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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1489	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeesa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida	Rhogeessa tumida		[MSW2] Subgenus Rhogeessa. Listed as a subspecies of parvula by Hall and Kelson (1959:196); but also see LaVal (1973b) and Hall (1981:228), who included bombyx and velilla.; [MSW3] Subgenus Rhogeessa. Listed as a subspecies of parvula by Hall and Kelson (1959), but see LaVal (1973b) and Hall (1981). Does not include aeneus; see Audet et al. (1993) and Genoways and Baker (1996). Does not include io; see Genoways and Baker (1996). See Vonhof (2000).; [HMW] Rhogeessa tumida H. Allen, 1866 , “Mirador [ Veracruz ], Mexico .” Previously listed as a race of R. parvula . Until recently included R. bickhami and R. menchuae , which were split based on mitochondrial and nuclear genetic distances; the distances between R. tumida , R. aenea , and R. menchuae were low, and suggest that the first two might be treated as synonyms, although karyological and morphological data do not support this; R. aenea appears to be the most recently evolved of the three. Monotypic.; [batnames2022] Subgenus Rhogeessa . Listed as a subspecies of parvula by Hall and Kelson (1959), but see LaVal (1973 b ) and Hall(1981). Does not include aeneus ; see Audet et al. (1993) and Genoways and Baker (1996). Does not include io ; see Genoways andBaker (1996). See Vonhof (2000).; [IUCN] Listed as a subspecies of parvula . Does not include aeneus ; see Genoways and Baker (1996). Does not include io ; see Genoways and Baker (1996).; [batnames2023] Subgenus Rhogeessa . Listed as a subspecies of parvula by Hall and Kelson (1959), but see LaVal (1973 b ) and Hall(1981). Does not include aeneus ; see Audet et al. (1993) and Genoways and Baker (1996). Does not include io ; see Genoways andBaker (1996). See Vonhof (2000).; [batnames2025_1.7] Subgenus Rhogeessa. Listed as a subspecies of parvula by Hall and Kelson (1959), but see LaVal (1973b) and Hall(1981). Does not include aeneus; see Audet et al. (1993) and Genoways and Baker (1996). Does not include io; see Genoways andBaker (1996). See Vonhof (2000).						aeneus, bombyx, io, riparia, velilla.								tumida	Listed as a subspecies of parvula . Does not include aeneus ; see Genoways and Baker (1996). Does not include io ; see Genoways and Baker (1996).			tumida	tumida			tumida H. Allen, 1866		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		E Mexico – Ecuador, S Brazil, Bolivia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhogeessa tumida	Mexico, Veracruz, Mirador.	H. Allen	1866	Proc. Acad. Nat. Sei. Phila., 18:286.	Distribution: Ranging from northeastern Mexico to western Ecuador and northeastern Brazil.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Central American yellow bat	E Mexico – Ecuador, S Brazil, Bolivia; Trinidad	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	H. Allen	1866	Proc. Acad. Nat. Sci. Philadelphia, 18:286.	Subgenus Rhogeessa. Listed as a subspecies of parvula by Hall and Kelson (1959:196); but also see LaVal (1973b) and Hall (1981:228), who included bombyx and velilla.	Tamaulipas (Mexico) to Ecuador, Bolivia, and NE Brazil; Trinidad and Tobago.	Mexico, Veracruz, Mirador.		H. ALLEN	1866	Ears relatively short. Hairs of dorsum dark with two bands. Size varying from large to small (forearm length, 25-34 mm). Lingual cingulum of upper canine always with at least some indication of cusps. Uropatagium mostly naked. Outer lower incisor usually not reduced. Postorbital constriction poorly developed. Third metacarpal relatively short.	Distribution: Ranging from northeastern Mexico to western Ecuador and northeastern Brazil.	No currently recognized subspecies.		