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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1460	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi	Rhinolophus thomasi		[MSW3] rouxii species group. Does not include septentrionalis, see Csorba et al. (2003). Also see Corbet and Hill (1992) and Hendrichsen et al. (2001b).; [HMW] Rhinolophus thomasi K . Andersen, 1905 , “ Karin Hills, Burma [= Myanmar ]. ” Rhinolophus thomasi is included in the rouxii species group and appears to be sister to R. sinicus septentrionalis , which is likely a distinct species pending revision. Two subspecies are sometimes recognized. Here considered to be monotypic until further studies validate the existence of either subspecies.; [batnames2022]  rouxii species group. Does not include septentrionalis ; see Csorba (2002) and Csorba et al. (2003, 2019). Also see Corbet and Hill (1992) and Hendrichsen et al.(2001). Zhang et al (2009) suggested that thomasi and sinicus may be conspecific (see also Csorba, 2002), but data from Mao et al. (2019) support the distinctiveness of both taxa. Following Csorba et al. (2003, 2019) we do not recognize any subspecies of thomasi .; [IUCN] Wang (2002) assigns Chinese material to the subspecies R. t. latifolius , but we follow Csorba et al . (2003) in relegating latifolius as a synonym of the nominate form.; [batnames2023]  rouxii species group. Does not include septentrionalis ; see Csorba (2002) and Csorba et al. (2003, 2019). Also see Corbet and Hill (1992) and Hendrichsen et al.(2001). Zhang et al (2009) suggested that thomasi and sinicus may be conspecific (see also Csorba, 2002), but data from Mao et al. (2019) support the distinctiveness of both taxa. Following Csorba et al. (2003, 2019) we do not recognize any subspecies of thomasi .; [batnames2025_1.7] rouxii species group. Does not include septentrionalis; see Csorba (2002), Csorba et al. (2003) and Burgin (2019). Also see Corbet and Hill (1992) and Hendrichsen et al.(2001). Zhang et al (2009) suggested that thomasi and sinicus may be conspecific (see also Csorba, 2002), but data from Mao et al. (2019) support the distinctiveness of both taxa. Following Csorba et al. (2003) and Burgin (2019) we do not recognize any subspecies of thomasi.						latifolius, septentrionalis.	thomasi, septentrionalis, latifolius	thomasi, latifolius				thomasi, latifolius		thomasi, latifolius	Wang (2002) assigns Chinese material to the subspecies R. t. latifolius , but we follow Csorba et al . (2003) in relegating latifolius as a synonym of the nominate form.	thomasi	thomasi - latifolius	thomasi, latifolius 	thomasi, latifolius 	thomasi 	thomasi - latifolius 	thomasi Andersen, 1905|latifolius Sanborn, 1939		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Burma, Yunnan, Indochina	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinolophus thomasi	Burma, Karin Hills.	K. Andersen	1905	Proc. Zool. Soc. Lond., 1905:100.	Distribution: Known from Burma, southwestern China, Thailand, and Vietnam.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Thomas's horseshoe bat	Burma, Yunnan – Indochina	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	K. Andersen	1905	Proc. Zool. Soc. Lond., 1905:100.		Burma, Vietnam, Thailand, Yunnan (China).	Burma, Karin Hills.		ANDERSEN	1905	Lancet tip extremely short (very has tate). Metacarpals relatively short. Anterior up per premolar greatly reduced, though in tooth row. Size medium to fairly large (forearm length, 42-55 mm).	Distribution: Known from Burma, southwestern China, Thailand, and Vietnam.	Three subspecies are recognized:	R. t. thomasi (Burma, Thailand), R. t. septentrionalis (Yun nan in China), R. t. latifolius (Vietnam).	54	species	R. thomasi	ANDERSEN	1905	Rhinolophus	genus	Rhinolophus thomasi				Lancet tip extremely short (very has tate). Metacarpals relatively short. Anterior up per premolar greatly reduced, though in tooth row. Size medium to fairly large (forearm length, 42-55 mm).	Three subspecies are recognized:		15. R. thomasi ANDERSEN 1905 [ferrumequinum group].	15	_R. t. latifolius_ Sanborn, 1939; _R. t. thomasi_ Andersen, 1905			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus thomasi	Rhinolophus		thomasi	K. Andersen		1905		Proc. Zool. Soc. Lond.	1905		100		Thomas's Horseshoe Bat	Burma, Karin Hills.	Burma, Vietnam, Thailand, Laos.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	latifolius Sanborn, 1939.	