http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1499	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis	Rousettus madagascariensis		[MSW2] Subgenus Rousettus. Considered a subspecies of lanosus by Hayman and Hill (1971:12); but see Bergmans (1977).; [MSW3] Subgenus Stenonycteris. Considered a subspecies of lanosus by Hayman and Hill (1977), but see Bergmans (1977). Revised by Peterson et al. (1995). Does not include obliviosus; see Bergmans (1994), but also see Peterson et al. (1995).; [HMW] Rousettus madagascariensis G. Grandidier, 1929 , surroundings of Beforona, Madagascar . Rousettus madagascariensis is closely related to, but clearly distinct from, other African rousettes ( R. aegyptiacus and R. obliviosus ). Monotypic.; [batnames2022] Subgenus Stenonycteris . Considered a subspecies of Stenonycteris lanosus by Hayman and Hill (1977), but see Bergmans (1977). Revised by Peterson et al. (1995). Does not include obliviosus ; see Bergmans (1994)and Goodman et al. (2010). See Peterson et al. (1995) for an alternative view. Issue date of the publication may be as late as 1930; see Helgen and McFadden (2001). We follow Bergmans (1994) in considering the issue date 1929.; [batnames2023] Subgenus Stenonycteris . Considered a subspecies of Stenonycteris lanosus by Hayman and Hill (1977), but see Bergmans (1977). Revised by Peterson et al. (1995). Does not include obliviosus ; see Bergmans (1994)and Goodman et al. (2010). See Peterson et al. (1995) for an alternative view. Issue date of the publication may be as late as 1930; see Helgen and McFadden (2001). We follow Bergmans (1994) in considering the issue date 1929.; [batnames2025_1.7] Subgenus Stenonycteris. Considered a subspecies of Stenonycterislanosus by Hayman and Hill (1977), but see Bergmans (1977). Revised by Peterson et al. (1995). Does not include obliviosus; see Bergmans (1994)and Goodman et al. (2010). See Peterson et al. (1995) for an alternative view. Issue date of the publication may be as late as 1930; see Helgen and McFadden (2001). We follow Bergmans (1994) in considering the issue date 1929.														madagascariensis				madagascariensis	madagascariensis			madagascariensis G. Grandidier, 1929		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Madagascar; ref. 4.17	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rousettus madagascariensis	Madagascar, Beforona (between Tananarive and Andevoranto).	G. Grandidier	1928	Bull. Acad. Malgache, 11:91.	Distribution: The species is known only from Madagascar.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Madagascar rousette	Madagascar	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	G. Grandidier	1928	Bull. Acad. Malgache, 11:91.	Subgenus Rousettus. Considered a subspecies of lanosus by Hayman and Hill (1971:12); but see Bergmans (1977).	Madagascar.	Madagascar, Beforona (between Tananarive and Andevoranto).		GRANDIDIER	1929	A small species (forearm length, 65-76 mm). Molars relatively narrow. Median edge of pla giopatagium attached to side of body. Uropatagium sparsely haired. Body fur fairly short.	Distribution: The species is known only from Madagascar.	No subspecies.		20	species	R. madagascariensis	GRANDIDIER	1929	Rousettus	subgenus	Rousettus madagascariensis				A small species (forearm length, 65-76 mm). Molars relatively narrow. Median edge of pla giopatagium attached to side of body. Uropatagium sparsely haired. Body fur fairly short.	No subspecies.		6. R. madagascariensis GRANDIDIER 1929.	6	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Rousettus madagascariensis	Rousettus	Stenonycteris	madagascariensis	G. Grandidier		1928		Bull. Acad. Malgache, N.S.	11		91		Malagasy Rousette	Madagascar, Beforona (between Tananarive [= Antananarivo] and Andevoranto).	Madagascar except SW region.	IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Lower Risk (nt).		Subgenus Stenonycteris. Considered a subspecies of lanosus by Hayman and Hill (1977), but see Bergmans (1977). Revised by Peterson et al. (1995). Does not include obliviosus; see Bergmans (1994), but also see Peterson et al. (1995).	