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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1446	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Rhinolophus macrotis [synonym of]	Rhinolophus macrotis siamensis	Rhinolophus siamensis	Rhinolophus siamensis	Rhinolophus siamensis	Rhinolophus siamensis	Rhinolophus siamensis	Rhinolophus siamensis	Rhinolophus siamensis	Rhinolophus siamensis	Rhinolophus siamensis	Rhinolophus siamensis		[MSW3] Formerly included in macrotis but apparently distinct, see Francis et al. (1999b) and Hendrichsen et al. (2001b).; [HMW] Rhinolophus macrotis siamensis Gyldenstolpe, 1917 , “ Doi Par Sakeng, Northwestern Siam [= Thailand ]. ” Rhinolophus siamensis is in the macrotis species group and is genetically sister to R episcopus , which was recently validated as a distinct species from R macrotis . Rhinolophus siamensis generally has been included as a subspecies of R macrotis , but is now regarded as a separate species based on broad sympatry and lack of hybridization. Recently described. huananus is now considered a synonym of AE siamensis based on genetic data and apparent lack of reliable differentiating morphological traits. Monotypic.; [batnames2022] Formerly included in macrotis but apparently distinct, see Francis et al. (1999b), Hendrichsen et al. (2001b), and Liu et al. (2019). Includes huananus ; see Zhang et al. (2018); [MDD2022] includes the recently described huananus; [IUCN] Originally described as a subspecies of R. macrotis but proved to be a distinct species (see Francis 1999, Hendrichsen et al. 2001, Simmons 2005). This includes huananus (see Zhang et al. 2009, Zhang et al. 2018).; [batnames2023] Formerly included in macrotis but apparently distinct, see Francis et al. (1999b), Hendrichsen et al. (2001b), and Liu et al. (2019). Includes huananus ; see Zhang et al. (2018); [MDD2023] includes the recently described huananus; [MDD2025_2.0] includes the recently described huananus; [batnames2025_1.7] Formerly included in macrotis but apparently distinct, see Francis et al. (1999b), Hendrichsen et al. (2001b), and Liu et al. (2019). Includes huananus; see Zhang et al. (2018); [MDD2025_2.2] includes the recently described huananus											huananus	siamensis 	siamensis - huananus	siamensis, huananus	Originally described as a subspecies of R. macrotis but proved to be a distinct species (see Francis 1999, Hendrichsen et al. 2001, Simmons 2005). This includes huananus (see Zhang et al. 2009, Zhang et al. 2018).	siamensis 	siamensis - huananus	siamensis, huananus	siamensis, huananus	siamensis	siamensis - huananus	siamensis Gyldenstolpe, 1916|huananus Wu Yi, Motokawa, & Harada, 2008						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus siamensis	Rhinolophus		siamensis	Gyldenstolpe		1917		Kungliga Svenska VetenskAkad. Handl.	57		12		Thai Horseshoe Bat	Thailand, NW Thailand, Doi Par Sakang.	Thailand, Laos, Vietnam.	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).		Formerly included in macrotis but apparently distinct, see Francis et al. (1999b) and Hendrichsen et al. (2001b).	885887A2FFC08A26FF1CF68FF7B5DA4F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	301	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFD98A3EF89CEFB2FE0DD1D4.xml	Rhinolophus siamensis	Rhinolophidae	Rhinolophus	siamensis	Gyldenstolpe	1916	Thai Horseshoe Bat @en | Rhinolophe siamois @fr | Thai-Hufeisennase @de | Herradura siamesa @es | Siamese Horseshoe Bat @en	Rhinolophus macrotis siamensis Gyldenstolpe, 1917 , “ Doi Par Sakeng, Northwestern Siam [= Thailand ]. ” Rhinolophus siamensis is in the macrotis species group and is genetically sister to R episcopus , which was recently validated as a distinct species from R macrotis . Rhinolophus siamensis generally has been included as a subspecies of R macrotis , but is now regarded as a separate species based on broad sympatry and lack of hybridization. Recently described. huananus is now considered a synonym of AE siamensis based on genetic data and apparent lack of reliable differentiating morphological traits. Monotypic.	