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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1441	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Rhinolophus sakejiensis	Rhinolophus sakejiensis	Rhinolophus sakejiensis	Rhinolophus sakejiensis	Rhinolophus sakejiensis	Rhinolophus sakejiensis	Rhinolophus sakejiensis	Rhinolophus sakejiensis	Rhinolophus sakejiensis	Rhinolophus sakejiensis		[MSW3] ferrumequinum species group.; [HMW] Rhinolophus sakejiensis Cotterill, 2002 , “ Kavunda ( 11°17’ S ; 24°2 ' E ) between the Sakeji and Zambezi rivers: c. 11 km north-north-east of the source of the Zambezi River in the Ikelenge Pedicle, Mwinilunga District of north-west Zambia .” Rhinolophus sakejiensis is in the ferrumequinum species group based on morphology, but its exact relationships are uncertain. Monotypic.; [batnames2022]  ferrumequinum species group.; [batnames2023]  ferrumequinum species group.; [batnames2025_1.7] ferrumequinumspecies group.														sakejiensis				sakejiensis	sakejiensis			sakejiensis Cotterill, 2002						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus sakejiensis	Rhinolophus		sakejiensis	Cotterill		2002		J. Zool.	256		166		Sakeji Horseshoe Bat	Zambia, Mwinilunga District, Ikelenge Pedicle between the Sakeji and Zambezi Rivers, approx. 11 km NNE of source of Zambezi River, Kavunda, 11°17'S, 24°21'E, 1,388 m.	Known only from the type locality.	IUCN 2003 – Not evaluated (new species).		ferrumequinum species group.	885887A2FFD88A3EFF1BF687F405DBD8	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	294	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFC08A39F8B3F157FAAEDE93.xml	Rhinolophus sakejiensis	Rhinolophidae	Rhinolophus	sakejiensis	Cotterill	2002	Sakeji Horseshoe Bat @en | Rhinolophe de la Sakeji @fr | Sakeji-Hufeisennase @de | Herradura de Sakeji @es	Rhinolophus sakejiensis Cotterill, 2002 , “ Kavunda ( 11°17’ S ; 24°2 ' E ) between the Sakeji and Zambezi rivers: c. 11 km north-north-east of the source of the Zambezi River in the Ikelenge Pedicle, Mwinilunga District of north-west Zambia .” Rhinolophus sakejiensis is in the ferrumequinum species group based on morphology, but its exact relationships are uncertain. Monotypic.	Known only from type locality in NW Zambia .	Head—body c .57-58 nun, tail 29—31 mm , ear 20—22 mm , hindfoot 14—15 mm , forearm 53—55 mm ; weight 19—24 g . Dorsal pelage is bright brownish orange (hairs are orange with darker tips); venter is bright yellowish orange; and eyes are surrounded by darker brown fur. No gray morph has been reported. Males seem to lack axillary tufts. Ears are short and dark brown. Noseleaf has hastate lancet; connecting process rises steeply to high, narrow, and rounded tip, being much higher than height of sella; sella is naked with broad, rounded tip that is angled forward and with slightly concave sides; and horseshoe is medium in width at c . 10-11 mm , does not cover entire muzzle and has lateral leaflets and wide, deep median emargination. Lower lip has one conspicuous median groove and two faint lateral grooves. Wings and uropatagium are dark blackish brown. Baculum is trumpet-shaped, with dorso-ventrally flattened shaft. Skull is extremely robust, with robust zygomatic arches (zygomatic width is wider than mastoid width); nasal swellings are relatively low and not smoothly rounded; frontal depression is very shallow and supraorbital ridges are inconspicuous; sagittal crest is well developed anteriorly but weakly developed posteriorly ; and interpterygoid groove is shallow. P2 is absent, allowing C1 and P4 to touch, and P3 is absent, allowing P2 and P4 to touch. Dental formula is 11/2, C 1/1, P 1/2, M 3/3 (x2) = 28.	Wet miombo woodland mosaic at an elevation of 1388 m .	No information.	No information.	Only known specimens of the Sakeji Horseshoe Bat were collected while roosting during the day, hanging in evergreen foliage on the underside of a branch at base of a large tree. This kind of day roost is rather unusual for a rhinolophid.	All known specimens of the Sakeji Horseshoe Bat were captured from a day roost that had a group of six individuals.	Classified as Data Deficient on The IUCNRed List. The Sakeji Horseshoe Bat is known from only three specimens at a single location, and no information on its natural history and ecology is known. It might be threatened by logging and agricultural expansion, but additional research is needed.	