126	species	R. tumida	H. ALLEN	1866	Rhogeesa	subgenus	Rhogeesa tumida				Ears relatively short. Hairs of dorsum dark with two bands. Size varying from large to small (forearm length, 25-34 mm). Lingual cingulum of upper canine always with at least some indication of cusps. Uropatagium mostly naked. Outer lower incisor usually not reduced. Postorbital constriction poorly developed. Third metacarpal relatively short.	No currently recognized subspecies.		1. R. tumida H. ALLEN 1866.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Nycticeiini	Rhogeessa tumida	Rhogeessa	Rhogeessa	tumida	H. Allen		1866		Proc. Acad. Nat. Sci. Phil.	18		286		Black-winged Little Yellow Bat	Mexico, Veracruz, Mirador.	Tamaulipas (Mexico) to N Nicaragua and NW Costa Rica.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Subgenus Rhogeessa. Listed as a subspecies of parvula by Hall and Kelson (1959), but see LaVal (1973b) and Hall (1981). Does not include aeneus; see Audet et al. (1993) and Genoways and Baker (1996). Does not include io; see Genoways and Baker (1996). See Vonhof (2000).	4C3D87E8FF9F6A20FF4593DB1B6BB1DE	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	857	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF986A27FF8B94291A7AB02E.xml	Rhogeessa tumida	Vespertilionidae	Rhogeessa	tumida	H. Allen	1866	Rhogeessa a ailes noires @fr | Schwarzfligel-Gelbfledermaus @de | Rogesade alas negras @es | Central American Yellow Bat @en	Rhogeessa tumida H. Allen, 1866 , “Mirador [ Veracruz ], Mexico .” Previously listed as a race of R. parvula . Until recently included R. bickhami and R. menchuae , which were split based on mitochondrial and nuclear genetic distances; the distances between R. tumida , R. aenea , and R. menchuae were low, and suggest that the first two might be treated as synonyms, although karyological and morphological data do not support this; R. aenea appears to be the most recently evolved of the three. Monotypic.	Atlantic coast of Mexico , from Tamaulipas S to C Tabasco and N Chiapas .	Head-body ¢. 36—44 mm , tail 27-35 mm , ear 11-14- 2 mm , hindfoot 6- 4-8 mm , forearm 27-5-31- 6 mm ; weight 3-5 g . Dorsal fur bicolored, with buff-gray to buff-yellow bases and fuscous-black to pinkish-cinnamon tips (bases and tips not always contrasted); ventral fur unicolored or with buff-brown to light ocherous-buff tips and slightly paler bases; hairs are 3-4 mm long (sometimes up to 5 mm ). Ears short and triangular, with rounded tips; tragus long with rounded tip. In male, prominent odoriferous glands are present on dorsal side of lower pinna. Uropatagium is almost naked, furred only at base; wing membranes also naked and relatively thick; calcar well developed, extending a little over free margin of uropatagium, with developed keel. Skull inclined vertically from anterior border of nasal to occiput, without horizontal portions; sagittal crest present or absent; braincase low; helmet formed by sagittal (if present) and occipital crests is found above occiput region. Lingual cingulum of C' has two small cusps, differing from other congeners, which have smooth canines; I, has smaller cusps than I,, and is about same size or slightly smaller than I. Dental formula for all species of Rhogeessais11/3,C1/1,P 1/2, M 3/3 (x2) = 30. Chromosomal complement has 2n = 34 and FN = 50, with a subtelocentric X-chromosome and a metacentric Y-chromosome.	All major tropical vegetation associations in North America, notably evergreen forests, subdeciduous forests, deciduous forests, open areas, and villages; commonest in slightly disturbed deciduous forests. Sometimes associated with small streams, along trails in ravines, and around trees in anthropic areas. Occurs from sea level to 1800 m , typically below 500 m .	Insectivorous;flies low along streams.	Pregnant females have been captured in May in Tabasco . Lactating females April to June in Oaxaca , Tabasco , Tamaulipas , and Veracruz . Juveniles were captured from May to August.	