03AD87FAFFDAF6358CBB3313FEF3FB7E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	88	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFDAF63489B63D13F685F959.xml	Rousettus madagascariensis	Pteropodidae	Rousettus	madagascariensis	G. Grandidier	1929	Roussette de Madagascar @fr | Madagaskar Flughund @de | Rosetus de Madagascar @es | Madagascan Rousette @en | Madagascar Rousette @en	Rousettus madagascariensis G. Grandidier, 1929 , surroundings of Beforona, Madagascar . Rousettus madagascariensis is closely related to, but clearly distinct from, other African rousettes ( R. aegyptiacus and R. obliviosus ). Monotypic.	Madagascar except C highlands and arid SW region.	Head-body 119- 143 mm , tail 6~13- 5 mm , ear 13- 5-17 mm , hindfoot 16-19 mm , forearm 66-80 mm ; weight 31-81- 5 g . Skull is delicately built, premaxillae are not co-ossified and projected forward, braincase is moderately deflect ed, and alveolar line projects backward passing through upper part of or just above condylus occipitalis or just above foramen magnum. Fur is somewhat longer, denser, and more persistent than that of the Egyptian Rousette ( R. aegyptiacus ). Dorsal hairs and on top of head are light grayish to brownish at their bases and darker brown at tips. Overall color is brown, with reddish brown and grayish hues. Fur is lighter on venter and sparser on neck, shoulders, and throat. Uropatagium is sparsely furred in some zones. Dorsal side oftibia is practically naked. Wings attach between first and second toes. Postorbital width is larger than interorbital width in most specimens, supraoccipital crest is low, and zygomatic arches are slender. Palatal ridges arranged ina 4 + 3 + 1 pattern. Cheekteeth are relatively narrow, with widths of premolars and molars about equal to or slightly longer than one-half their lengths, except for M* and M, in which width is distinctly longer. Diploid number is 2n = 36.	Eastern humid forests, western, and north-western dry deciduous forests, plantations and other agricultural areas, and villages from sea level up to elevations of c. 1150 m . The Malagasy Rousette is rare or absent in central Madagascar Highlands and arid south-western Madagascar .	Few seeds were found in feces of Malagasy Rousettes, suggesting that they either feed on fruits with seeds too large to ingest or they use other food sources preferentially, such as nectar, flower parts, and leaves. Ficus sp. ( Moraceae ) seeds were found in feces in March-April, and individuals were captured near fruiting fig trees. Feeding observation in captivity showed that Madagascan fruit bats feed on fruits that they consume by ingesting fruit juice and pulp and spitting out the fiber. They preferred lipid- and calcium-rich fruits, and fruits of no commercial value over commercially important ones. Nectar appears to be an important dietary element; bats have been observed feeding on kapok flowers ( Ceiba , Malvaceae ) and on nectar of banana flowers.	Pregnancy and lactation of Malagasy Rousettes coincide with summer and higher food availability. In Maromizaha Forest (eastern Madagascar ), pregnancy was recorded in October and December, and lactating females were caught in December—January. Weaning of young occurs at c.8 weeks old and sexual maturity at c.1 year.	The Malagasy Rousette roosts in caves and rock shelters. Because it occurs in areas far from caves, it might roost at othersites such as tree holes. It is high mobile below forest canopies.	Roosting sites of Malagasy Rousettes can be 8 km from food sources. Foraging routes can include round trips of up to 27 km , according to radio-tracking observations. Colonies range from a few hundred individuals to over a thousand. A genetic study using cytochrome-b and six microsatellite loci surveyed individuals throughout the distribution and found no geographic structure in populations.	Classified as Near Threatened on The IUCN Red List. Main threat to the Malagasy Rousette is subsistence hunting that is common in roost caves. Disturbance of caves by tourism and mining activities also are probably threats to colonies. Although Malagasy Rousettes are not strictly dependent on intact forests, impact of forest loss has not been evaluated but might impact their foraging behavior. Presently recorded in six protect areas: Ankarana, Analamerana, and Tsingy de Namoroka reserves and Ankarafantsika, Tsingy de Bemaraha, and Isalo national parks.	