S China ( Yunnan , Guangxi , Hunan , and Guangdong ), N Thailand , N & C Laos , and N & C Vietnam .	Head-body 33- 5— 40 mm , tail 13-22 mm , ear 19-22- 6 mm , hindfoot 8-9 mm , forearm 36-41 mm ; weight 4-5—6- 4 g . The Thai Horseshoe Bat is similar to the Big-eared Horseshoe Bat ( AE macrotis ) but distinguished by its much higher frequency calls and smaller size. Dorsal pelage is pale brown; venter is paler buff. There is no known orange morph. Males lack axillary tufts. Ears are proportionally long. Noseleaf has subtriangular lancet, with narrow, rounded tip and convex sides; connecting process is rounded with straight anterior edge when viewed laterally; sella is hairy, broad, rounded at tip, long, and upward facing; and horseshoe is broad at 6-7-8- 3 mm , completely covers muzzle, has moderately developed lateral leaflets concealed by horseshoe, and has small but distinct median emargination. Lower lip has three medial grooves. Skull is similar to the Big-eared Horseshoe Bat. P2 is in tooth row and relatively large, separating C1 and P4, and P3 is small and labially displaced from tooth row.	Apparently tropical and subtropical moist forests in lowland or middle elevation habitats.	No information.	No information.	Thai Horseshoe Bats are nocturnal. They are known to roost in caves. Call shape is FM/CF/FM, with F component recordeed at 66-71 -8 kHz in China and 68—74 kHz in Laos .	No information.	Classified as Least Concern on The IUCN Red List. There are currently no known threats to the Thai Horseshoe Bat, although little is known about its ecology and natural history. Additional research is needed.	Chiozza (2008a) | Csorba et al. (2003) | Francis (2008a) | Liu Tong et al. (2019) | Smith & XieYan (2008) | Tu Vuong Tan, Hassanin et al. (2017) | WuYi et al. (2008) | Zhang Libiao, Jones et al. (2009) | Zhang Lin et al. (2018)	https://zenodo.org/record/3749994/files/figure.png	45 . Thai Horseshoe Bat Rhinolophus siamensis French: Rhinolophe siamois / German: Thai-Hufeisennase / Spanish: Herradura siamesa Other common names: Siamese Horseshoe Bat Taxonomy. Rhinolophus macrotis siamensis Gyldenstolpe, 1917 , “ Doi Par Sakeng, Northwestern Siam [= Thailand ]. ” Rhinolophus siamensis is in the macrotis species group and is genetically sister to R episcopus , which was recently validated as a distinct species from R macrotis . Rhinolophus siamensis generally has been included as a subspecies of R macrotis , but is now regarded as a separate species based on broad sympatry and lack of hybridization. Recently described. huananus is now considered a synonym of AE siamensis based on genetic data and apparent lack of reliable differentiating morphological traits. Monotypic. Distribution. S China ( Yunnan , Guangxi , Hunan , and Guangdong ), N Thailand , N & C Laos , and N & C Vietnam . Descriptive notes. Head-body 33- 5— 40 mm , tail 13-22 mm , ear 19-22- 6 mm , hindfoot 8-9 mm , forearm 36-41 mm ; weight 4-5—6- 4 g . The Thai Horseshoe Bat is similar to the Big-eared Horseshoe Bat ( AE macrotis ) but distinguished by its much higher frequency calls and smaller size. Dorsal pelage is pale brown; venter is paler buff. There is no known orange morph. Males lack axillary tufts. Ears are proportionally long. Noseleaf has subtriangular lancet, with narrow, rounded tip and convex sides; connecting process is rounded with straight anterior edge when viewed laterally; sella is hairy, broad, rounded at tip, long, and upward facing; and horseshoe is broad at 6-7-8- 3 mm , completely covers muzzle, has moderately developed lateral leaflets concealed by horseshoe, and has small but distinct median emargination. Lower lip has three medial grooves. Skull is similar to the Big-eared Horseshoe Bat. P2 is in tooth row and relatively large, separating C1 and P4, and P3 is small and labially displaced from tooth row. Habitat. Apparently tropical and subtropical moist forests in lowland or middle elevation habitats. Food and Feeding. No information. Breeding. No information. Activity patterns. Thai Horseshoe Bats are nocturnal. They are known to roost in caves. Call shape is FM/CF/FM, with F component recordeed at 66-71 -8 kHz in China and 68—74 kHz in Laos . Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. There are currently no known threats to the Thai Horseshoe Bat, although little is known about its ecology and natural history. Additional research is needed. Bibliography. Chiozza (2008a), Csorba eta/. (2003), Francis (2008a), Liu Tong eta/. (2019), Smith & XieYan (2008), Tu Vuong Tan, Hassanin et al. (2017), WuYi et al. (2008), Zhang Libiao, Jones et al. (2009), Zhang Lin eta/. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus siamensis	Rhinolophus		siamensis	Gyldenstolpe	1917	0	Kungliga Svenska VetenskAkad. Handl.	######	Northern Thai Horseshoe Bat	 huananus Wu, Motokawa & Harada, 2008.	Thailand, NW Thailand, Doi Par Sakang.	S China, Thailand, Laos, Vietnam.	Not listed.	Least Concern	Formerly included in macrotis but apparently distinct, see Francis et al. (1999b), Hendrichsen et al. (2001b), and Liu et al. (2019). Includes huananus ; see Zhang et al. (2018)	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus siamensis	23	Thai Horseshoe Bat	Siamese Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	siamensis	Gyldenstolpe	1917	0	Rhinolophus_macrotis_siamensis	Gyldenstolpe, N. (1917). Zoological results of the Swedish zoological expeditions to Siam 1911-1912 & 1914-1915. V. Mammals ii. Kungliga Svenska vetenskapsakademiens handlingar, 57, 12.	https://www.biodiversitylibrary.org/item/131280#page/84/mode/1up			"Doi Par Sakeng, Northwestern Siam [= Thailand]."			siamensis Gyldenstolpe, 1917|huananus Wu Yi, Motokawa, & Harada, 2008	includes the recently described huananus	Wu, Y., Motokawa, M., & Harada, M. (2008). A new species of horseshoe bat of the genus Rhinolophus from China (Chiroptera: Rhinolophidae). Zoological Science, 25(4), 438-443.|Zhang, L., Sun, K., Liu, T., Zhao, H., Csorba, G., Jin, L., ... & Feng, J. (2018). Multilocus phylogeny and species delimitation within the philippinensis group (Chiroptera: Rhinolophidae). Zoologica Scripta, 47(6), 655-672.	China|Thailand|Laos|Vietnam	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_siamensis	0	sciname match	Rhinolophus_siamensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136651	Rhinolophus siamensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	siamensis	Gyldenstolpe, 1917	Originally described as a subspecies of R. macrotis but proved to be a distinct species (see Francis 1999, Hendrichsen et al. 2001, Simmons 2005). This includes huananus (see Zhang et al. 2009, Zhang et al. 2018).	20000000	Rhinolophus siamensis	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Confirmed as Least Concern, because its extent of occurrence is much larger than 20,000 km², and although there is no recent information on its population status it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species has been found broadly in sympatry with other species of the R. macrotis complex in karst areas in southern China, Vietnam, Laos, and Thailand. It was usually found roosting in limestone caves and forages in different habitats ranging from heavily disturbed to primary evergreen forests (Francis 1999, Tu et al. 2017).	This species is vulnerable to cave disturbance or destruction and forest degradation mainly through tourist and extractive industry (Furey et al. 2010). In many regions in Vietnam and possibly in Laos, illegal harvesting for local consumption is also identified as additional threats to this species (V.T. Tu. pers. comm.).	The abundance and population size of this species are not known.	Unknown	This species ranges from Yunnan Province in China, southwards extending in to Thailand, Lao PDR, and Viet Nam.	