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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1458	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Rhinolophus euryotis [synonym of]	Rhinolophus euryotis tatar	Rhinolophus euryotis tatar	Rhinolophus tatar	Rhinolophus tatar	Rhinolophus tatar	Rhinolophus tatar	Rhinolophus tatar	Rhinolophus tatar	Rhinolophus tatar	Rhinolophus tatar	Rhinolophus tatar		[HMW] Rhinolophus tatar Bergmans & Rozendaal, 1982 , “ 5 m above Moinakom River ( 00°41’N , 124°3’E ), Dumoga Nature Reserve , North Sulawesi , Indonesia , altitude 525 m .” Rhinolophus tatar is included in the euryotis species group and is sister to the clade that includes. euryotis and the new central Sulawesi species. Rhinolophus tatar was previously included in R. euryotis but was recently elevated to species level by L. E. Patrick and coworkers in 2013, based on genetic and morphometric data. The islands in which R. euryotis occurs between Sulawesi and New Guinea have not been tested phylogenetically or morphometrically, and some or all of these populations could potentially refer to R. tatar . Monotypic.; [batnames2022] Formerly included in euryotis, but see Patrick et al. 2013.; [MDD2022] split from R. euryotis; [IUCN] <span lang="EN-US">This species was considered to be conspecific with R. euryotis <span lang="EN-US">(</span><span lang="EN-US">Hill 1992<span lang="EN-US">)</span><span lang="EN-US">. However, genetic work by Patrick et al . <span lang="EN-US">(</span><span lang="EN-US">2013<span lang="EN-US">) ;</span><span lang="EN-US">demonstrated that specimens from Sulawesi were genetically distinct from those collected on Papua New Guinea and deserving of distinct species status.  </span>; [batnames2023] Formerly included in euryotis, but see Patrick et al. 2013.; [MDD2023] split from R. euryotis; [MDD2025_2.0] split from R. euryotis; [batnames2025_1.7] Formerly included in euryotis, but see Patrick et al. 2013.; [MDD2025_2.2] split from R. euryotis														tatar	<span lang="EN-US">This species was considered to be conspecific with R. euryotis <span lang="EN-US">(</span><span lang="EN-US">Hill 1992<span lang="EN-US">)</span><span lang="EN-US">. However, genetic work by Patrick et al . <span lang="EN-US">(</span><span lang="EN-US">2013<span lang="EN-US">) ;</span><span lang="EN-US">demonstrated that specimens from Sulawesi were genetically distinct from those collected on Papua New Guinea and deserving of distinct species status.  </span>			tatar	tatar			tatar Bergmans & F. G. Rozendaal, 1982						N/A																																								NA																											885887A2FFFB8A1DFF7CF5D2F77FDCC8	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	318	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFE88A0EF8B7FD62F247D5CA.xml	Rhinolophus tatar	Rhinolophidae	Rhinolophus	tatar	Bergmans & Rozendaal	1982	Sulawesi Broad-eared Horseshoe Bat @en | Rhinolophe d'Archbold @fr | Tate-Archbold-Hufeisennase @de | Herradura de Archbold @es | Tatar Horseshoe Bat @en	Rhinolophus tatar Bergmans & Rozendaal, 1982 , “ 5 m above Moinakom River ( 00°41’N , 124°3’E ), Dumoga Nature Reserve , North Sulawesi , Indonesia , altitude 525 m .” Rhinolophus tatar is included in the euryotis species group and is sister to the clade that includes. euryotis and the new central Sulawesi species. Rhinolophus tatar was previously included in R. euryotis but was recently elevated to species level by L. E. Patrick and coworkers in 2013, based on genetic and morphometric data. The islands in which R. euryotis occurs between Sulawesi and New Guinea have not been tested phylogenetically or morphometrically, and some or all of these populations could potentially refer to R. tatar . Monotypic.	Sulawesi .	