rouxii species group. Does not include septentrionalis, see Csorba et al. (2003). Also see Corbet and Hill (1992) and Hendrichsen et al. (2001b).	885887A2FFFE8A18F8B4FCA2F30BD6BB	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	326	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFE08A06FF07F506F2B4DAA9.xml	Rhinolophus thomasi	Rhinolophidae	Rhinolophus	thomasi	K. Andersen	1905	Rhinolophe de Thomas @fr | Thomas-Hufeisennase @de | Herradura de Thomas @es	Rhinolophus thomasi K . Andersen, 1905 , “ Karin Hills, Burma [= Myanmar ]. ” Rhinolophus thomasi is included in the rouxii species group and appears to be sister to R. sinicus septentrionalis , which is likely a distinct species pending revision. Two subspecies are sometimes recognized. Here considered to be monotypic until further studies validate the existence of either subspecies.	S China , E Myanmar , Thailand , Laos , and Vietnam .	Head—body 48—50 mm , tail 18—28 mm , ear 16—24 mm , hindfoot 8—10 mm , forearm 40-48 mm ; weight 7—11 g . Dorsal pelage is rich brown with a russet tinge (hairs light yellow basally with medium brown tips); ventral pelage is slightly paler. Ears are small. Noseleaf has a short lancet, with an almost rudimentary tip in some cases; connecting process is rounded, as in the Indian Rufous Horseshoe Bat ( R.rouxii ) ; sella is virtually parallel-sided with a broadly rounded-off tip; horseshoe is moderately wide (7-2-8- 9 mm ) but does not cover the muzzle completely , and welldeveloped lateral leaflets are usually present. Lower lip has three mental grooves. Baculum is lightly built with double curvature, making an elongated S-shape; basal cone is dorso-ventrally flattened and smaller than in the Indian Rufous Horseshoe Bat; basal emargination is deeper ventrally than dorsally; shaft is laterally compressed, and its proximal ventral constriction is adjacent to the basal cone. Skull is relatively robust (zygomatic width is much greater than mastoid width); anterior median swellings are small; posterior swellings are more developed than the anterior ones; rostral profile is clearly sloped rearward; sagittal crest is moderately high; frontal depression is shallow to moderately deep; supraorbital crests are obliterated or well demarcated. C1 is moderately long; P2 is small and partially to completely extruded from the tooth row; C1 and P4 are touching or separated, depending on how extruded P2 is; P3 is small and completely extruded from the tooth row; P2 and P are touching or nearly so.	Often associated with limestone areas, and has been recorded in dry forests and disturbed areas. Reported at elevations of 400-1100 m .	No information.	Pregnant females have been recorded in late March and April, in Vietnam .	Thomas’s Horseshoe Bat roosts in caves. Call shape is FM/CF/FM with peak F reported at 76-77 kHz in males and 83-85-5 kHz in females, in Laos .	Thomas’s Horseshoe Bats roost in colonies.	Classified as Least Concern on The IUCNed List. Thomas’s Horseshoe Bat is widespread and does not seem to be facing any major threats, although it may be locally threatened by roost disturbance. Cave destruction in some regions has negatively affected this species, causing local population declines.	Bates, Bumrungsri & Csorba (2008f) | Bates, Thi Mar-Mar et al. (2004) | Csorba et al. (2003) | ran ­ cis (2008a) | Kruskop (2013a) | Smith & XieYan (2008) | Stoffberg et al. (2010) | Zhang Lin et al. (2018)	https://zenodo.org/record/3750098/files/figure.png	94 . Thomas’s Horseshoe Bat Rhinolophus thomasi French: Rhinolophe deThomas /German: Thomas-Hufeisennase / Spanish: Herradura de Thomas Taxonomy. Rhinolophus thomasi K . Andersen, 1905 , “ Karin Hills, Burma [= Myanmar ]. ” Rhinolophus thomasi is included in the rouxii species group and appears to be sister to R. sinicus septentrionalis , which is likely a distinct species pending revision. Two subspecies are sometimes recognized. Here considered to be monotypic until further studies validate the existence of either subspecies. Distribution. S China , E Myanmar , Thailand , Laos , and Vietnam . Descriptive notes. Head—body 48—50 mm , tail 18—28 mm , ear 16—24 mm , hindfoot 8—10 mm , forearm 40-48 mm ; weight 7—11 g . Dorsal pelage is rich brown with a russet tinge (hairs light yellow basally with medium brown tips); ventral pelage is slightly