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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1491	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso	Rhynchonycteris naso		[MSW3] See Plumpton and Jones (1992); see Emmons (1997) for an updated distribution map.; [HMW] Vespertilio naso Wied-Neuwied , 1820 , “ Mucuri.” Restricted by F. D. de Avila- Pires in 1965 to “Vcinity of Morro d’Arara, Rio Mucuri , State of Bahia , Brazil .” This species is monotypic.; [batnames2022] See Plumpton and Jones (1992); see Emmons (1997) for an updated distribution map.; [IUCN] The genus is monotypic.; [batnames2023] See Plumpton and Jones (1992); see Emmons (1997) for an updated distribution map.; [MDD2025_2.0] likely represents a species compex; [batnames2025_1.7] See Plumpton and Jones (1992); see Emmons (1997) for an updated distribution map.; [MDD2025_2.2] likely represents a species compex		(Tufted bat)				lineata, priscus, rivalis, saxatilis, villosa.			lineata, priscus, rivalis, saxatilis, villosa			naso	naso - lineata, priscus, rivalis, saxatilis, villosa	naso, rivalis, saxatilis, lineata, villosa, priscus	The genus is monotypic.	naso	naso - lineata, priscus, rivalis, saxatilis, villosa	naso, rivalis, saxatilis, lineata, villosa, priscus	naso, rivalis, saxatilis, lineata, villosa, prisca, nasio	naso 	naso - lineata, priscus, rivalis, saxatilis, villosa	naso (zu Wied-Neuwied, 1820)|rivalis (von Spix, 1823)|saxatilis (von Spix, 1823)|lineata (Temminck, 1841)|villosa (P. Gervais, 1855)|prisca (G. M. Allen, 1914)|nasio (Dalquest, 1957) [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Proboscis bat	S Mexico – Peru, Brazil	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhynchonycteris naso	Brazil, Bahia, Rio Mucuri, near Morro d'Arara.	Wied-Neuwied	1820	Reise nach Brasilien, 1:251.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Long-nosed bat (Tufted bat)	S Mexico – Bolivia, Brazil; Trinidad	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Wied-Neuwied	1820	Reise nach Brasilien, 1:251.		E Oaxaca and C Veracruz (Mexico) to C and E Brazil, Peru, Bolivia, French Guiana, Guyana, and Surinam; Trinidad.	Brazil, Bahia, Rio Mucuri, near Morro d'Arara; for clarification see Avila-Pires (1965:9).		WIED-NEUWIED	1820	Size relatively small (forearm length, 35-41 mm).	Distribution: Same as for genus.	No subspecies.		46	species	R. naso	WIED-NEUWIED	1820	Rhynchonycteris	genus	Rhynchonycteris naso				Size relatively small (forearm length, 35-41 mm).	No subspecies.		1. R. naso (WIED-NEUWIED 1820).	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Emballonurinae		Rhynchonycteris naso	Rhynchonycteris		naso	Wied-Neuwied	y	1820		Reise nach Brasilien	1		251		Proboscis Bat	Brazil, Bahia, Rio Mucuri, near Morro d'Arara; for clarification see Avila-Pires (1965:9).	E Oaxaca and C Veracruz (Mexico) to C and E Brazil, Peru, Bolivia, French Guiana, Guyana, and Surinam; Trinidad.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	lineata Temminck, 1838; priscus G. M. Allen, 1914; rivalis Spix, 1823; saxatilis Spix, 1823; villosa Gervais, 1855.	See Plumpton and Jones (1992); see Emmons (1997) for an updated distribution map.	4C3D87E8FFB86A00FA7D9B781A21B08B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	370	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFDA4C12F829258DFA62F493.xml	Rhynchonycteris naso	Emballonuridae	Rhynchonycteris	naso	Wied-Neuwied	1820	Proboscis Bat @en | Nasin des rivières @fr | Nasenfledermaus @de | Murciélago narigón @es | Brazilian Long-nosed Bat @en | Long-nosed Proboscis Bat @en | River Bat @en | Sharp-nosed Bat @en	Vespertilio naso Wied-Neuwied , 1820 , “ Mucuri.” Restricted by F. D. de Avila- Pires in 1965 to “Vcinity of Morro d’Arara, Rio Mucuri , State of Bahia , Brazil .” This species is monotypic.	