Head-body 51-2-61- 6 mm , tail 16-2-21- 6 mm , ear 17-2-22- 3 mm , hindfoot 9-2-12- 4 mm , forearm 47-3-51- 8 mm ; weight 9-13 g . Dorsal pelage is dark brown (hairs with whitish base and dark brown shaft), whereas ventral pelage is a litde lighter. Ears are moderately long. Noseleaf has more or less straight-sided and densely haired lancet; connecting process is semicircular and is hairy; sella is wide but relatively short and almost parallel-sided with convex outlined tip; horseshoe has longitudinal groove with raised edges extending from median edge to intemarial region, is dark in color with exception of whitish stripe on median longitudinal groove, is wide (9—9- 6 mm ), covering muzzle, and has very shallow median emargination. Lower lip has three mental grooves. Skull is large but slender in build (zygomatic breadth is subequal to mastoid breadth); anterior median swellings are prominent and bulbous; posterior swellings are reduced; frontal depression extends anteriorly to medium swellings and is elongated and moderately deep; supraorbital crests are conspicuous but not very sharp; sagittal crest is moderately developed. C1 is long and strong; P2 is medium-sized and within tooth row; P3 is very small and completely extruded from tooth row, or sometimes missing entirely; P and P4 are usually very close to one another or touching. Dental formula is the usual of 32 teeth for the genus, or only 30 teeth when a lower premolar is missing.	Recorded from slightly disturbed lowland rainforest to lower montane forests.	No information.	A pregnant female with one embryo was collected in November.	Sulawesi Broad-eared Horseshoe Bats probably roost in trees, or possibly in caves.	No information.	Classified as Least Concern on The IUCN ed List. The Sulawesi Broad-eared Horseshoe Bat is relatively widespread but is likely to be threatened by roost disturbance and habitat destruction and alteration.	Bergmans & Rozendaal (1982) | Patrick & Ruedas (2017d) | Patrick et al. (2013) | Wiantoro et at (2017)	https://zenodo.org/record/3750068/files/figure.png	79 . Sulawesi Broad-eared Horseshoe Bat Rhinolophus tatar French: Rhinolophe d'Archbold I German: Tate-Archbold-Hufeisennase / Spanish: Herradura de Archbold Other common names: Tatar Horseshoe Bat Taxonomy. Rhinolophus tatar Bergmans & Rozendaal, 1982 , “ 5 m above Moinakom River ( 00°41’N , 124°3’E ), Dumoga Nature Reserve , North Sulawesi , Indonesia , altitude 525 m .” Rhinolophus tatar is included in the euryotis species group and is sister to the clade that includes. euryotis and the new central Sulawesi species. Rhinolophus tatar was previously included in R. euryotis but was recently elevated to species level by L. E. Patrick and coworkers in 2013, based on genetic and morphometric data. The islands in which R. euryotis occurs between Sulawesi and New Guinea have not been tested phylogenetically or morphometrically, and some or all of these populations could potentially refer to R. tatar . Monotypic. Distribution. Sulawesi . Descriptive notes. Head-body 51-2-61- 6 mm , tail 16-2-21- 6 mm , ear 17-2-22- 3 mm , hindfoot 9-2-12- 4 mm , forearm 47-3-51- 8 mm ; weight 9-13 g . Dorsal pelage is dark brown (hairs with whitish base and dark brown shaft), whereas ventral pelage is a litde lighter. Ears are moderately long. Noseleaf has more or less straight-sided and densely haired lancet; connecting process is semicircular and is hairy; sella is wide but relatively short and almost parallel-sided with convex outlined tip; horseshoe has longitudinal groove with raised edges extending from median edge to intemarial region, is dark in color with exception of whitish stripe on median longitudinal groove, is wide (9—9- 6 mm ), covering muzzle, and has very shallow median emargination. Lower lip has three mental grooves. Skull is large but slender in build (zygomatic breadth is subequal to mastoid breadth); anterior median swellings are prominent and