paler. Ears are small. Noseleaf has a short lancet, with an almost rudimentary tip in some cases; connecting process is rounded, as in the Indian Rufous Horseshoe Bat ( R.rouxii ) ; sella is virtually parallel-sided with a broadly rounded-off tip; horseshoe is moderately wide (7-2-8- 9 mm ) but does not cover the muzzle completely , and welldeveloped lateral leaflets are usually present. Lower lip has three mental grooves. Baculum is lightly built with double curvature, making an elongated S-shape; basal cone is dorso-ventrally flattened and smaller than in the Indian Rufous Horseshoe Bat; basal emargination is deeper ventrally than dorsally; shaft is laterally compressed, and its proximal ventral constriction is adjacent to the basal cone. Skull is relatively robust (zygomatic width is much greater than mastoid width); anterior median swellings are small; posterior swellings are more developed than the anterior ones; rostral profile is clearly sloped rearward; sagittal crest is moderately high; frontal depression is shallow to moderately deep; supraorbital crests are obliterated or well demarcated. C1 is moderately long; P2 is small and partially to completely extruded from the tooth row; C1 and P4 are touching or separated, depending on how extruded P2 is; P3 is small and completely extruded from the tooth row; P2 and P are touching or nearly so. Habitat. Often associated with limestone areas, and has been recorded in dry forests and disturbed areas. Reported at elevations of 400-1100 m . Food and Feeding. No information. Breeding. Pregnant females have been recorded in late March and April, in Vietnam . Activity patterns. Thomas’s Horseshoe Bat roosts in caves. Call shape is FM/CF/FM with peak F reported at 76-77 kHz in males and 83-85-5 kHz in females, in Laos . Movements, Home range and Social organization. Thomas’s Horseshoe Bats roost in colonies. Status and Conservation. Classified as Least Concern on The IUCNed List. Thomas’s Horseshoe Bat is widespread and does not seem to be facing any major threats, although it may be locally threatened by roost disturbance. Cave destruction in some regions has negatively affected this species, causing local population declines. Bibliography. Bates, Bumrungsri & Csorba (2008f), Bates, Thi Mar-Mar eta/. (2004), Csorba eta/. (2003), ran ­ cis (2008a), Kruskop (2013a), Smith & XieYan (2008), Stoffberg eta/. (2010), Zhang Lin et al. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus thomasi	Rhinolophus		thomasi	K. Andersen	1905	0	Proc. Zool. Soc. Lond.	1905, II: 100, pl. III, fig. 10	Thomas's Horseshoe Bat	<b> latifolius </b>Sanborn, 1939.	Burma, Karin Hills	S China, E Myanmar, Laos, Vietnam, Thailand	Not listed.	Least Concern	 rouxii species group. Does not include septentrionalis ; see Csorba (2002) and Csorba et al. (2003, 2019). Also see Corbet and Hill (1992) and Hendrichsen et al.(2001). Zhang et al (2009) suggested that thomasi and sinicus may be conspecific (see also Csorba, 2002), but data from Mao et al. (2019) support the distinctiveness of both taxa. Following Csorba et al. (2003, 2019) we do not recognize any subspecies of thomasi .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus thomasi	23	Thomas's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	thomasi	K. Andersen	1905	0	Rhinolophus_thomasi	Andersen, K. (1905). On Some Bats of the Genus Rhinolophus, with Remarks on Their Natural Affinities, and Descriptions of Twenty-six New Forms. Proceedings of the Zoological Society of London, 1905(2), 100.	https://www.biodiversitylibrary.org/item/99487#page/156/mode/1up	BM 1890.4.7.10		"Karin Hills, Burma [= Myanmar]."			thomasi K. Andersen, 1905|latifolius Sanborn, 1939	NA	NA	China|Myanmar|Thailand|Laos|Vietnam	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_thomasi	0	sciname match	Rhinolophus_thomasi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19573	Rhinolophus thomasi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	thomasi	K. Andersen, 1905	Wang (2002) assigns Chinese material to the subspecies R. t. latifolius , but we follow Csorba et al . (2003) in relegating latifolius as a synonym of the nominate form.	20000000	Rhinolophus thomasi	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Confirmed as Least Concern as recent records show that this species is even more widespread than previously thought, it is locally uncommon to common, and it is unlikely to be declining at nearly the rate required to qualify for listing in a more threatened category.	