From S Mexico (Veracruz and Oaxaca) S throughout CentraAmerica to Colombia, E to C & E Brazil, and S to E Ecuador, E Peru, and N & NE Bolivia and also on Trinidad I.	Head-body 36-48 mm, tail 11-17 mm, ear 11-14 mm, hindfoot 6-8 mm, forearm 36-40 mm; weight 3-6 g. Long, fluffy dorsal fur of the Proboscis Bat is grizzled gray-brown, with yellow tones and two faint, wavy, cream-colored stripes extend from shoulders to rump. Venter is pale yellow or grayish. Nose is long and projects well beyond lower lip. Flight membranes and facial skin are black. Tufts of whitish fur on forearms are diagnostic. Wing sacs are absent. Calcar is about three times as long as feet, and uropatagium is longer than legs. Dental formula is 11/3, C 1/1, P 2/2, M 3/3 (x2) = 32.	Almost always associated with water and wedands (streams, rivers, mangroves, and lakes) near multi-strata evergreen forests, semideciduous forests, dry forests, secondary forests, croplands, or pastures from sea level to elevations of c.1500 m ( generally most common below 500 m).	Proboscis Bats are aerial insectivores and usually forage within 3 m of surface of slow moving, shallow water. Diet includes small dipterans including mosquitoes.	Proboscis Bats are monoestrous; single young are bom once a year in early wet season.	Proboscis Bats are crepuscular, and their foraging flights usually begin well before sunset and continue into darkness. They roost on tree trunks, exposed logs, boulders, and steep banks; under bridges; and in tree cavities, always near or direcdy above water. Roosting individuals usually form a single vertical line. Color pattern of the Proboscis Bat makes it extremely cryptic when it roosts on tree bark. Echolocation call contains a F component at c.100 kHz.	Proboscis Bats usually roost in small, single-species colonies of c.10-24 individuals, but groups of up to 100 individuals have been noted. When roosting, they are often aligned in vertical rows with individual spacing of c.20-40 mm. Several males occur in a roosting group, and there appears to be no harem formation or defense. If disturbed, a group flies together to an alternate roost. At times, all group members rock back and forth like leaves blowing in the wind. Spiders frequently prey on Proboscis Bats. The orb-web spider Argiope submaronica has been observed to feed on adult Proboscis Bats. In Costa Rica, a Proboscis Bat near a roost under eaves of a building was entangled in the center of a web; it was wrapped in silk and fed upon by the spider. The orb-weaver Nephila clavipes also was observed capturing and feeding on Proboscis Bats in Peru.	Classified as Least Concern on 77 zr IUCN Red List. The Proboscis Bat has a large distribution and presumably large and stable overall population, and it is common in areas with suitable wetland habitats. It can be locally common and found in many protected areas in Mexico, Central America, and South America. In Mexico, it is provided special protection.	Avila-Pires (1965) | Dalquest (1957) | Eisenberg (1989) | Emmons & Feer (1997) | Goodwin & Greenhall (1961) | Husson (1978) | Nyffeler & Knörnschild (2013) | Timm & Losilla (2007)	https://zenodo.org/record/3747997/files/figure.png	49 . Proboscis Bat Rhynchonycteris naso French: Nasin des rivières I German: Nasenfledermaus / Spanish: Murciélago narigón Other common names: Brazilian Long-nosed Bat, Long-nosed Proboscis Bat, River Bat, Sharp-nosed Bat Taxonomy . Vespertilio naso Wied-Neuwied , 1820 , “ Mucuri.” Restricted by F. D. de Avila- Pires in 1965 to “Vcinity of Morro d’Arara, Rio Mucuri , State of Bahia , Brazil .” This species is