bulbous; posterior swellings are reduced; frontal depression extends anteriorly to medium swellings and is elongated and moderately deep; supraorbital crests are conspicuous but not very sharp; sagittal crest is moderately developed. C1 is long and strong; P2 is medium-sized and within tooth row; P3 is very small and completely extruded from tooth row, or sometimes missing entirely; P and P4 are usually very close to one another or touching. Dental formula is the usual of 32 teeth for the genus, or only 30 teeth when a lower premolar is missing. Habitat. Recorded from slightly disturbed lowland rainforest to lower montane forests. Food and Feeding. No information. Breeding. A pregnant female with one embryo was collected in November. Activity patterns. Sulawesi Broad-eared Horseshoe Bats probably roost in trees, or possibly in caves. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN ed List. The Sulawesi Broad-eared Horseshoe Bat is relatively widespread but is likely to be threatened by roost disturbance and habitat destruction and alteration. Bibliography. Bergmans & Rozendaal (1982), Patrick & Ruedas (2017d), Patrick eta/. (2013), Wiantoro et at (2017).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus tatar	Rhinolophus		tatar	Bergmans and Rozendaal	1982	0	Bijdr. Dierk.	54:50:00	Tatar Horseshoe Bat	None.		Sulawesi	Not listed.	Least Concern	Formerly included in euryotis, but see Patrick et al. 2013.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus tatar	23	Sulawesi Broad-eared Horseshoe Bat	Tatar Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	tatar	Bergmans & Rozendaal	1982	0	Rhinolophus_tatar	Bergmans, W., & Rozendaal, F. G. (1982). Notes on Rhinolophus Lacépède, 1799 from Sulawesi, Indonesia, with the description of a new species (Mammalia, Microchiroptera). Bijdragen tot de Dierkunde, 52(2), 170.	https://brill.com/view/journals/btd/52/2/article-p169_7.xml	ZMA 21.838		"5 m above Moinakom River (00°41'N, 124°3'E), Dumoga Nature Reserve, North Sulawesi, Indonesia, altitude 525 m."	0.6833	124.05	tatar Bergmans & Rozendaal, 1982	split from R. euryotis	Patrick, L.E., McCulloch, E.S., Ruedas, L.A. 2013. Systematics and biogeography of the arcuate horseshoe bat species complex (Chiroptera, Rhinolophidae). Zoologica Scripta. doi: 10.1111/zsc.12026.	Indonesia	Asia	Australasia/Oceania	LC	0	0	0	Rhinolophus_tatar	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Rhinolophus tatar	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	tatar	Bergmans and Rozendaal, 1982	<span lang="EN-US">This species was considered to be conspecific with R. euryotis <span lang="EN-US">(</span><span lang="EN-US">Hill 1992<span lang="EN-US">)</span><span lang="EN-US">. However, genetic work by Patrick et al . <span lang="EN-US">(</span><span lang="EN-US">2013<span lang="EN-US">) ;</span><span lang="EN-US">demonstrated that specimens from Sulawesi were genetically distinct from those collected on Papua New Guinea and deserving of distinct species status.  </span>	80000000	Rhinolophus tatar	Least Concern		2017	2016-06-25 00:00:00 UTC	3.1	English	This species is listed as Least Concern (LC) on the basis of its widespread distribution in Sulawesi, its stable population trend and the absence of threats affecting the whole population. D<span lang="EN-US">eforestation and human disturbance of roost sites, however, could still adversely affect populations at a local scale.	<p><span lang="EN-US">This species is a gleaning or aerial insectivore, as are all members of the genus <span lang="EN-US">(<span lang="EN-US">Hill 1992<span lang="EN-US">)<span lang="EN-US">. ;It is possible that it roosts in caves although more likely a tree rooster. This species has been mistnetted in slightly disturbed rainforest lowland to lower montane forest <span lang="EN-US">(<span lang="EN-US">Bergmans and Rozendaal 1982<span lang="EN-US">)<span lang="EN-US">. ;</span></span></span></span></p>	<p><span lang="EN-US">While this species is not currently under threat throughout its range, deforestation and human disturbance of roost sites could adversely affect local populations <span lang="EN-US">(<span lang="EN-US">Wiles and Brooke 2009<span lang="EN-US">)<span lang="EN-US">.