Andriafidison et al. (2008a) | Andrianaivoarivelo, Jenkins et al. (2012) | Andrianaivoarivelo, Ramilijaona et al. (2011) | Cardiff et al. (2009) | Goodman, Andriafidison et al. (2005) | Goodman, Chan et al. (2010) | Goodman, Rajemison et al. (2017) | Jenkins & Racey (2008) | Richards et al. (2016)		47. Malagasy Rousette Rousettus madagascariensis French: Roussette de Madagascar / German: MadagaskarFlughund / Spanish: Rosetus de Madagascar Other common names: Madagascan Rousette , Madagascar Rousette Taxonomy. Rousettus madagascariensis G. Grandidier, 1929 , surroundings of Beforona, Madagascar . Rousettus madagascariensis is closely related to, but clearly distinct from, other African rousettes ( R. aegyptiacus and R. obliviosus ). Monotypic. Distribution. Madagascar except C highlands and arid SW region. Descriptive notes. Head-body 119- 143 mm , tail 6~13- 5 mm , ear 13- 5-17 mm , hindfoot 16-19 mm , forearm 66-80 mm ; weight 31-81- 5 g . Skull is delicately built, premaxillae are not co-ossified and projected forward, braincase is moderately deflect ed, and alveolar line projects backward passing through upper part of or just above condylus occipitalis or just above foramen magnum. Fur is somewhat longer, denser, and more persistent than that of the Egyptian Rousette ( R. aegyptiacus ). Dorsal hairs and on top of head are light grayish to brownish at their bases and darker brown at tips. Overall color is brown, with reddish brown and grayish hues. Fur is lighter on venter and sparser on neck, shoulders, and throat. Uropatagium is sparsely furred in some zones. Dorsal side oftibia is practically naked. Wings attach between first and second toes. Postorbital width is larger than interorbital width in most specimens, supraoccipital crest is low, and zygomatic arches are slender. Palatal ridges arranged ina 4 + 3 + 1 pattern. Cheekteeth are relatively narrow, with widths of premolars and molars about equal to or slightly longer than one-half their lengths, except for M* and M, in which width is distinctly longer. Diploid number is 2n = 36. Habitat. Eastern humid forests, western, and north-western dry deciduous forests, plantations and other agricultural areas, and villages from sea level up to elevations of c. 1150 m . The Malagasy Rousette is rare or absent in central Madagascar Highlands and arid south-western Madagascar . Food and Feeding. Few seeds were found in feces of Malagasy Rousettes, suggesting that they either feed on fruits with seeds too large to ingest or they use other food sources preferentially, such as nectar, flower parts, and leaves. Ficus sp. ( Moraceae ) seeds were found in feces in March-April, and individuals were captured near fruiting fig trees. Feeding observation in captivity showed that Madagascan fruit bats feed on fruits that they consume by ingesting fruit juice and pulp and spitting out the fiber. They preferred lipid- and calcium-rich fruits, and fruits of no commercial value over commercially important ones. Nectar appears to be an important dietary element; bats have been observed feeding on kapok flowers ( Ceiba , Malvaceae ) and on nectar of banana flowers. Breeding. Pregnancy and lactation of Malagasy Rousettes coincide with summer and higher food availability. In Maromizaha Forest (eastern Madagascar ), pregnancy was recorded in October and December, and lactating females were caught in December—January. Weaning of young occurs at c.8 weeks old and sexual maturity at c.1 year. Activity patterns. The Malagasy Rousette roosts in caves and rock shelters. Because it occurs in areas far from caves, it might roost at othersites such as tree holes. It is high mobile below forest canopies. Movements, Home range and Social organization. Roosting sites of Malagasy Rousettes can be 8 km from food sources. Foraging routes can include round trips of up to 27 km , according to radio-tracking observations. Colonies range from a few hundred individuals to over a thousand. A genetic study using cytochrome-b and six microsatellite loci surveyed individuals throughout the distribution and found no geographic structure in populations. Status and Conservation. Classified as Near Threatened on The IUCN Red List. Main threat to the Malagasy Rousette is subsistence