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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1468	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Rhinolophus ziama	Rhinolophus ziama	Rhinolophus ziama	Rhinolophus ziama	Rhinolophus ziama	Rhinolophus ziama	Rhinolophus ziama	Rhinolophus ziama	Rhinolophus ziama	Rhinolophus ziama		[MSW3] maclaudi species group. See Fahr et al. for a key to species of the maclaudi species group.; [HMW] Rhinolophus ziama Fahr et al, 2002 , “ western edge of Sérédou near park station, border of‘Réserve de la Biosphère du Massifdu Ziama’, Guinée Forestière , Guinea . ” Rhinolophus ziama is in the maclaudi species group and is probably most closely related to R maclaudi . Monotypic.; [batnames2022]  maclaudi species group. See Fahr et al. for a key to species of the maclaudi species group.; [IUCN] A split from the Rhinolophus maclaudi complex. Other similar species include R. ruwenzorii and R. hilli .; [batnames2023]  maclaudi species group. See Fahr et al. for a key to species of the maclaudi species group.; [batnames2025_1.7] maclaudispecies group. See Fahr et al. for a key to species of the maclaudi species group.														ziama	A split from the Rhinolophus maclaudi complex. Other similar species include R. ruwenzorii and R. hilli .			ziama	ziama			ziama Fahr, Vierhaus, Hutterer, & Kock, 2002						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus ziama	Rhinolophus		ziama	Fahr, Vierhaus, Hutterer, and Kock		2002		Myotis	40		109		Ziama Horseshoe Bat	Guinea, Guinée Forestière, Réserve de la Biosphère du Massif du Ziama, western edge of Sérédou near park station.	SE Guinea, NW Liberia.	IUCN 2003– Not evaluated (new species). Fahr et al. (2002) suggested that this species be listed as Data Deficient.		maclaudi species group. See Fahr et al. for a key to species of the maclaudi species group.	0383245F2221977F8ED7FFC8FBC1F320	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinonycteridae_194.pdf.imf	hash://md5/ffba5c272223977c8e73f509ffb1ffd0	298	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFDC8A3DF8B9EEB0FA2CD015.xml	Rhinolophus ziama	Rhinolophidae	Rhinolophus	ziama	Fahr et al.	2002	Ziama Horseshoe Bat @en | Rhinolophe du Ziama @fr | Ziama-Hufeisennase @de | Herradura de Ziama @es | German @en	Rhinolophus ziama Fahr et al, 2002 , “ western edge of Sérédou near park station, border of‘Réserve de la Biosphère du Massifdu Ziama’, Guinée Forestière , Guinea . ” Rhinolophus ziama is in the maclaudi species group and is probably most closely related to R maclaudi . Monotypic.	SE Guinea (Ziama Massif) and NW Liberia ( Wonegizi Mts).	Head-body c. 74- 76 mm , tail 35-37 mm , ear 35-36 mm , hindfoot 14 mm , forearm 60 mm ; weight 20- 5 g . The Ziama Horseshoe Bat is one of the largest horseshoe bats in Africa. Dorsal pelage is buffy brown (hairs are pale buff basally and pale brown at tips); venter is paler dirty buff, with pale brown sheen. There is no known orange morph. Males lack axillary tufts. Ears are very long (39-63% of forearm length), with 11—12 internal folds and very large antitragus. Noseleaf has long and narrow subtriangular lancet, with nearly parallel sides distally and rounded tip; connecting process is largely reduced, low, and concave, leaving large depression between sella and lancet; sella is inclined forward, with nearly parallel sides and enlarged lobes at base, forming heart-shaped, corolla-like cup; nostrils are bordered laterally by raised and nearly straight rims that almost reach anterior margin of horseshoe; and horseshoe is of medium width at 11- 5 mm , almost covers muzzle, and has no lateral leaflets and inconspicuous or absent median emarginadon. Lower lip has one medial groove. Wings and uropatagium are dark gray. Skull is large and slender, with weak zygomatic arches that are dorsally notched (zygomatic width is subequal to mastoid width); nasal swellings are high domed, with roughly heart-shaped chambers in dorsal view; frontal depression is very deep; sagittal crest is moderately developed anteriorly and poorly developed posteriorly; infraorbital bridge is very elongated and slender; and interpterygoid groove is very deep. P2 is small and in tooth row, separating C1 and P4; P3 is small and slightly to completely displaced labially; and P2 and P4 are in contact or separate.	