This species is recorded from different habitats and found in caves from 400 up to 1,100 m asl. It can also occur in degraded habitats. Nothing is known from China, but this species is known to roost in caves with other species of Rhinolophus in other areas.	There are no major threats, although the species appears to be dependent on caves and their destruction in some parts of the range may cause localised population declines (Bates et al. 2008).	There is no information available on the population and abundance of this species. However, it is known to be locally uncommon to common, but never observed in large colonies.	Unknown	This species occurs in south west China, extending down the Indochina Peninsula in Myanmar, Viet Nam, Thailand, and Lao PDR.		Terrestrial	It is known from a number of protected and unprotected areas throughout its range. The species is listed as Vulnerable in the current Red Data Book of Vietnam (MOST and VAST 2007). However, conservation of the species is still limited nationwide.	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		thomasi	K. Andersen	1905	0	Proc. Zool. Soc. Lond.	1905, II: 100, pl. III, fig. 10	Thomas's Horseshoe Bat	<b> latifolius </b>Sanborn, 1939.	Burma, Karin Hills	S China, E Myanmar, Laos, Vietnam, Thailand	Not listed.	Least Concern	 rouxii species group. Does not include septentrionalis ; see Csorba (2002) and Csorba et al. (2003, 2019). Also see Corbet and Hill (1992) and Hendrichsen et al.(2001). Zhang et al (2009) suggested that thomasi and sinicus may be conspecific (see also Csorba, 2002), but data from Mao et al. (2019) support the distinctiveness of both taxa. Following Csorba et al. (2003, 2019) we do not recognize any subspecies of thomasi .	Rhinolophus thomasi	1004754	23	Thomas's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	thomasi	K. Andersen	1905	0	Rhinolophus_thomasi	Andersen, K. (1905). On Some Bats of the Genus Rhinolophus, with Remarks on Their Natural Affinities, and Descriptions of Twenty-six New Forms. Proceedings of the Zoological Society of London, 1905(2), 100.	https://www.biodiversitylibrary.org/item/99487#page/156/mode/1up	BM 1890.4.7.10		"Karin Hills, Burma [= Myanmar]."			thomasi K. Andersen, 1905|latifolius Sanborn, 1939	NA	NA				China|Myanmar|Thailand|Laos|Vietnam	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_thomasi	0	sciname match	Rhinolophus_thomasi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_thomasi	1004754	23	Thomas's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	thomasi	Andersen	0	Rhinolophus thomasi	Andersen, K.C. 1905-10-07. On some bats of the genus _Rhinolophus_, with remarks on their mutual affinities, and descriptions of twenty-six new forms. Proceedings of the Zoological Society of London 1905-II(1):75-145.	https://www.biodiversitylibrary.org/page/31207561	BMNH:Mamm:1890.4.7.10	holotype	https://data.nhm.ac.uk/object/21052960-a0b6-456a-80c9-494eae6fe439	"Karin Hills, Burma [= Myanmar]."			NA	NA				China|Myanmar|Thailand|Laos|Vietnam	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_thomasi	0	sciname match	Rhinolophus_thomasi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		thomasi	K. Andersen	1905	0	Proc. Zool. Soc. Lond.	1905, II: 100, pl. III, fig. 10	Thomas's Horseshoe Bat	Yes.	Burma, Karin Hills	S China, E Myanmar, Laos, Vietnam, Thailand	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19573/21990671/' target='_blank'>Least Concern</a>	rouxii species group. Does not include septentrionalis; see Csorba (2002), Csorba et al. (2003) and Burgin (2019). Also see Corbet and Hill (1992) and Hendrichsen et al.(2001). Zhang et al (2009) suggested that thomasi and sinicus may be conspecific (see also Csorba, 2002), but data from Mao et al. (2019) support the distinctiveness of both taxa. Following Csorba et al. (2003) and Burgin (2019) we do not recognize any subspecies of thomasi.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus thomasi; Rhinolophus thomasi; Rhinolophus thomasi; Rhinolophus thomasi; Rhinolophus thomasi; Rhinolophus thomasi; thomasi; latifolius; latifolius; thomasi; latifolius; Rhinolophe de Thomas; Thomas-Hufeisennase; Herradura de Thomas; Thomas's Horseshoe Bat; Thomas's Horseshoe Bat; Thomas's Horseshoe Bat; R. thomasi