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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1470	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina	Rhinophylla alethina		[HMW] Rhinophylla alethina Handley, 1966 , “Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca ], Colombia .” This species is monotypic.														alethina				alethina	alethina			alethina Handley, 1966		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		W Colombia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinophylla alethina	Colombia, Valle, 27 km S. Buenaventura, Raposo River.	Handley	1966	Proc. Biol. Soc. Wash., 79:86.	Distribution: Confined to the Pacific slope of western Colombia and Ecuador.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		W Colombia, Ecuador	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Handley	1966	Proc. Biol. Soc. Washington, 79:86.		W Colombia, W Ecuador.	Colombia, Valle, 27 km S Buenaventura, Raposo River.		HANDLEY	1966	First upper incisor with less than three well-defined lobes, leaving a gap between upper incisors and canine. Margin of uropatagium with a conspicuous fringe of hair. Size relatively large (forearm length, 33-38 mm; condylobasal length, 16-19 mm).	Distribution: Confined to the Pacific slope of western Colombia and Ecuador.	No subspecies.		84	species	R. alethina	HANDLEY	1966	Rhinophylla	genus	Rhinophylla alethina				First upper incisor with less than three well-defined lobes, leaving a gap between upper incisors and canine. Margin of uropatagium with a conspicuous fringe of hair. Size relatively large (forearm length, 33-38 mm; condylobasal length, 16-19 mm).	No subspecies.		2. R. alethina HANDLEY 1966.	2	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Carolliinae		Rhinophylla alethina	Rhinophylla		alethina	Handley		1966		Proc. Biol. Soc. Wash.	79		86		Hairy Little Fruit Bat	Colombia, Valle, 27 km S Buenaventura, Raposo River.	W Colombia, W Ecuador.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).			03A687BCFF89FF891389F8CFF5A7F36B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	539	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF86FF891640F335FCF3F574.xml	Rhinophylla alethina	Phyllostomidae	Rhinophylla	alethina	Handley	1966	Rhinophylle de Handley @fr | Wollhaarige Kleinfruchtfledermaus @de | Rhinofilo de Handley @es	Rhinophylla alethina Handley, 1966 , “Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca ], Colombia .” This species is monotypic.	Pacific slope of Colombia (Choco S to Narino departments) and Ecuador ( Esmeraldas S to Chimborazo provinces).	Head-body 47-60 mm (tailless), ear 12-18 mm, hindfoot 8-12 mm, forearm 34-9-37-2 mm; weight 12-16 g. Dorsalfur of the Hairy Little Fruit Batis blackish. Head pelage is darker than dorsum. Underparts are black on chin, fuscous black on chest, and brownish on abdomen. Dorsal hairs are tricolored. Body fur is woollier in texture than in the Dwarf Little Fruit Bat ( R. pumilio ) and Fischer’s Little Fruit Bat ( R. fischerae ). Tragus is small and blackish. Noseleat is simple and blackish, with base of horseshoe fused to upperlip. Chin has triangular central protuberance bordered by two longitudinalfleshy pads, converging ventrally, and small, circular median ventral protuberance. Wing membranes and uropatagium are blackish. Uropatagium is narrow, ¢. 5 mm from midline. Proximal two-thirds of forearm is hairy. Uropatagium is hairy and fringed. Calcar is smaller than foot. Tail is absent. Rostrum is relatively short, ¢.50% the length of braincase. Zygomatic arches are absent. Sagittal crest is low. Palate is relatively broad and short. Mandible has small angular process. Condyle is level with tooth row or slightly below. Dental formula for all species of Rhinophyllais 12/2, C1/1,P 2/2, M 3/3 ( x2 ) = 32. I' is much larger than I?, P° is small, M? is peg-like, [, is large and trilobed, and I,is small and unicuspid.	Moist tropical forests from sea level to elevations of ¢. 1700 m (most commonly 200-600 m).	The Hairy Little Fruit Bat is a frugivore that eats fruits and infructescences of Ficus ( Moraceae ), Philodendron ( Araceae ), and Piper ( Piperaceae ).	Two reproductive periods of the Hairy Little Fruit Bat have been suggested in February-May and July-November, with birth peaks in October. One young is born per pregnancy.	No information.	No information.	