Primary and secondary montane and lowland tropical moist forests, recorded at elevations of c. 600 m .	No information.	No information.	Day roosts of Ziama Horseshoe Bats are likely caves, although no records from roosts are known.	No information.	Classified as Endangered on The IUCN ed List. The Ziama Horseshoe Bat is known from only a few specimens in a restricted distribution. It might be threatened by habitat destruction from logging, mining, and agricultural expansion.	ACR (2018) | Fahr (2008c, 2013i) | ahr et al. (2002) | Weber & Fahr (2007b)	https://zenodo.org/record/3749978/files/figure.png	36 . Ziama Horseshoe Bat Rhinolophus ziama French: Rhinolophe du Ziama / German : Ziama-Hufeisennase / Spanish: Herradura de Ziama Taxonomy. Rhinolophus ziama Fahr et al, 2002 , “ western edge of Sérédou near park station, border of‘Réserve de la Biosphère du Massifdu Ziama’, Guinée Forestière , Guinea . ” Rhinolophus ziama is in the maclaudi species group and is probably most closely related to R maclaudi . Monotypic. Distribution. SE Guinea (Ziama Massif) and NW Liberia ( Wonegizi Mts). Descriptive notes. Head-body c. 74- 76 mm , tail 35-37 mm , ear 35-36 mm , hindfoot 14 mm , forearm 60 mm ; weight 20- 5 g . The Ziama Horseshoe Bat is one of the largest horseshoe bats in Africa. Dorsal pelage is buffy brown (hairs are pale buff basally and pale brown at tips); venter is paler dirty buff, with pale brown sheen. There is no known orange morph. Males lack axillary tufts. Ears are very long (39-63% of forearm length), with 11—12 internal folds and very large antitragus. Noseleaf has long and narrow subtriangular lancet, with nearly parallel sides distally and rounded tip; connecting process is largely reduced, low, and concave, leaving large depression between sella and lancet; sella is inclined forward, with nearly parallel sides and enlarged lobes at base, forming heart-shaped, corolla-like cup; nostrils are bordered laterally by raised and nearly straight rims that almost reach anterior margin of horseshoe; and horseshoe is of medium width at 11- 5 mm , almost covers muzzle, and has no lateral leaflets and inconspicuous or absent median emarginadon. Lower lip has one medial groove. Wings and uropatagium are dark gray. Skull is large and slender, with weak zygomatic arches that are dorsally notched (zygomatic width is subequal to mastoid width); nasal swellings are high domed, with roughly heart-shaped chambers in dorsal view; frontal depression is very deep; sagittal crest is moderately developed anteriorly and poorly developed posteriorly; infraorbital bridge is very elongated and slender; and interpterygoid groove is very deep. P2 is small and in tooth row, separating C1 and P4; P3 is small and slightly to completely displaced labially; and P2 and P4 are in contact or separate. Habitat. Primary and secondary montane and lowland tropical moist forests, recorded at elevations of c. 600 m . Food and Feeding. No information. Breeding. No information. Activity patterns. Day roosts of Ziama Horseshoe Bats are likely caves, although no records from roosts are known. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Endangered on The IUCN ed List. The Ziama Horseshoe Bat is known from only a few specimens in a restricted distribution. It might be threatened by habitat destruction from logging, mining, and agricultural expansion. Bibliography. ACR (2018), Fahr (2008c, 2013i ), ahr et al. (2002), Weber & Fahr (2007b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus ziama	Rhinolophus		ziama	Fahr, Vierhaus, Hutterer & Kock	2002	0	Myotis	41:49:00	Ziama Horseshoe Bat	None.	Guinea, GuinÃ©e ForestiÃ¨re, RÃ©serve de la BiosphÃ¨re du Massif du Ziama, western edge of SÃ©rÃ©dou near park station.	SE Guinea, NW Liberia.	Not listed.	Endangered	 maclaudi species group. See Fahr et al. for a key to species of the maclaudi species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus ziama	23	Ziama Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	ziama	Fahr, Vierhaus, Hutterer, & Kock	2002	0	Rhinolophus_ziama	Fahr, J., Vierhaus, H., Hutterer, R. & Kock, D. (2002). A revision of the Rhinolophus maclaudi species group with the description of a new species from West Africa (Chiroptera: Rhinolophidae). Myotis, 40, 109.	https://www.researchgate.net/profile/Jakob_Fahr/publication/209869220_A_revision_of_the_Rhinolophus_maclaudi_species_group_with_the_description_of_a_new_species_from_West_Africa_Chiroptera_Rhinolophidae/links/0912f50cd9f7351c53000000.pdf	ZFMK 99.934		"western edge of SÃ©rÃ©dou near park station, border of â€˜RÃ©serve de la BiosphÃ¨re du Massif du Ziama', GuinÃ©e ForestiÃ¨re, Guinea."			ziama Fahr, Vierhaus, Hutterer, & Kock, 2002	NA	NA	Guinea|Liberia|Sierra Leone	Africa	Afrotropic	EN	0	0	0	Rhinolophus_ziama	0	sciname match	Rhinolophus_ziama	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	44786	Rhinolophus ziama	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	ziama	Fahr, Vierhaus, Hutterer &; Kock, 2002	A split from the Rhinolophus maclaudi complex. Other similar species include R. ruwenzorii and R. hilli .	20000000	Rhinolophus ziama	Endangered	B2ab(iii)	2020	2019-06-21 00:00:00 UTC	3.1	English	This species is listed as Endangered under criterion B2ab(iii) as it is only known from three locations in Guinea, Liberia, and Sierra Leone where its area of occupancy (AOO) is 150-300 kmÂ². The global population for this species is projected to continue to decline as the quality, area, and extent of its forest and cave habitats continues to be lost and/or degraded.	This species is endemic to West Africa and appears to be restricted to Afromontane-Afroalpine and lowland tropical moist forest (rainforest) of Guinea, Liberia, and Sierra Leone (Fahr 2013, Decher 2010). The Guinea specimens were captured in a secondary forest close to primary forest, while the Liberian specimen was captured in upland "undisturbed High Forest" (Fahr et al. 2002). The Sierra Leone specimen was captured in riparian forest in the transition zone between forest and savanna (Decher et al. 2010). The day roosts of this species are not known, although it is suspected to roost as small colonies in caves, mine shafts and similar habitats based on assumed similarities with Rhinolophus maclaudi (Fahr et al. 2002, Happold 2013).	The species is threatened by past and continued deforestation of its roosts and foraging habitats, largely through logging and mining operations as well as inundation of forest for the generation of hydro-electric power, and conversion of land to agricultural use. It is also considered possible that the species could be threatened by overharvesting for subsistence food in the future (Happold 2013).	This species is known only from five specimens (Fahr et al. 2002, Decher et al. 2010). The global population is thought to be declining due to ongoing loss and degradation of forests and cave/mine habitats.	