monotypic. Distribution. From S Mexico (Veracruz and Oaxaca) S throughout CentraAmerica to Colombia, E to C & E Brazil, and S to E Ecuador, E Peru, and N & NE Bolivia and also on Trinidad I. Descriptive notes. Head-body 36-48 mm, tail 11-17 mm, ear 11-14 mm, hindfoot 6-8 mm, forearm 36-40 mm; weight 3-6 g. Long, fluffy dorsal fur of the Proboscis Bat is grizzled gray-brown, with yellow tones and two faint, wavy, cream-colored stripes extend from shoulders to rump. Venter is pale yellow or grayish. Nose is long and projects well beyond lower lip. Flight membranes and facial skin are black. Tufts of whitish fur on forearms are diagnostic. Wing sacs are absent. Calcar is about three times as long as feet, and uropatagium is longer than legs. Dental formula is 11/3, C 1/1, P 2/2, M 3/3 (x2) = 32. Habitat . Almost always associated with water and wedands (streams, rivers, mangroves, and lakes) near multi-strata evergreen forests, semideciduous forests, dry forests, secondary forests, croplands, or pastures from sea level to elevations of c.1500 m ( generally most common below 500 m). Food and Feeding . Proboscis Bats are aerial insectivores and usually forage within 3 m of surface of slow moving, shallow water. Diet includes small dipterans including mosquitoes. Breeding . Proboscis Bats are monoestrous; single young are bom once a year in early wet season. Activity patterns. Proboscis Bats are crepuscular, and their foraging flights usually begin well before sunset and continue into darkness. They roost on tree trunks, exposed logs, boulders, and steep banks; under bridges; and in tree cavities, always near or direcdy above water. Roosting individuals usually form a single vertical line. Color pattern of the Proboscis Bat makes it extremely cryptic when it roosts on tree bark. Echolocation call contains a F component at c.100 kHz. Movements, Home range and Social organization. Proboscis Bats usually roost in small, single-species colonies of c.10-24 individuals, but groups of up to 100 individuals have been noted. When roosting, they are often aligned in vertical rows with individual spacing of c.20-40 mm. Several males occur in a roosting group, and there appears to be no harem formation or defense. If disturbed, a group flies together to an alternate roost. At times, all group members rock back and forth like leaves blowing in the wind. Spiders frequently prey on Proboscis Bats. The orb-web spider Argiope submaronica has been observed to feed on adult Proboscis Bats. In Costa Rica, a Proboscis Bat near a roost under eaves of a building was entangled in the center of a web; it was wrapped in silk and fed upon by the spider. The orb-weaver Nephila clavipes also was observed capturing and feeding on Proboscis Bats in Peru. Status and Conservation . Classified as Least Concern on 77 zr IUCN Red List. The Proboscis Bat has a large distribution and presumably large and stable overall population, and it is common in areas with suitable wetland habitats. It can be locally common and found in many protected areas in Mexico, Central America, and South America. In Mexico, it is provided special protection. Bibliography. Avila-Pires (1965), Dalquest (1957), Eisenberg (1989), Emmons & Feer (1997), Goodwin & Greenhall (1961), Husson (1978), Nyffeler & Knörnschild (2013), Timm & Losilla (2007).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Rhynchonycteris naso	Rhynchonycteris		naso	Wied-Neuwied	1820	1	Reise nach Brasilien	0.216	Proboscis Bat	 lineata Temminck, 1838; priscus G. M. Allen, 1914; rivalis Spix, 1823; saxatilis Spix, 1823; villosa Gervais, 1855.	Brazil, Bahia, Rio Mucuri, near Morro d'Arara; for clarification see Avila-Pires (1965:9).	