In many regions in Vietnam and possibly in Laos species is illegal harvested for local consumption (V.T. Tu. pers. comm.).	Terrestrial	It is not known if the species has been recorded from any protected areas. Further studies are needed into the distribution, abundance, natural history and threats to this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		siamensis	Gyldenstolpe	1917	0	Kungliga Svenska VetenskAkad. Handl.	57:12:00	Northern Thai Horseshoe Bat	 huananus Wu, Motokawa & Harada, 2008.	Thailand, NW Thailand, Doi Par Sakang.	S China, Thailand, Laos, Vietnam.	Not listed.	Least Concern	Formerly included in macrotis but apparently distinct, see Francis et al. (1999b), Hendrichsen et al. (2001b), and Liu et al. (2019). Includes huananus ; see Zhang et al. (2018)	Rhinolophus siamensis	1004743	23	Thai Horseshoe Bat	Siamese Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	siamensis	Gyldenstolpe	1917	0	Rhinolophus_macrotis_siamensis	Gyldenstolpe, N. (1917). Zoological results of the Swedish zoological expeditions to Siam 1911-1912 & 1914-1915. V. Mammals ii. Kungliga Svenska vetenskapsakademiens handlingar, 57, 12.	https://www.biodiversitylibrary.org/item/131280#page/84/mode/1up			"Doi Par Sakeng, Northwestern Siam [= Thailand]."			siamensis Gyldenstolpe, 1917|huananus Wu Yi, Motokawa, & Harada, 2008	includes the recently described huananus	Wu, Y., Motokawa, M., & Harada, M. (2008). A new species of horseshoe bat of the genus Rhinolophus from China (Chiroptera: Rhinolophidae). Zoological Science, 25(4), 438-443.|Zhang, L., Sun, K., Liu, T., Zhao, H., Csorba, G., Jin, L., ... & Feng, J. (2018). Multilocus phylogeny and species delimitation within the philippinensis group (Chiroptera: Rhinolophidae). Zoologica Scripta, 47(6), 655-672.				China|Thailand|Laos|Vietnam	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_siamensis	0	sciname match	Rhinolophus_siamensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_siamensis	1004743	23	Thai Horseshoe Bat	Siamese Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	siamensis	Gyldenstolpe	0	Rhinolophus macrotis siamensis	Gyldenstolpe, N. 1916. Zoological results of the Swedish zoological expeditions to Siam. 1911–1912 & 1914–1915. V. Mammals II. Kunglika Svenska Vetenskapsakademiens Handlingar 57(2):1-59.	https://www.biodiversitylibrary.org/page/41891600		holotype		"Doi Par Sakeng, Northwestern Siam [= Thailand]."			includes the recently described huananus	Wu, Y., Motokawa, M., & Harada, M. (2008). A new species of horseshoe bat of the genus Rhinolophus from China (Chiroptera: Rhinolophidae). Zoological Science, 25(4), 438-443.|Zhang, L., Sun, K., Liu, T., Zhao, H., Csorba, G., Jin, L., ... & Feng, J. (2018). Multilocus phylogeny and species delimitation within the philippinensis group (Chiroptera: Rhinolophidae). Zoologica Scripta, 47(6), 655-672.				China|Thailand|Laos|Vietnam	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_siamensis	0	sciname match	Rhinolophus_siamensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		siamensis	Gyldenstolpe	1917	0	Kungliga Svenska VetenskAkad. Handl.	57:12:00	Northern Thai Horseshoe Bat	huananus Wu, Motokawa & Harada, 2008.	Thailand, NW Thailand, Doi Par Sakang.	S China, Thailand, Laos, Vietnam.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136651/21990143/' target='_blank'>Least Concern</a>	Formerly included in macrotis but apparently distinct, see Francis et al. (1999b), Hendrichsen et al. (2001b), and Liu et al. (2019). Includes huananus; see Zhang et al. (2018)		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus siamensis; Rhinolophus siamensis; Rhinolophus siamensis; Rhinolophus siamensis; Rhinolophus siamensis; Rhinolophus siamensis; huananus; huananus; siamensis; huananus; Thai Horseshoe Bat; Rhinolophe siamois; Thai-Hufeisennase; Herradura siamesa; Siamese Horseshoe Bat; Thai Horseshoe Bat; Siamese Horseshoe Bat; Thai Horseshoe Bat; Northern Thai Horseshoe Bat; R. siamensis