Nocturnal; shows two peaks of activity, one in first hour after sunset and second an hour before dawn. It seems to use torporas a response to acute thermal stress, but data are lacking on limits of torpor depth and duration. Roostsites include tree branches, hollow trees, and buildings. Echolocation calls are short, broadband, and with maximum energy at frequencies of 50-60 kHz and lowest energy at frequencies of 40-50 kHz.	May form large colonies.	Classified as Least Concern on The IUCN Red List. Widespread but not abundant and generally considered rare. It occurs in protected areas and appears to be tolerant to some degree of habitat modification.	Allen (1866) | Alvarez-Castafieda & Gonzalez-Ruiz (2018) | Arroyo-Cabrales & Baker ( 2014g) | Baird, Hillis et al. (2008, 2009) | Baird, Marchan-Rivadeneira et al. (2012) | Bickham & Baker (1977) | Emmons & Feer 1997) | Genoways & Baker (1996b) | Hall (1981) | Hall & Dalquest (1963) | Hall & Kelson (1959) | Jones, Smith & Turner (1971) | LaVal (1973a) | Miller et al. (2016b) | Reid (2009) | Simmons (2005) | Vonhof (2000)	https://zenodo.org/record/6398246/files/figure.png	206. Black-winged Little Yellow Bat Rhogeessa tumida French: Rhogeessa a ailes noires / German: Schwarzfligel-Gelbfledermaus / Spanish: Rogesa de alas negras Other common names: Central American Yellow Bat Taxonomy. Rhogeessa tumida H. Allen, 1866 , “Mirador [ Veracruz ], Mexico .” Previously listed as a race of R. parvula . Until recently included R. bickhami and R. menchuae , which were split based on mitochondrial and nuclear genetic distances; the distances between R. tumida , R. aenea , and R. menchuae were low, and suggest that the first two might be treated as synonyms, although karyological and morphological data do not support this; R. aenea appears to be the most recently evolved of the three. Monotypic. Distribution. Atlantic coast of Mexico , from Tamaulipas S to C Tabasco and N Chiapas . Descriptive notes. Head-body ¢. 36—44 mm , tail 27-35 mm , ear 11-14- 2 mm , hindfoot 6- 4-8 mm , forearm 27-5-31- 6 mm ; weight 3-5 g . Dorsal fur bicolored, with buff-gray to buff-yellow bases and fuscous-black to pinkish-cinnamon tips (bases and tips not always contrasted); ventral fur unicolored or with buff-brown to light ocherous-buff tips and slightly paler bases; hairs are 3-4 mm long (sometimes up to 5 mm ). Ears short and triangular, with rounded tips; tragus long with rounded tip. In male, prominent odoriferous glands are present on dorsal side of lower pinna. Uropatagium is almost naked, furred only at base; wing membranes also naked and relatively thick; calcar well developed, extending a little over free margin of uropatagium, with developed keel. Skull inclined vertically from anterior border of nasal to occiput, without horizontal portions; sagittal crest present or absent; braincase low; helmet formed by sagittal (if present) and occipital crests is found above occiput region. Lingual cingulum of C' has two small cusps, differing from other congeners, which have smooth canines; I, has smaller cusps than I,, and is about same size or slightly smaller than I. Dental formula for all species of Rhogeessais11/3,C1/1,P 1/2, M 3/3 (x2) = 30. Chromosomal complement has 2n = 34 and FN = 50, with a subtelocentric X-chromosome and a metacentric Y-chromosome. Habitat. All major tropical vegetation associations in North America, notably evergreen forests, subdeciduous forests, deciduous forests, open areas, and villages; commonest in slightly disturbed deciduous forests. Sometimes associated with small streams, along trails in ravines, and around trees in anthropic areas. Occurs from sea level to 1800 m , typically below 500 m . Food and Feeding. Insectivorous;flies low along