E Oaxaca and C Veracruz (Mexico) to C and E Brazil, Peru, Bolivia, French Guiana, Guyana, and Surinam; Trinidad.	Not listed.	Least Concern	See Plumpton and Jones (1992); see Emmons (1997) for an updated distribution map.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhynchonycteris naso	23	Proboscis Bat	Brazilian Long-nosed Bat|Long-nosed Proboscis Bat|River Bat|Sharp-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	EMBALLONURINAE	DICLIDURINI	Rhynchonycteris	NA	naso	Wied-Neuwied	1820	1	Vespertilio_naso	Wied-Neuwied, M. (1820). Reise nach Brasilien in den Jahren 1815 bis 1817, Heinrich Ludwig Brönner, Frankfurt, 1, 251 (footnote).	https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t4gm9j910&view=1up&seq=307	AMNH 386		"Mucuri." Restricted by F. D. de Ávila-Pires in 1965 to "Vicinity of Morro d'Arara, Rio Mucuri, State of Bahia, Brazil."			naso (Wied-Neuwied, 1820)|rivalis (Spix, 1823)|saxatilis (Spix, 1823)|lineata (Temminck, 1841)|villosa (P. Gervais, 1856)|priscus (G. M. Allen, 1914)	NA	NA	Mexico|Belize|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Rhynchonycteris_naso	0	sciname match	Rhynchonycteris_naso	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19714	Rhynchonycteris naso	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Rhynchonycteris	naso	(Wied-Neuwied, 1820)	The genus is monotypic.	20000000	Rhynchonycteris naso	Least Concern		2016	2016-07-01 00:00:00 UTC	3.1	English	This species is listed as Least Concern because it is widely distributed, common in areas with water and suitable habitat and unlikely to be declining at a rate which would qualify the species for inclusion in one of the threatened categories in the near future.	It is almost always associated with moist areas near multistratal evergreen forests. These bats tend to roost in small, single-species colonies of about ten to twenty-four, on tree trunks, in tree cavities, or in rock caves (Eisenberg 1989). When roosting they are often aligned in vertical rows with individuals about 10 mm apart. Several males occur in a roosting group, and there appears to be no harem formation or defence. These bats are aerial insectivores (Husson 1978, Goodwin and Greenhall 1961); and they tend to feed over water, flying only a short distance above the surface (Eisenberg 1989). In Mexico they have been also reported from secondary forests, crop-lands and grasslands (de Grammont pers. comm.)	There are no major threats to this species. There may be some issues associated with the water bodies nearby their roosts, and from which they obtain their prey (insects).	These bats are often common in lowland forest near water (streams, rivers, mangroves, and lakes) (Reid 2009); widespread (Emmons and Feer 1997). Colonies vary from a few individuals to 100 individuals (Dalquest 1957), although the usual includes less than a dozen bats (Hood and Gardner 2008).	Unknown	This species ranges from Oaxaca and Veracruz, Mexico, to central and eastern Brazil, Peru, Bolivia, Guianas, Suriname and Trinidad (Simmons 2005). It is widely distributed at low elevations, generally below 500 m (Eisenberg 1989) but up to 1,500 m.		Terrestrial	This species occurs in protected areas. It is widely distributed throughout the Neotropics. In Mexico it is listed as subject to special protection under NOM - 059 - SEMARNAT - 2001 (Arroyo-Cabrales pers. comm.).	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Rhynchonycteris		naso	Wied-Neuwied	1820	1	Reise nach Brasilien	1: 248 [footnote]	Proboscis Bat	 lineata Temminck, 1838; priscus G. M. Allen, 1914; rivalis Spix, 1823; saxatilis Spix, 1823; villosa Gervais, 1855.	