Decreasing	This West African bat is known from three (3) locations: one each in southeast Guinea, northwest Liberia, and Sierra Leone. Two of the locations are in the Guinea Highlands (ca 600 m asl) of southeast Guinea (Ziama Forest) and north-west Liberia (Wonegizi Mountains) (Fahr et al. 2002). The third location in Sierra Leone is in the upper Seli valley (Decher et al. 2010). Although surrounding areas have been surveyed for the species, such as at Mount Simandou, Mount Nimba, and the Gola Forest, it remains undetected (Fahr et al. 2006, Decher 2015, Denys et al. 2013, Monadjem et al. 2016).		Terrestrial	It is not known if the species is present in any protected areas. There is a need to protect suitable areas of forest habitat for this species, and to initiate appropriate bat conservation awareness programmes among local people. In addition, further studies are needed into the distribution, abundance, natural history, and threats to this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		ziama	Fahr, Vierhaus, Hutterer & Kock	2002	0	Myotis	41:49:00	Ziama Horseshoe Bat	None.	Guinea, GuinÃ©e ForestiÃ¨re, RÃ©serve de la BiosphÃ¨re du Massif du Ziama, western edge of SÃ©rÃ©dou near park station.	SE Guinea, NW Liberia.	Not listed.	Endangered	 maclaudi species group. See Fahr et al. for a key to species of the maclaudi species group.	Rhinolophus ziama	1004760	23	Ziama Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	ziama	Fahr, Vierhaus, Hutterer, & Kock	2002	0	Rhinolophus_ziama	Fahr, J., Vierhaus, H., Hutterer, R. & Kock, D. (2002). A revision of the Rhinolophus maclaudi species group with the description of a new species from West Africa (Chiroptera: Rhinolophidae). Myotis, 40, 109.	https://www.researchgate.net/profile/Jakob_Fahr/publication/209869220_A_revision_of_the_Rhinolophus_maclaudi_species_group_with_the_description_of_a_new_species_from_West_Africa_Chiroptera_Rhinolophidae/links/0912f50cd9f7351c53000000.pdf	ZFMK 99.934		"western edge of SÃ©rÃ©dou near park station, border of â€˜RÃ©serve de la BiosphÃ¨re du Massif du Ziama', GuinÃ©e ForestiÃ¨re, Guinea."			ziama Fahr, Vierhaus, Hutterer, & Kock, 2002	NA	NA				Guinea|Liberia|Sierra Leone	Africa	Afrotropic	EN	0	0	0	Rhinolophus_ziama	0	sciname match	Rhinolophus_ziama	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_ziama	1004760	23	Ziama Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	ziama	Fahr, Vierhaus, Hutterer, & Kock	0	Rhinolophus ziama	Fahr, J., Vierhaus, H., Hutterer, R. and Kock, D. 2002-12. A revision of the _Rhinolophus maclaudi_ species group with the description of a new species from West Africa (Chiroptera: Rhinolophidae). Myotis 40:95-126.	https://www.researchgate.net/profile/Jakob_Fahr/publication/209869220_A_revision_of_the_Rhinolophus_maclaudi_species_group_with_the_description_of_a_new_species_from_West_Africa_Chiroptera_Rhinolophidae/links/0912f50cd9f7351c53000000.pdf	ZFMK:Mamm:1999.934	holotype		"western edge of SÃ©rÃ©dou near park station, border of â€˜RÃ©serve de la BiosphÃ¨re du Massif du Ziama', GuinÃ©e ForestiÃ¨re, Guinea."	8.383333	-9.3	NA	NA				Guinea|Liberia|Sierra Leone	Africa	Afrotropic	EN	0	0	0	Rhinolophus_ziama	0	sciname match	Rhinolophus_ziama	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		ziama	Fahr, Vierhaus, Hutterer & Kock	2002	0	Myotis	41:49:00	Ziama Horseshoe Bat	None.	Guinea, GuinÃ©e ForestiÃ¨re, RÃ©serve de la BiosphÃ¨re du Massif du Ziama, western edge of SÃ©rÃ©dou near park station.	SE Guinea, NW Liberia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/44786/22068674/' target='_blank'>Endangered</a>	maclaudispecies group. See Fahr et al. for a key to species of the maclaudi species group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus ziama; Rhinolophus ziama; Rhinolophus ziama; Rhinolophus ziama; Rhinolophus ziama; Rhinolophus ziama; ziama; Ziama Horseshoe Bat; Rhinolophe du Ziama; Ziama-Hufeisennase; Herradura de Ziama; German; Ziama Horseshoe Bat; Ziama Horseshoe Bat; Ziama Horseshoe Bat; R. ziama
