rousette	None.	Comoro Isls, Grand Comoro, near Boboni, 640 m.	Comoro Isls.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href=â€https://www.iucnredlist.org/species/19757/166527449/â€ target=â€_blank>Vulnerable</a>	Subgenus Rousettus. Considered a subspecies of madagascarensis by Peterson et al. (1995), but clearly distinct see Bergmans (1994) and Goodman et al. (2010).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rousettus obliviosus; Rousettus obliviosus; Rousettus obliviosus; Rousettus obliviosus; Rousettus obliviosus; Rousettus obliviosus; obliviosus; Roussette des Comores; Komoren-Flughund; Rosetus de las Comoras; Comoros Rousette; Comoro Rousette; Comoros Rousette; Comoro Rousette; Comoro Rousette; R. obliviosus