hunting that is common in roost caves. Disturbance of caves by tourism and mining activities also are probably threats to colonies. Although Malagasy Rousettes are not strictly dependent on intact forests, impact of forest loss has not been evaluated but might impact their foraging behavior. Presently recorded in six protect areas: Ankarana, Analamerana, and Tsingy de Namoroka reserves and Ankarafantsika, Tsingy de Bemaraha, and Isalo national parks. Bibliography. Andriafidison et al. (2008a), Andrianaivoarivelo, Jenkins et al. (2012), Andrianaivoarivelo, Ramilijaona et al. (2011), Cardiff et al. (2009), Goodman, Andriafidison et al. (2005), Goodman, Chan et al. (2010), Goodman, Rajemison et al. (2017), Jenkins & Racey (2008), Richards et al. (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Rousettus madagascariensis	Rousettus	Stenonycteris	madagascariensis	G. Grandidier	1929	0	Bull. Acad. Malgache, N.S.	0.5215	Malagasy Rousette	None.	Madagascar, Beforona (between Tananarive [= Antananarivo] and Andevoranto).	Madagascar except SW region.	Not listed.	Vulnerable	Subgenus Stenonycteris . Considered a subspecies of Stenonycteris lanosus by Hayman and Hill (1977), but see Bergmans (1977). Revised by Peterson et al. (1995). Does not include obliviosus ; see Bergmans (1994)and Goodman et al. (2010). See Peterson et al. (1995) for an alternative view. Issue date of the publication may be as late as 1930; see Helgen and McFadden (2001). We follow Bergmans (1994) in considering the issue date 1929.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rousettus madagascariensis	23	Malagasy Rousette	Madagascan Rousette|Madagascar Rousette	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	ROUSETTINAE	ROUSETTINI	Rousettus	NA	madagascariensis	G. Grandidier	1929	0	Rousettus_madagascariensis	Grandidier, G. (1929). Nouvelle espece de Chauve-Souris frugivore Rousettus madagascariensis G. Grand. Bulletin AcadÃ©mie Malgache (n.s.), 11 (for 1928), 91.		MCZ 45432		surroundings of Beforona, Madagascar.			madagascariensis G. Grandidier, 1929	NA	NA	Madagascar	Africa	Afrotropic	VU	0	0	0	Rousettus_madagascariensis	0	sciname match	Rousettus_madagascariensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19750	Rousettus madagascariensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Rousettus	madagascariensis	G. Grandidier, 1928		20000000	Rousettus madagascariensis	Vulnerable	A2d	2019	2018-08-31 00:00:00 UTC	3.1	English	This species is listed as Vulnerable as it is suspected to have undergone a decline in the region of more than 30% over the last three generations (15 years) as a result of chronic hunting. Evidence has accumulated on the widespread hunting of this species for bushmeat throughout Madagascar (Andrianaivoarivelo et al. 2012a; Cardiff et al. 2009; Golden 2005, 2009; Jenkins and Racey 2008; Rakotondrasana and Goodman 2007).	It roosts in caves well beyond the twilight zone (MacKinnon et al. 2003). Few colonies are known to biologists even in areas with extensive underground cavities (MacKinnon et al. 2003, Kofoky et al. 2007) suggesting that they have specific roosting requirements (Andrianaivoarivelo et al. 2011a). It is also recorded from a number of locations that are without known roosts or caves (Andriafidison et al. 2006, Rakotoarivelo and Randrianandriananina 2007, Rakotonandrasana and Goodman 2007) and it may therefore roost in other sites such as tree holes. Although it is widely distributed it appears to be associated with forests (MacKinnon et al. 2003), is able to fly within relatively intact forest (Kofoky et al. 2007) and has been trapped inside Eucalyptus plantations (Randrianandriananina et al. 2006), in agricultural settings (Andrianaivoarivelo et al. 2011b, Randrianandriananina et al. 2006, Kofoky et al. 2007, Rakotoarivelo and Randrianandriananina 2007, Ralisata 2014) and in villages (Andriafidison et al. 2006). There have been several dietary studies. It feed on the fruit of endemic forest trees (Razafindrakoto 2006). Nectar appears to be an important dietary constituent and bats have been observed feeding on banana and kapok flowers (Andrianaivoarivelo et al. 2011b, Andriafidison et al. 2006). In the Maromizha forest in eastern Madagascar, Andrianaivoarivelo et