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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1445	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Rhinolophus lepidus [synonym of]	Rhinolophus lepidus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei	Rhinolophus shortridgei		[MSW3] pusillus species group. Formerly considered a subspecies of lepidus, but recently captured in sympatry with that species and thus clearly distinct; see Csorba (2002) and Csorba et al. (2003).; [HMW] Rhinolophus lepidus shortridgei K. Andersen, 1918 , “ Pagan, R[ iver ]. Irrawaddy , Burma [= Myanmar ].” Rhinolophus shortridgei is included in the pusiUus species group; it is close to R monticolus, and together these two make R pusiUus paraphyletic. Rhinolophus shortridgei was previously included in R lepidus , but is recognized as a distinct species based on morphological and genetic differences presented by G. Csorba and colleagues in 2003 and by P. Soisook and others in 2016. The precise distribution of this species has not been well established in the areas where it comes together with R lepidus , especially as the two are sympatric in some areas. Monotypic.; [batnames2022]  pusillus species group. Formerly considered a subspecies of lepidus, but recently captured in sympatry with that species and thus clearlydistinct; see Csorba (2002) and Csorba et al. (2003).; [IUCN] Until recently this species was considered to be a subspecies of Rhinolophus lepidus , but it was recently captured in sympatry with R. lepidus and is therefore obviously distinct (Csorba 2002, Csorba et al . 2003).; [batnames2023]  pusillus species group. Formerly considered a subspecies of lepidus, but recently captured in sympatry with that species and thus clearlydistinct; see Csorba (2002) and Csorba et al. (2003).; [batnames2025_1.7] pusillusspecies group. Formerly considered a subspecies of lepidus, but recently captured in sympatry with that species and thus clearlydistinct; see Csorba (2002) and Csorba et al. (2003).														shortridgei	Until recently this species was considered to be a subspecies of Rhinolophus lepidus , but it was recently captured in sympatry with R. lepidus and is therefore obviously distinct (Csorba 2002, Csorba et al . 2003).			shortridgei 	shortridgei 			shortridgei Andersen, 1918						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus shortridgei	Rhinolophus		shortridgei	K. Andersen		1918		Ann. Mag. Nat. Hist., ser. 9	2		376		Shortridge's Horseshoe Bat	Burma, Irrawaddy River, Pagan (= Bagan).	N India, Burma	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).		pusillus species group. Formerly considered a subspecies of lepidus, but recently captured in sympatry with that species and thus clearly distinct; see Csorba (2002) and Csorba et al. (2003).	885887A2FFD98A3EF89CEFB2FE0DD1D4	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	312	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFEE8A08F8B6FB8EF622D41C.xml	Rhinolophus shortridgei	Rhinolophidae	Rhinolophus	shortridgei	K. Andersen	1918	Rhinolophede Shortridge @fr | Shortridge-Hufeisennase @de | Herradura de Shortridge @es	Rhinolophus lepidus shortridgei K. Andersen, 1918 , “ Pagan, R[ iver ]. Irrawaddy , Burma [= Myanmar ].” Rhinolophus shortridgei is included in the pusiUus species group; it is close to R monticolus, and together these two make R pusiUus paraphyletic. Rhinolophus shortridgei was previously included in R lepidus , but is recognized as a distinct species based on morphological and genetic differences presented by G. Csorba and colleagues in 2003 and by P. Soisook and others in 2016. The precise distribution of this species has not been well established in the areas where it comes together with R lepidus , especially as the two are sympatric in some areas. Monotypic.	NE India (Falta, West Bengal ), N & C Myanmar , and S & SE China .			No information.	No information.	Shortridge’s Horseshoe Bat is nocturnal. Search call shape is FM/ CF/FM with a peak F recorded at 94-5-100-7 kHz in Myanmar , females having higher frequency calls than males.	