ACR (2018) | Cotterill (2002a, 2008a, 2013c) | Csorba et al. (2003)	https://zenodo.org/record/3749966/files/figure.png	29 . Sakeji Horseshoe Bat Rhinolophus sakejiensis French: Rhinolophe de la Sakeji / German: Sakeji-Hufeisennase / Spanish: Herradura de Sakeji Taxonomy. Rhinolophus sakejiensis Cotterill, 2002 , “ Kavunda ( 11°17’ S ; 24°2 ' E ) between the Sakeji and Zambezi rivers: c. 11 km north-north-east of the source of the Zambezi River in the Ikelenge Pedicle, Mwinilunga District of north-west Zambia .” Rhinolophus sakejiensis is in the ferrumequinum species group based on morphology, but its exact relationships are uncertain. Monotypic. Distribution. Known only from type locality in NW Zambia . Descriptive notes. Head—body c .57-58 nun, tail 29—31 mm , ear 20—22 mm , hindfoot 14—15 mm , forearm 53—55 mm ; weight 19—24 g . Dorsal pelage is bright brownish orange (hairs are orange with darker tips); venter is bright yellowish orange; and eyes are surrounded by darker brown fur. No gray morph has been reported. Males seem to lack axillary tufts. Ears are short and dark brown. Noseleaf has hastate lancet; connecting process rises steeply to high, narrow, and rounded tip, being much higher than height of sella; sella is naked with broad, rounded tip that is angled forward and with slightly concave sides; and horseshoe is medium in width at c . 10-11 mm , does not cover entire muzzle and has lateral leaflets and wide, deep median emargination. Lower lip has one conspicuous median groove and two faint lateral grooves. Wings and uropatagium are dark blackish brown. Baculum is trumpet-shaped, with dorso-ventrally flattened shaft. Skull is extremely robust, with robust zygomatic arches (zygomatic width is wider than mastoid width); nasal swellings are relatively low and not smoothly rounded; frontal depression is very shallow and supraorbital ridges are inconspicuous; sagittal crest is well developed anteriorly but weakly developed posteriorly ; and interpterygoid groove is shallow. P2 is absent, allowing C1 and P4 to touch, and P3 is absent, allowing P2 and P4 to touch. Dental formula is 11/2, C 1/1, P 1/2, M 3/3 (x2) = 28. Habitat. Wet miombo woodland mosaic at an elevation of 1388 m . Food and Feeding. No information. Breeding. No information. Activity patterns. Only known specimens of the Sakeji Horseshoe Bat were collected while roosting during the day, hanging in evergreen foliage on the underside of a branch at base of a large tree. This kind of day roost is rather unusual for a rhinolophid. Movements, Home range and Social organization. All known specimens of the Sakeji Horseshoe Bat were captured from a day roost that had a group of six individuals. Status and Conservation. Classified as Data Deficient on The IUCNRed List. The Sakeji Horseshoe Bat is known from only three specimens at a single location, and no information on its natural history and ecology is known. It might be threatened by logging and agricultural expansion, but additional research is needed. Bibliography. ACR (2018), Cotterill (2002a, 2008a, 2013c), Csorba et al. (2003).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus sakejiensis	Rhinolophus		sakejiensis	Cotterill	2002	0	J. Zool.	258:46:00	Sakeji Horseshoe Bat	None.	Zambia, Mwinilunga District, Ikelenge Pedicle between the Sakeji and Zambezi Rivers, approx. 11 km NNE of source of Zambezi River, Kavunda, 11Â°17'S, 24Â°21'E, 1,388 m.	Known only from the type locality.	Not listed.	Data Deficient	 ferrumequinum species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus sakejiensis	23	Sakeji Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	sakejiensis	Cotterill	2002	0	Rhinolophus_sakejiensis	Cotterill, F. P. D. (2002). A new species of horseshoe bat (Microchiroptera: Rhinolophidae) from south-central Africa: with comments on its affinities and evolution, and the characterization of a rhinolophid species. Journal of Zoology, 256(2), 166.	https://zslpublications.onlinelibrary.wiley.com/doi/abs/10.1017/S0952836902000201?casa_token=EOpiDcWU2kkAAAAA:y_n1b89mAPQezq5hI2X_gf48BStkzwjjs35Thrgkpl2lYXe8gtrfK0_aFn0k5MQZajy-b1LpT9GyqA	NMZB 29153		"Kavunda (11Â°17' S; 24Â°21' E) between the Sakeji and Zambezi rivers: c. 11 km north-north-east of the source of the Zambezi River in the Ikelenge Pedicle, Mwinilunga District of north-west Zambia."	