al. (2011b) found relatively few seeds (several Ficus species , Dypsi ssp and Rubus mollucanus ) in the faeces and inferred that R. madagascariensi s fed also on nectar, flowers, leaves and fruit with seeds too large to swallow. Flight cage experiments by Andrianaivoarivelo et al. , (2102b) revealed that it preferred native or introduced fruit of no commercial value (F. polita , Syzigium jambos and S. malaccense ) to commercial species (Litchi chinensis and Diospyros kaki ). In similar experiments, bats were offered F. sakalavarum and Ziziphus jujube . Juveniles fed extensively on the former while adults preferred unripe Ziziphus , whose seeds are too large to swallow. It was concluded that R. madagascariensis is an effective disperser of F. sakalavaru m particularly for isolated trees or those in forest fragments where other frugivores are rare (Andrianaivoarivelo et al. 2012a). Rousettus madagascariensis feeds especially on autochthonous or introduced plant species which grow inside or outside the natural forest ecosystems (Andrianaivoarivelo et al. 2012a, Andrianaivoarivelo et al. 2011b, Andriafidison et al. 2006). It appears to have no predators when foraging, but the barn owl (Tyto alba ) is a predator at roost sites (Goodman and Griffiths 2006). Its feeding behavior seems to be well adapted to the anthropogenic changes of the environment and it contributes to the dispersion of forest seeds (Andrianaivoarivelo et al. 2011b). The morphometric variations of this species are associated with sexual dimorphism and with geographical and altitudinal locations. The body mass also varies with food availability which in turns depends on season. However there was contrast between the morphometric and genetic diversities. Populations sampled at low altitude and in the North have a longer forearm and lower body mass than those at higher altitude, in the South and East of Madagascar. However, molecular genetic studies of samples from throughout Madagascar showed that there was no genetic variations between populations and the species has a panmictic structure (Andrianaivoarivelo 2012, Goodman et al. 2010). The lactation period coincides with the rainy season and weaning is completed by the end of the rain period. The growth of young bats takes place during the dry season and somatic growth is completed within a year (Andrianaivoarivelo 2012).	Although Goodman et al. (2005) suggested that Rousettus madagascariensis  is not dependent on relatively intact forest, the extent to which the destruction and degradation of natural forest threatens this species is not clear. Recent research demonstrated that the removal of forest cover in the east led the bats to abandon their roosts (Andrianaivoarivelo et al. 2011a). Some colonies are located away from intact forest and additional study is therefore needed on the mobility of this species. The main threats to R. madagascariensis are to its roosts where it is subject to hunting pressure (Golden 2005, 2009; Jenkins et al. 2007; Rakotonandrasana and Goodman 2007; Jenkins and Racey 2008) in virtually all sites that are not inside protected areas (Goodman et al. 2005) or considered sacred (Rakotoarivelo and Randrianandriananina 2007). It appears to be hunted exclusively for subsistence and bats are harvested using locally made traps as well as being knocked down from the cave ceiling with wooden batons (Andrianaivoarivelo et al. 2011a, Rakotonandrasana and Goodman 2007). On Ile Sainte Marie the reported offtake was between 360 and 480 bats per year (Rakotonandrasana and Goodman 2007).	It roosts in aggregations of over a thousand individuals (MacKinnon et al. 2003, Rakotoarivelo and Randrianandriananina 2007) but smaller colonies of a few hundred are also known (Andrianaivoarivelo et al. 2011a, Jenkins et al. 2007, Kofoky et al. 2007). It can be locally abundant and is often the most commonly trapped species in mist-netting surveys in the west of Madagascar (Kofoky et al. 2007, Rakotoarivelo and Randrianandriananina 2007) and in the southeast (Ralisata 2014).	Decreasing	This species is endemic to Madagascar and the adjacent islands Nosy Be, Nosy Komba and Sainte Marie. It is generally widespread but rare or absent from the central highlands and the arid south-west (MacKinnon et al. 2003, Goodman et al. 2005).	