streams. Breeding. Pregnant females have been captured in May in Tabasco . Lactating females April to June in Oaxaca , Tabasco , Tamaulipas , and Veracruz . Juveniles were captured from May to August. Activity patterns. Nocturnal; shows two peaks of activity, one in first hour after sunset and second an hour before dawn. It seems to use torporas a response to acute thermal stress, but data are lacking on limits of torpor depth and duration. Roostsites include tree branches, hollow trees, and buildings. Echolocation calls are short, broadband, and with maximum energy at frequencies of 50-60 kHz and lowest energy at frequencies of 40-50 kHz. Movements, Home range and Social organization. May form large colonies. Status and Conservation. Classified as Least Concern on The IUCN Red List. Widespread but not abundant and generally considered rare. It occurs in protected areas and appears to be tolerant to some degree of habitat modification. Bibliography. Allen (1866), Alvarez-Castafieda & Gonzalez-Ruiz (2018), Arroyo-Cabrales & Baker ( 2014g ), Baird, Hillis et al. (2008, 2009), Baird, Marchan-Rivadeneira et al. (2012), Bickham & Baker (1977), Emmons & Feer 1997), Genoways & Baker (1996b), Hall (1981), Hall & Dalquest (1963), Hall & Kelson (1959), Jones, Smith & Turner (1971), LaVal (1973a), Miller et al. (2016b), Reid (2009), Simmons (2005), Vonhof (2000).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Rhogeessa tumida	Rhogeessa		tumida	H. Allen	1866	0	Proc. Acad. Nat. Sci. Phil.	0.9486	Black-winged Little Yellow Bat	None.	Mexico, Veracruz, Mirador.	Tamaulipas to Tehuantepec (Mexico).	Not listed.	Least Concern	Subgenus Rhogeessa . Listed as a subspecies of parvula by Hall and Kelson (1959), but see LaVal (1973 b ) and Hall(1981). Does not include aeneus ; see Audet et al. (1993) and Genoways and Baker (1996). Does not include io ; see Genoways andBaker (1996). See Vonhof (2000).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhogeessa tumida	23	Black-winged Little Yellow Bat	Central American Yellow Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	ANTROZOINI	Rhogeessa	Rhogeessa	tumida	H. Allen	1866	0	Rhogeessa_tumida	Allen, H. (1866). Notes on the Vespertilionidae of Tropical America. Proceedings of the Academy of Natural Sciences of Philadelphia, 18, 286.	https://www.biodiversitylibrary.org/page/1861739#page/294/mode/1up	ANSP 1831/USNM 84021		"Mirador [Veracruz], Mexico."			tumida H. Allen, 1866	NA	NA	Mexico	North America	Nearctic|Neotropic	LC	0	0	0	Rhogeessa_tumida	0	sciname match	Rhogeessa_tumida	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19685	Rhogeessa tumida	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Rhogeessa	tumida	H. Allen, 1866	Listed as a subspecies of parvula . Does not include aeneus ; see Genoways and Baker (1996). Does not include io ; see Genoways and Baker (1996).	20000000	Rhogeessa tumida	Least Concern		2016	2016-08-08 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its wide distribution, presumed large population, occurrence in a number of protected areas, tolerance to some degree of habitat modification, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	This bat inhabits in a variety of habitats, including evergreen and deciduous forest, open areas, and villages. It appears to favor slightly disturbed, deciduous forest (Reid 2009). Roosts in buildings and hollow trees, and colonies may be large. This is one of the first bats to appear at sunset, often flying low to the ground along wide trails on roads. There are two peaks of activity, for an hour after sunset and within an hour of dawn. It feeds on small flying insects, and individuals appear to have established hunting routes. Echolocation calls are short, with maximum energy at 50 to 60 kHz. In Belize, synchronized births occur once a year in the rainy season, and litter size