</span></span></p>	<p><span lang="EN-US">The species is recorded from throughout the island of Sulawesi <span lang="EN-US">(<span lang="EN-US">Hill 1992<span lang="EN-US">)<span lang="EN-US">; no data are available at the current time with respect to population sizes and trends, however, although these parameters are unknown, populations are assumed to be large and stable.</span></span></p>	Unknown	<p><span lang="EN-US">The species as currently defined has been found across the island of Sulawesi <span lang="EN-US">(Hill 1992<span lang="EN-US">)<span lang="EN-US">.</span></span></p>		Terrestrial		Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		tatar	Bergmans and Rozendaal	1982	0	Bijdr. Dierk.	54:50:00	Tatar Horseshoe Bat	None.		Sulawesi	Not listed.	Least Concern	Formerly included in euryotis, but see Patrick et al. 2013.	Rhinolophus tatar	1004752	23	Sulawesi Broad-eared Horseshoe Bat	Tatar Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	tatar	Bergmans & Rozendaal	1982	0	Rhinolophus_tatar	Bergmans, W., & Rozendaal, F. G. (1982). Notes on Rhinolophus Lacépède, 1799 from Sulawesi, Indonesia, with the description of a new species (Mammalia, Microchiroptera). Bijdragen tot de Dierkunde, 52(2), 170.	https://brill.com/view/journals/btd/52/2/article-p169_7.xml	ZMA 21.838		"5 m above Moinakom River (00°41'N, 124°3'E), Dumoga Nature Reserve, North Sulawesi, Indonesia, altitude 525 m."	0.683333	124.05	tatar Bergmans & Rozendaal, 1982	split from R. euryotis	Patrick, L.E., McCulloch, E.S., Ruedas, L.A. 2013. Systematics and biogeography of the arcuate horseshoe bat species complex (Chiroptera, Rhinolophidae). Zoologica Scripta. doi: 10.1111/zsc.12026.				Indonesia	Asia	Australasia/Oceania	LC	0	0	0	Rhinolophus_tatar	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_tatar	1004752	23	Sulawesi Broad-eared Horseshoe Bat	Tatar Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	tatar	Bergmans & F. G. Rozendaal	0	Rhinolophus tatar	Bergmans, W. and Rozendaal, F.G. 1982. Notes on Rhinolophus _Lacépède_, 1799 from Sulawesi, Indonesia, with the description of a new species (Mammalia, Microchiroptera). Bijdragen tot de Dierkunde 52(2):169-174.	https://brill.com/view/journals/btd/52/2/article-p169_7.xml	ZMA.MAM.21838.a, ZMA.MAM.21838.b	holotype	https://data.biodiversitydata.nl/naturalis/specimen/ZMA.MAM.21838.a | https://data.biodiversitydata.nl/naturalis/specimen/ZMA.MAM.21838.b	"5 m above Moinakom River (00°41'N, 124°3'E), Dumoga Nature Reserve, North Sulawesi, Indonesia, altitude 525 m."	0.683333	124.05	split from R. euryotis	Patrick, L.E., McCulloch, E.S., Ruedas, L.A. 2013. Systematics and biogeography of the arcuate horseshoe bat species complex (Chiroptera, Rhinolophidae). Zoologica Scripta. doi: 10.1111/zsc.12026.				Indonesia	Asia	Australasia	LC	0	0	0	Rhinolophus_tatar	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		tatar	Bergmans & Rozendaal	1982	0	Bijdr.    Dierk.	54:50:00	Tatar Horseshoe Bat	None.		Sulawesi	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84372447/84372450/' target='_blank'>Least Concern</a>	Formerly included in euryotis, but see Patrick et al. 2013.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus tatar; Rhinolophus tatar; Rhinolophus tatar; Rhinolophus tatar; Rhinolophus tatar; tatar; Sulawesi Broad-eared Horseshoe Bat; Rhinolophe d'Archbold; Tate-Archbold-Hufeisennase; Herradura de Archbold; Tatar Horseshoe Bat; Sulawesi Broad-eared Horseshoe Bat; Tatar Horseshoe Bat; Tatar Horseshoe Bat; R. tatar