No information.	Classified as Least Concern on The IUCNed List. Shortridge’s Horseshoe Bat is widespread, and there are currently no major threats known to be affecting it. Virtually nothing is known of this species’ ecology, and further studies are needed.	Bates et al. (2004) | Chiozza (2008b) | Csorba (2002) | Csorba et al. (2003) | rancis (2008a) | Smith & XieYan (2008) | Soisook et al. (2016)	https://zenodo.org/record/3750040/files/figure.png	67 . Shortridge’s Horseshoe Bat Rhinolophus shortridgei French: Rhinolophe de Shortridge / German: Shortridge-Hufeisennase / Spanish: Herradura de Shortridge Taxonomy. Rhinolophus lepidus shortridgei K. Andersen, 1918 , “ Pagan, R[ iver ]. Irrawaddy , Burma [= Myanmar ].” Rhinolophus shortridgei is included in the pusiUus species group; it is close to R monticolus, and together these two make R pusiUus paraphyletic. Rhinolophus shortridgei was previously included in R lepidus , but is recognized as a distinct species based on morphological and genetic differences presented by G. Csorba and colleagues in 2003 and by P. Soisook and others in 2016. The precise distribution of this species has not been well established in the areas where it comes together with R lepidus , especially as the two are sympatric in some areas. Monotypic. Distribution. NE India (Falta, West Bengal ), N & C Myanmar , and S & SE China . Descriptive notes. Head-body 40- 6-59 mm , tail 16-1-29- 4 mm , ear 14-20 mm , hindfoot 6-9- 7 mm , forearm 38- 2—43 mm ; weight 5-5—8- 8 g . Shortridge’s Horseshoe Bat is at present externally inseparable from Blyth’s Horseshoe Bat ( lepidus ); only dried study skins are known for the species, so the external characteristics remain rather uncertain. Dorsal pelage is fight brown on the upper side, whereas ventral pelage is much paler, dirty white or gray . Ears are moderate in size. Noseleaf has a concave-sided lancet; connecting process is a pointed triangle; sella is more or less parallel-sided and widely rounded above; horseshoe does not cover muzzle (6-7— 7-9 mm wide) and has small lateral leaflets. Lower lip has three mental grooves. Skull is robust (zygomatic width is larger than mastoid width); anterior median swellings are low and very small; posterior compartments are relatively large; rostral profile is slightly curved upward near tip; sagittal crest is very well developed; frontal depression is shallow; supraorbital crests are conspicuous. C1 is widebased and long; P2 is medium-sized and within tooth row; P3 is small to moderate in size and half to fully displaced from tooth row, usually separating P2 from P4. Habitat In Myanmar , Shortridge’s Horseshoe Bat has been collected in seasonally dry dipterocarp forest. Food and Feeding. No information. Breeding. No information. Activity patterns. Shortridge’s Horseshoe Bat is nocturnal. Search call shape is FM/ CF/FM with a peak F recorded at 94-5-100-7 kHz in Myanmar , females having higher frequency calls than males. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCNed List. Shortridge’s Horseshoe Bat is widespread, and there are currently no major threats known to be affecting it. Virtually nothing is known of this species’ ecology, and further studies are needed. Bibliography. Bates et al. (2004), Chiozza (2008b), Csorba (2002), Csorba eta/. (2003), rancis (2008a), Smith & XieYan (2008), Soisook et al. (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus shortridgei	Rhinolophus		shortridgei	K. Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 376	Shortridge's Horseshoe Bat	None.	Burma, Irrawaddy River, Pagan (= Bagan).	N India, Burma	Not listed.	Data Deficient	 pusillus species group. Formerly considered a subspecies of lepidus, but recently captured in sympatry with that species and thus clearlydistinct; see Csorba (2002) and Csorba et al. (2003).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus shortridgei	23	Shortridge's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	shortridgei	K. Andersen	1918	0	Rhinolophus_lepidus_shortridgei	Andersen, K. (1918). Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History; Zoology, Botany, and Geology, Ser. 9, 2, 376.	https://www.biodiversitylibrary.org/item/53366#page/408/mode/1up	BM 1918.8.3.1		"Pagan, R[iver]. Irrawaddy, Burma [= Myanmar]."			shortridgei K. Andersen, 1918	NA	NA	India|Myanmar|China	Asia	Indomalaya|Palearctic	DD	0	0	0	Rhinolophus_shortridgei	0	sciname match	Rhinolophus_shortridgei	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136631	Rhinolophus shortridgei	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	shortridgei	K. Andersen, 1918	Until recently this species was considered to be a subspecies of Rhinolophus lepidus , but it was recently captured in sympatry with R. lepidus and is therefore obviously distinct (Csorba 2002, Csorba et al . 2003).	20000000	Rhinolophus shortridgei	Data Deficient		2020	2018-09-15 00:00:00 UTC	3.1	English	Listed as Data Deficient, because there is no information available on the population and abundance of this species.	There is little information available for this species. In Myanmar, it has been collected in seasonally dry dipterocarp forest.	The threats to this species are not known.	There is no information available on the population and abundance of this species.	Unknown	<p><a style="mso-comment-reference:DW_1;mso-comment-date:20190412T2030">This species occurs in northern India and Myanmar (Csorba et al . 2003, Simmons 2005).</a></p>		Terrestrial	It is not known if the species is present in any protected areas. Further studies are needed into the taxonomy, distribution, abundance, ecology, and threats to this species.	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		shortridgei	K. Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 376	Shortridge's Horseshoe Bat	None.	Burma, Irrawaddy River, Pagan (= Bagan).	N India, Burma	Not listed.	Data Deficient	 pusillus species group. Formerly considered a subspecies of lepidus, but recently captured in sympatry with that species and thus clearlydistinct; see Csorba (2002) and Csorba et al. (2003).	Rhinolophus shortridgei	1004742	23	Shortridge's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	shortridgei	K. Andersen	1918	0	Rhinolophus_lepidus_shortridgei	Andersen, K. (1918). Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History; Zoology, Botany, and Geology, Ser. 9, 2, 376.	https://www.biodiversitylibrary.org/item/53366#page/408/mode/1up	BM 1918.8.3.1		"Pagan, R[iver]. Irrawaddy, Burma [= Myanmar]."			shortridgei K. Andersen, 1918	NA	NA				India|Myanmar|China	Asia	Indomalaya|Palearctic	DD	0	0	0	Rhinolophus_shortridgei	0	sciname match	Rhinolophus_shortridgei	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_shortridgei	1004742	23	Shortridge's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	shortridgei	Andersen	0	Rhinolophus lepidus shortridgei	Andersen, K.C. 1918-10-01. Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History (9)2(10):374-384.	https://www.biodiversitylibrary.org/page/22103259	BMNH:Mamm:1918.8.3.1	holotype	https://data.nhm.ac.uk/object/e365f5e1-6fea-43ab-8413-7fe3154d4455	"Pagan, R[iver]. Irrawaddy, Burma [= Myanmar]."			NA	NA				India|Myanmar|China	Asia	Indomalaya|Palearctic	DD	0	0	0	Rhinolophus_shortridgei	0	sciname match	Rhinolophus_shortridgei	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		shortridgei	K. Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 376	Shortridge's Horseshoe Bat	None.	Burma, Irrawaddy River, Pagan (= Bagan).	N India, Burma	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136631/21987430/' target='_blank'>Data Deficient</a>	pusillusspecies group. Formerly considered a subspecies of lepidus, but recently captured in sympatry with that species and thus clearlydistinct; see Csorba (2002) and Csorba et al. (2003).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus shortridgei; Rhinolophus shortridgei; Rhinolophus shortridgei; Rhinolophus shortridgei; Rhinolophus shortridgei; Rhinolophus shortridgei; shortridgei; Rhinolophede Shortridge; Shortridge-Hufeisennase; Herradura de Shortridge; Shortridge's Horseshoe Bat; Shortridge's Horseshoe Bat; Shortridge's Horseshoe Bat; R. shortridgei