It appears to be hunted exclusively for subsistence and bats are harvested using locally made traps as well as being knocked down from the cave ceiling with wooden batons (Andrianaivoarivelo et al. 2011a, Rakotonandrasana and Goodman 2007).	Terrestrial	As a game species under Malagasy law (Durbin 2007), R. madagascariensis is only protected when it occurs in nature reserves. In a survey of western Madagascar it was found in six protected areas: Parcs National dâ€™Ankarana, Ankarafantsika, Namoroka, Tsingy de Bemaraha, Isalo and RÃ©serve SpÃ©ciale dâ€™Analamerana (Goodman et al. 2005). Roosts within existing protected areas should receive close attention from park staff to discourage hunting. Other roosts need to be conserved and this might be best achieved through their inclusion within new protected areas and with the cooperation of local communities.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Rousettus	Stenonycteris	madagascariensis	G. Grandidier	1929	0	Bull. Acad. Malgache, N.S.	0.521528	Malagasy Rousette	None.	Madagascar, Beforona (between Tananarive [= Antananarivo] and Andevoranto).	Madagascar except SW region.	Not listed.	Vulnerable	Subgenus Stenonycteris . Considered a subspecies of Stenonycteris lanosus by Hayman and Hill (1977), but see Bergmans (1977). Revised by Peterson et al. (1995). Does not include obliviosus ; see Bergmans (1994)and Goodman et al. (2010). See Peterson et al. (1995) for an alternative view. Issue date of the publication may be as late as 1930; see Helgen and McFadden (2001). We follow Bergmans (1994) in considering the issue date 1929.	Rousettus madagascariensis	1004544	23	Malagasy Rousette	Madagascan Rousette|Madagascar Rousette	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	ROUSETTINAE	ROUSETTINI	Rousettus	NA	madagascariensis	G. Grandidier	1929	0	Rousettus_madagascariensis	Grandidier, G. (1929). Nouvelle espece de Chauve-Souris frugivore Rousettus madagascariensis G. Grand. Bulletin AcadÃ©mie Malgache (n.s.), 11 (for 1928), 91.		MCZ 45432		surroundings of Beforona, Madagascar.			madagascariensis G. Grandidier, 1929	NA	NA				Madagascar	Africa	Afrotropic	VU	0	0	0	Rousettus_madagascariensis	0	sciname match	Rousettus_madagascariensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rousettus_madagascariensis	1004544	23	Malagasy Rousette	Madagascan Rousette|Madagascar Rousette	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Epomophorinae	Rousettini	Rousettus	NA	madagascariensis	G. Grandidier	0	Rousettus madagascariensis	Grandidier, G. 1929. Nouvelle espÃ¨ce Chauve-Souris frugivore, _Rousettus madagascariensis_ G. Grand. Bulletin de l'AcadÃ©mie malgache (n.s.)11:91-93.		MCZ:Mamm:45432	holotype	https://mczbase.mcz.harvard.edu/guid/MCZ:Mamm:45432	surroundings of Beforona, Madagascar.			NA	NA				Madagascar	Africa	Afrotropic	VU	0	0	0	Rousettus_madagascariensis	0	sciname match	Rousettus_madagascariensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Rousettus	Rousettus	madagascariensis	G. Grandidier	1929	0	Bull. Acad. Malgache, N.S.	0.521528	Malagasy Rousette	None.	Madagascar, Beforona (between Tananarive [= Antananarivo] and Andevoranto).	Madagascar except SW region.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href=â€https://www.iucnredlist.org/species/19750/22002909/â€ target=â€_blank>Vulnerable</a>	Subgenus Stenonycteris. Considered a subspecies of Stenonycterislanosus by Hayman and Hill (1977), but see Bergmans (1977). Revised by Peterson et al. (1995). Does not include obliviosus; see Bergmans (1994)and Goodman et al. (2010). See Peterson et al. (1995) for an alternative view. Issue date of the publication may be as late as 1930; see Helgen and McFadden (2001). We follow Bergmans (1994) in considering the issue date 1929.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rousettus madagascariensis; Rousettus madagascariensis; Rousettus madagascariensis; Rousettus madagascariensis; Rousettus madagascariensis; Rousettus madagascariensis; madagascariensis; Roussette de Madagascar; Madagaskar Flughund; Rosetus de Madagascar; Madagascan Rousette; Madagascar Rousette; Malagasy Rousette; Madagascan Rousette; Madagascar Rousette; Malagasy Rousette; Malagasy Rousette; R. madagascariensis