Classified as Near Threatened on The IUCN Red List. Although the Hairy Little Fruit Bat is relatively common in some areas, such as in the Cotacachi-Cayapas Ecological Reserve ( Ecuador ), it is otherwise known from relatively few specimens. Moreover,its habitat, Chocoan forests, suffers from intense deforestation, and future scenarios of climate change predict that it will lose mostits preferred habitat in the next four decades.	Albuja (1999) | Albuja & Mena (1999) | Baud (1982) | Handley (19664) | Iturralde-Pélit et al. (2017) | McLellan & Koopman (2008) | Zapata-Mesa et al. (2017)	https://zenodo.org/record/6458841/files/figure.png	116. Hairy Little Fruit Bat Rhinophylla alethina French: Rhinophylle de Handley / German: Wollhaarige Kleinfruchtfledermaus / Spanish: Rhinofilo de Handley Taxonomy. Rhinophylla alethina Handley, 1966 , “Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca ], Colombia .” This species is monotypic. Distribution. Pacific slope of Colombia (Choco S to Narino departments) and Ecuador ( Esmeraldas S to Chimborazo provinces). Descriptive notes. Head-body 47-60 mm (tailless), ear 12-18 mm, hindfoot 8-12 mm, forearm 34-9-37-2 mm; weight 12-16 g. Dorsalfur of the Hairy Little Fruit Batis blackish. Head pelage is darker than dorsum. Underparts are black on chin, fuscous black on chest, and brownish on abdomen. Dorsal hairs are tricolored. Body fur is woollier in texture than in the Dwarf Little Fruit Bat ( R. pumilio ) and Fischer’s Little Fruit Bat ( R. fischerae ). Tragus is small and blackish. Noseleat is simple and blackish, with base of horseshoe fused to upperlip. Chin has triangular central protuberance bordered by two longitudinalfleshy pads, converging ventrally, and small, circular median ventral protuberance. Wing membranes and uropatagium are blackish. Uropatagium is narrow, ¢. 5 mm from midline. Proximal two-thirds of forearm is hairy. Uropatagium is hairy and fringed. Calcar is smaller than foot. Tail is absent. Rostrum is relatively short, ¢.50% the length of braincase. Zygomatic arches are absent. Sagittal crest is low. Palate is relatively broad and short. Mandible has small angular process. Condyle is level with tooth row or slightly below. Dental formula for all species of Rhinophyllais 12/2, C1/1,P 2/2, M 3/3 ( x2 ) = 32. I' is much larger than I?, P° is small, M? is peg-like, [, is large and trilobed, and I,is small and unicuspid. Habitat. Moist tropical forests from sea level to elevations of ¢. 1700 m (most commonly 200-600 m). Food and Feeding. The Hairy Little Fruit Bat is a frugivore that eats fruits and infructescences of Ficus ( Moraceae ), Philodendron ( Araceae ), and Piper ( Piperaceae ). Breeding. Two reproductive periods of the Hairy Little Fruit Bat have been suggested in February-May and July-November, with birth peaks in October. One young is born per pregnancy. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Near Threatened on The IUCN Red List. Although the Hairy Little Fruit Bat is relatively common in some areas, such as in the Cotacachi-Cayapas Ecological Reserve ( Ecuador ), it is otherwise known from relatively few specimens. Moreover,its habitat, Chocoan forests, suffers from intense deforestation, and future scenarios of climate change predict that it will lose mostits preferred habitat in the next four decades. Bibliography. Albuja (1999), Albuja & Mena (1999), Baud (1982), Handley (19664), Iturralde-Pélit et al. (2017), McLellan & Koopman (2008), Zapata-Mesa et al. (2017).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Rhinophylla alethina	Rhinophylla		alethina	Handley	1966	0	Proc. Biol. Soc. Wash.	80:26:00	Hairy Little Fruit Bat	None.	Colombia, Valle, 27 km S Buenaventura, Raposo River.	W Colombia, W Ecuador.	Not listed.	Near Threatened		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinophylla alethina	23	Hairy Little Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	RHINOPHYLLINAE	NA	Rhinophylla	NA	alethina	Handley	1966	0	Rhinophylla_alethina	Handley, C. O., Jr. (1966). Descriptions of New Bats (Choeroniscus and Rhinophylla) from Colombia. Proceedings of the Biological Society of Washington, 79, 86.	https://www.biodiversitylibrary.org/page/34562753#page/442/mode/1up	USNM 324988		"RÃ­o Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca], Colombia."			