is usually two (LaVal 1973). Within limits imposed by its elevational distribution, this species is known from almost every major vegetation association in tropical North America (LaVal 1973). The single specimen reported by Carter and Jones (1978) in the state of Hidalgo, Mexico, was captured over a stream. At Diriamba, Nicaragua, R. tumida was captured over small streams, along a trail in a small ravine, and along trees bordering a hacienda (Jones et al. 1971). Roosting habits of R. tumida are unknown. The only recorded instance of predation on R. tumida is by a male Vampyrum spectrum in northern Guatemala (McCarthy 1987).	There are no major threats to this species throughout its range.	This bat is uncommon to locally common (Reid 2009); apparently rare but very widespread (Emmons and Feer 1997).	Stable	This species is known from Tamaulipas (Mexico) to north Nicaragua and northwestern Costa Rica (Vonhof 2000, Simmons 2005). It occurs from lowlands to 1,500 m (Reid 2009).		Terrestrial	Rhogeessa tumida occurs in protected areas.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Rhogeessa		tumida	H. Allen	1866	0	Proc. Acad. Nat. Sci. Phil.	0.948611	Black-winged Little Yellow Bat	None.	Mexico, Veracruz, Mirador.	Tamaulipas to Tehuantepec (Mexico).	Not listed.	Least Concern	Subgenus Rhogeessa . Listed as a subspecies of parvula by Hall and Kelson (1959), but see LaVal (1973 b ) and Hall(1981). Does not include aeneus ; see Audet et al. (1993) and Genoways and Baker (1996). Does not include io ; see Genoways andBaker (1996). See Vonhof (2000).	Rhogeessa tumida	1005506	23	Black-winged Little Yellow Bat	Central American Yellow Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	ANTROZOINI	Rhogeessa	NA	tumida	H. Allen	1866	0	Rhogeessa_tumida	Allen, H. (1866). Notes on the Vespertilionidae of Tropical America. Proceedings of the Academy of Natural Sciences of Philadelphia, 18, 286.	https://www.biodiversitylibrary.org/page/1861739#page/294/mode/1up	ANSP 1831/USNM 84021		"Mirador [Veracruz], Mexico."			tumida H. Allen, 1866	NA	NA				Mexico	North America	Nearctic|Neotropic	LC	0	0	0	Rhogeessa_tumida	0	sciname match	Rhogeessa_tumida	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhogeessa_tumida	1005506	23	Black-winged Little Yellow Bat	Central American Yellow Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Antrozoini	Rhogeessa	NA	tumida	H. Allen	0	Rhogeëssa tumida	Allen, H. 1866-08-28. Notes on the Vespertilionidae of tropical America. Proceedings of the Academy of Natural Sciences of Philadelphia 18:279-288.	https://www.biodiversitylibrary.org/page/1861746	USNM:MAMM:84021, ANSP 1831	holotype	http://n2t.net/ark:/65665/3199ca19d-b2d5-4286-86df-e86a980c9c9f	"Mirador [Veracruz], Mexico."			NA	NA				Mexico	North America	Nearctic|Neotropic	LC	0	0	0	Rhogeessa_tumida	0	sciname match	Rhogeessa_tumida	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Rhogeessa		tumida	H. Allen	1866	0	Proc. Acad. Nat. Sci. Phil.	0.948611	Black-winged Little Yellow Bat	None.	Mexico, Veracruz, Mirador.	Tamaulipas to Tehuantepec (Mexico).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19685/22006890/' target='_blank'>Least Concern</a>	Subgenus Rhogeessa. Listed as a subspecies of parvula by Hall and Kelson (1959), but see LaVal (1973b) and Hall(1981). Does not include aeneus; see Audet et al. (1993) and Genoways and Baker (1996). Does not include io; see Genoways andBaker (1996). See Vonhof (2000).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhogeessa tumida; Rhogeessa tumida; Rhogeessa tumida; Rhogeessa tumida; Rhogeessa tumida; Rhogeessa tumida; tumida; Rhogeessa a ailes noires; Schwarzfligel-Gelbfledermaus; Rogesade alas negras; Central American Yellow Bat; Black-winged Little Yellow Bat; Central American Yellow Bat; Black-winged Little Yellow Bat; Black-winged Little Yellow Bat; R. tumida