-11.28	24.35	sakejiensis Cotterill, 2002	NA	NA	Zambia	Africa	Afrotropic	DD	0	0	0	Rhinolophus_sakejiensis	0	sciname match	Rhinolophus_sakejiensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	44785	Rhinolophus sakejiensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	sakejiensis	Cotterill, 2002		20000000	Rhinolophus sakejiensis	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	Confirmed as Data Deficient since it has only recently been discovered, and there is still very little information on its extent of occurrence, ecological requirements and threats.	Cotterill (2002) collected the species hanging within evergreen foliage on the underside of a branch at the base of a large tree, within a stand of gallery forest within mesic miombo woodland. It is also reported to be found within roosting within hollow trees (Cotterill 2002).	The threats to this species are not well known, but it is presumably threatened by the loss of roosting sites resulting from logging and the conversion of suitable habitat to agricultural use.	It is currently known only from the type collection of three bats, taken from a cluster of six roosting bats.	Unknown	This recently described bat species is known only from the type locality of Kavunda, between the Sakeji and Zambezi rivers: c. 11 km north-north-east of the source of the Zambezi River in the Ikelenge Pedicle, Mwinilunga District of north-west Zambia (Cotterill 2002). It is likely that the species is also present within similar suitable habitat in Angola and the Congo basin (Cotterill 2002).		Terrestrial	It is not known if the species is present in any protected areas. Further surveys are needed to better determine the extent of the distribution of this species, to gather more information on the species natural history, and to review the threats to this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		sakejiensis	Cotterill	2002	0	J. Zool.	258:46:00	Sakeji Horseshoe Bat	None.	Zambia, Mwinilunga District, Ikelenge Pedicle between the Sakeji and Zambezi Rivers, approx. 11 km NNE of source of Zambezi River, Kavunda, 11Â°17'S, 24Â°21'E, 1,388 m.	Known only from the type locality.	Not listed.	Data Deficient	 ferrumequinum species group.	Rhinolophus sakejiensis	1004738	23	Sakeji Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	sakejiensis	Cotterill	2002	0	Rhinolophus_sakejiensis	Cotterill, F. P. D. (2002). A new species of horseshoe bat (Microchiroptera: Rhinolophidae) from south-central Africa: with comments on its affinities and evolution, and the characterization of a rhinolophid species. Journal of Zoology, 256(2), 166.	https://zslpublications.onlinelibrary.wiley.com/doi/abs/10.1017/S0952836902000201?casa_token=EOpiDcWU2kkAAAAA:y_n1b89mAPQezq5hI2X_gf48BStkzwjjs35Thrgkpl2lYXe8gtrfK0_aFn0k5MQZajy-b1LpT9GyqA	NMZB 29153		"Kavunda (11Â°17' S; 24Â°21' E) between the Sakeji and Zambezi rivers: c. 11 km north-north-east of the source of the Zambezi River in the Ikelenge Pedicle, Mwinilunga District of north-west Zambia."	-11.2833	24.35	sakejiensis Cotterill, 2002	NA	NA				Zambia	Africa	Afrotropic	DD	0	0	0	Rhinolophus_sakejiensis	0	sciname match	Rhinolophus_sakejiensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_sakejiensis	1004738	23	Sakeji Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	sakejiensis	Cotterill	0	Rhinolophus sakejiensis	Cotterill, F.P.D. 2002-02. A new species of horseshoe bat (Microchiroptera: Rhinolophidae) from south-central Africa: with comments on its affinities and evolution, and the characterization of rhinolophid species. Journal of Zoology, London 256(2):165-179.	https://doi.org/10.1017/S0952836902000201	NHMBZ 29153	holotype		"Kavunda (11Â°17' S; 24Â°21' E) between the Sakeji and Zambezi rivers: c. 11 km north-north-east of the source of the Zambezi River in the Ikelenge Pedicle, Mwinilunga District of north-west Zambia."	-11.2833	24.35	NA	NA				Zambia	Africa	Afrotropic	DD	0	0	0	Rhinolophus_sakejiensis	0	sciname match	Rhinolophus_sakejiensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		sakejiensis	Cotterill	2002	0	J. Zool.	258:46:00	Sakeji Horseshoe Bat	None.	Zambia, Mwinilunga District, Ikelenge Pedicle between the Sakeji and Zambezi Rivers, approx. 11 km NNE of source of Zambezi River, Kavunda, 11Â°17'S, 24Â°21'E, 1,388 m.	Known only from the type locality.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/44785/22068998/' target='_blank'>Data Deficient</a>	ferrumequinumspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus sakejiensis; Rhinolophus sakejiensis; Rhinolophus sakejiensis; Rhinolophus schnitzleri; Rhinolophus sakejiensis; sakejiensis; Sakeji Horseshoe Bat; Rhinolophe de la Sakeji; Sakeji-Hufeisennase; Herradura de Sakeji; Sakeji Horseshoe Bat; Sakeji Horseshoe Bat; R. sakejiensis