Brazil, Bahia, Rio Mucuri, near Morro d'Arara; for clarification see Avila-Pires (1965:9)	E Oaxaca and C Veracruz (Mexico) to C and E Brazil, Peru, Bolivia, French Guiana, Guyana, and Surinam; Trinidad	Not listed.	Least Concern	See Plumpton and Jones (1992); see Emmons (1997) for an updated distribution map.	Rhynchonycteris naso	1004792	23	Proboscis Bat	Brazilian Long-nosed Bat|Long-nosed Proboscis Bat|River Bat|Sharp-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	EMBALLONURINAE	DICLIDURINI	Rhynchonycteris	NA	naso	Wied-Neuwied	1820	1	Vespertilio_naso	Wied-Neuwied, M. (1820). Reise nach Brasilien in den Jahren 1815 bis 1817, Heinrich Ludwig Brönner, Frankfurt, 1, 251 (footnote).	https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t4gm9j910&view=1up&seq=307	AMNH 386		"Mucuri." Restricted by F. D. de Ávila-Pires in 1965 to "Vicinity of Morro d'Arara, Rio Mucuri, State of Bahia, Brazil."			naso (Wied-Neuwied, 1820)|rivalis (Spix, 1823)|saxatilis (Spix, 1823)|lineata (Temminck, 1841)|villosa (P. Gervais, 1856)|priscus (G. M. Allen, 1914)	NA	NA				Mexico|Belize|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Rhynchonycteris_naso	0	sciname match	Rhynchonycteris_naso	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhynchonycteris_naso	1004792	23	Proboscis Bat	Brazilian Long-nosed Bat|Long-nosed Proboscis Bat|River Bat|Sharp-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Emballonurinae	Diclidurini	Rhynchonycteris	NA	naso	zu Wied-Neuwied	1	Vespertilio naso	Wied-Neuwied, M. zu. 1820. Reise nach Brasilien in den Jahren 1815 bis 1817. Erster Band. H. L. Brönner, Vienna, 376 pp.	https://www.biodiversitylibrary.org/page/45439689	AMNH MS-386	lectotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-ms-386	"Mucuri." Restricted by F. D. de Ávila-Pires in 1965 to "Vicinity of Morro d'Arara, Rio Mucuri, State of Bahia, Brazil."			likely represents a species compex	Biganzoli-Rangel, A. J., Leon-Alvarado, O. D., Robe, L. J., Meza, M. A., Gutiérrez, E. E., & Paladini, A. (2023). Systematics and phylogeography of bats of the genus Rhynchonycteris (Chiroptera: Emballonuridae): Integrating molecular phylogenetics, ecological niche modeling and morphometric data. Plos one, 18(5), e0285271.				Mexico|Belize|Guatemala|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad and Tobago|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia	North America|South America	Nearctic|Neotropic	LC	0	0	0	Rhynchonycteris_naso	0	sciname match	Rhynchonycteris_naso	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Rhynchonycteris		naso	Wied-Neuwied	1820	1	Reise nach Brasilien	1: 248 [footnote]	Proboscis Bat	lineata Temminck, 1838; priscus G. M. Allen, 1914; rivalis Spix, 1823; saxatilis Spix, 1823; villosa Gervais, 1855.	Brazil, Bahia, Rio Mucuri, near Morro d'Arara; for clarification see Avila-Pires (1965:9)	E Oaxaca and C Veracruz (Mexico) to C and E Brazil, Peru, Bolivia, French Guiana, Guyana, and Surinam; Trinidad	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19714/22010818/' target='_blank'>Least Concern</a>	See Plumpton and Jones (1992); see Emmons (1997) for an updated distribution map.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhynchonycteris naso; Rhynchonycteris naso; Rhynchonycteris naso; Rhynchonycteris naso; Rhynchonycteris naso; Rhynchonycteris naso; lineata; priscus; rivalis; saxatilis; villosa; lineata; priscus; rivalis; saxatilis; villosa; naso; rivalis; saxatilis; lineata; villosa; priscus; Proboscis Bat; Nasin des rivières; Nasenfledermaus; Murciélago narigón; Brazilian Long-nosed Bat; Long-nosed Proboscis Bat; River Bat; Sharp-nosed Bat; Proboscis Bat; Brazilian Long-nosed Bat; Long-nosed Proboscis Bat; River Bat; Sharp-nosed Bat; Proboscis Bat; Proboscis Bat; Rhynchiscus naso; Rynchiscus naso; R. naso