alethina Handley, 1966	NA	NA	Colombia|Ecuador	South America	Neotropic	NT	0	0	0	Rhinophylla_alethina	0	sciname match	Rhinophylla_alethina	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19591	Rhinophylla alethina	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Rhinophylla	alethina	Handley, 1966		20000000	Rhinophylla alethina	Near Threatened	A2c	2018	2017-11-08 00:00:00 UTC	3.1	English	This bat is listed as Near Threatened because this species could be in significant decline (but probably at a rate of less than 30% over three generations; 15 years in Pacifici et al. 2013) because of widespread habitat loss through much of its range - especially Chocoan forests, thus making the species close to qualifying for Vulnerable (A2c). The species should be monitored as it may qualify for a higher threat category given more information.	This bat is found in Chocoan forests; it is strongly associated with moist areas and tropical evergreen forests, prefers unaltered habitats (Albuja 1999). In northwestern Ecuador, this species was found in primary forest, secondary forest, and banana plantations (Carrera et al. 2010). Like other species in the genus, it is expected to be primarily frugivorous, feeding on fruits of understory shrubs, also eats insects (Handley 1976). ; In Ecuador, it is reported as showing bimodal reproduction (Albuja 1999). Limited data suggests that reproduction is either extended or asynchronous ;(Wilson 1979).	In general, Chocoan forest habitats are severely threatened by extensive logging and land use changes (agricultural).	It is not known from many specimens, thus population status and trend are unknown. Seems to be locally abundant in appropriate areas, for example, in Reserva Ecologica Cotacachi Cayapas it is one of the most abundant bats (Albuja 1999).	Unknown	Rhinophylla alethina is endemic to the ChocÃ³ region of western Colombia and Ecuador (Handley 1966, Albuja 1999, Simmons 2005, McLellan and Koopman 2008). It occurs between 0-1,000 m on the Pacific flanks of the Andes and adjacent lowlands. In Ecuador it has been reported up to 1,700 m (Burneo pers. comm.).		Terrestrial	The species is found in some protected areas, including National Parks.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Rhinophylla		alethina	Handley	1966	0	Proc. Biol. Soc. Wash.	80:26:00	Hairy Little Fruit Bat	None.	Colombia, Valle, 27 km S Buenaventura, Raposo River.	W Colombia, W Ecuador.	Not listed.	Near Threatened		Rhinophylla alethina	1004992	23	Hairy Little Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	RHINOPHYLLINAE	NA	Rhinophylla	NA	alethina	Handley	1966	0	Rhinophylla_alethina	Handley, C. O., Jr. (1966). Descriptions of New Bats (Choeroniscus and Rhinophylla) from Colombia. Proceedings of the Biological Society of Washington, 79, 86.	https://www.biodiversitylibrary.org/page/34562753#page/442/mode/1up	USNM 324988		"RÃ­o Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca], Colombia."			alethina Handley, 1966	NA	NA				Colombia|Ecuador	South America	Neotropic	NT	0	0	0	Rhinophylla_alethina	0	sciname match	Rhinophylla_alethina	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinophylla_alethina	1004992	23	Hairy Little Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Rhinophyllinae	NA	Rhinophylla	NA	alethina	Handley	0	Rhinophylla alethina	Handley, C.O., Jr. 1966-05-23. Descriptions of new bats (_Choeroniscus_ and _Rhinophylla_) from Colombia. Proceedings of the Biological Society of Washington 79:83-88.	https://www.biodiversitylibrary.org/page/34562756	USNM:MAMM:324988	holotype	http://n2t.net/ark:/65665/38256280b-fa91-4081-b556-8e50442d6789	"RÃ­o Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca], Colombia."			NA	NA				Colombia|Ecuador	South America	Neotropic	NT	0	0	0	Rhinophylla_alethina	0	sciname match	Rhinophylla_alethina	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Rhinophylla		alethina	Handley	1966	0	Proc. Biol. Soc. Wash.	80:26:00	Hairy Little Fruit Bat	None.	Colombia, Valle, 27 km S Buenaventura, Raposo River.	W Colombia, W Ecuador.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19591/21998419/' target='_blank'>Near Threatened</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinophylla alethina; Rhinophylla alethina; Rhinophylla alethina; Rhinophylla alethina; Rhinophylla alethina; Rhinophylla alethina; alethina; Rhinophylle de Handley; Wollhaarige Kleinfruchtfledermaus; Rhinofilo de Handley; Hairy Little Fruit Bat; Hairy Little Fruit Bat; Hairy Little Fruit Bat; R. alethina
