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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1420	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Rhinolophus cornutus [synonym of]	N/A	Rhinolophus imaizumii?	Rhinolophus perditus	Rhinolophus perditus	Rhinolophus perditus	Rhinolophus perditus	Rhinolophus perditus	Rhinolophus perditus	Rhinolophus perditus	Rhinolophus perditus	Rhinolophus perditus		[MSW3] pusillus species group. Reviewed by Yoshiyuki (1989).; [HMW] Rhinolophus perditus K. Andersen, 1918 , “ Ishigaki, southern Liu-Kiu [= Ryukyu Islands ] /’ Japan . Rhinolophus perditus is included in the pusillus group. It is occasionally treated as a subspecies or synonym of R. comutus , but it is morphologically distinguishable from R. comutus occurring allopatrically on other Ryukyu Islands, and can also be differentiated based on echolocation. There are differing opinions regarding the taxonomic position of R. imaizumii in relation to R.perditus . The two taxa are morphologically distinguishable based on craniodental morphometries, but no genetic data have yet been shown to differentiate them; they have variously been treated as distinct species or as a single species, R.perditus (including imaizumii as ajunior synonym). It is possible that both may occur on some of the islands, lending to potential further confusion, but this has not yet been proven. Until a focused study using both genetic and morphological data is undertaken, the two taxa are considered subspecies under R.perditus . Two subspecies are tentatively recognized.; [batnames2022]  pusillus species group; distinct from cornutus and imaizumii ; see Wu et al. (2012). See also Yoshiyuki (1989), Csorba et al. (2003), and Sano and Armstrong (2009). Csorba et al. (2019) consider imaizumii as subspecies of perditus , but given the data presented by Wu et al. (2012), we continue to recognize it as a distinct species.; [MDD2022] split from R. cornutus; may include imaizumi, but the species is tentatively retained here following Cirranello and Simmons Bat Database; [batnames2023]  pusillus species group; distinct from cornutus and imaizumii ; see Wu et al. (2012). See also Yoshiyuki (1989), Csorba et al. (2003), and Sano and Armstrong (2009). Csorba et al. (2019) consider imaizumii as subspecies of perditus , but given the data presented by Wu et al. (2012), we continue to recognize it as a distinct species.; [MDD2023] split from R. cornutus; may include imaizumi, but the species is tentatively retained here following Cirranello and Simmons Bat Database; [MDD2025_2.0] split from R. cornutus; may include imaizumi, but the species is tentatively retained here following Cirranello and Simmons Bat Database; [batnames2025_1.7] pusillus species group; distinct from cornutus and imaizumii; see Wu et al. (2012). See also Yoshiyuki (1989), Csorba et al. (2003), and Sano and Armstrong (2009). Burgin (2019) considers imaizumii as subspecies of perditus, but given the data presented by Wu et al. (2012), we continue to recognize it as a distinct species.; [MDD2025_2.2] split from R. cornutus; may include imaizumi, but the species is tentatively retained here following Cirranello and Simmons Bat Database										perditus, imaizumii				perditus				perditus	perditus			perditus Andersen, 1918						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus imaizumii	Rhinolophus		imaizumii	Hill and Yoshiyuki		1980		Bull. Natl. Sci. Mus. Tokyo, ser. A (Zool.)	6		180		Imaizumi's Horseshoe Bat	Japan, Ryukyu Isls, Yayeyama Isls, Iriomote Isl, Otomi-do cave.	Iriomote Isl and Yaeyama Isl (Japan: Ryukyu Isls).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Endangered.		pusillus species group. Reviewed by Yoshiyuki (1989).	885887A2FFFE8A1BF8B4EFE6FC7FC876	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	311	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFEF8A08F8ADF66FFC95D040.xml	Rhinolophus perditus	Rhinolophidae	Rhinolophus	perditus	K. Andersen	1918	Yaeyama Horseshoe Bat @en | Rhinolophe des Yaeyama @fr | Yaeyama-Hufeisennase @de | Herradura de Yaeyama @es | Spanish @en | Lost Horseshoe Bat @en | Ryukyu Horseshoe Bat @en | Yaeyama Little Horseshoe Bat; @en | maizumi's Horseshoe Bat (imaizumii) @en	Rhinolophus perditus K. Andersen, 1918 , “ Ishigaki, southern Liu-Kiu [= Ryukyu Islands ] /’ Japan . Rhinolophus perditus is included in the pusillus group. It is occasionally treated as a subspecies or synonym of R. comutus , but it is morphologically distinguishable from R. comutus occurring allopatrically on other Ryukyu Islands, and can also be differentiated based on echolocation. There are differing opinions regarding the taxonomic position of R. imaizumii in relation to R.perditus . The two taxa are morphologically distinguishable based on craniodental morphometries, but no genetic data have yet been shown to differentiate them; they have variously been treated as distinct species or as a single species, R.perditus (including imaizumii as ajunior synonym). It is possible that both may occur on some of the islands, lending to potential further confusion, but this has not yet been proven. Until a focused study using both genetic and morphological data is undertaken, the two taxa are considered subspecies under R.perditus . Two subspecies are tentatively recognized.	R.p. perditus K. Andersen, 1918 - Yaeyama Is (Ishigakijima and Taketomi), Ryukyu Is. R.p. imaizumii Hill & Yoshiyuki, 1980 — Yaeyama Is (Iriomotejima and Kohamajima), Ryukyu Is.	Head-body 39-47- 8 mm , tail 18-1-26- 1 mm , ear 17-9-18- 9 mm , hindfoot 8- 4—9 mm , forearm 40- 3—43 mm . The Yaeyama Horseshoe Bat is similar to the Littlejapanese Horseshoe Bat (. cornutus ). Subspecies imaizumii is larger than perditus and the two taxa do not overlap in craniodental morphometries. Pelage is woolly and glossy, dorsal pelage being bright brown (base of hairs light beige), ventral pelage similar but paler and creamier brown. Ears are moderate-sized. Noseleaf has a slightly concave-sided lancet; connecting process is hom-like, curved anteriorly, narrow, and slender; sella has a broad base with a slightly constricted center and a rounded, anteriorly deflected tip; horseshoe is not very broad (7- 3 mm ), with no lateral leaflets, and a short, narrow median emargination. Lower lip has three mental grooves. Baculum is 4-2-4- 4 mm long and has a very deep dorsal notch on the basal cone with a small ventral notch; shaft is nearly cylindrical and is bent on the distal one-half. Skull is similar to that of the LittleJapanese Horseshoe Bat but can be distinguished based on craniodental measurements; skull is long and narrow, and zygomatic width is slighdy greater than ( imaizumii ) or subequal to ( perditus ) mastoid width; anterior median swellings are small and subcircular; posterior compartments are moderate in size; rostral profile slopes posteriorly ; sagittal crest is weakly to moderately developed; supraorbital crests are weak; frontal depression is shallow. Dental distinctions between subspecies are small but present; imaizumii has a higher crown portion of C1 compared to perditus , this being comparatively stout in perditus , C-M3 tooth row is slightly shorter in perditus but zygomatic width is greater than in imaizumii ; P2 and P3 are within the tooth row in perditus but both are extruded from the tooth row internally in imaizumii . Chromosomal complement has 2n = 62 and FNa = 60.	Forested habitats.	The Yaeyama Horseshoe Bat feeds on arthropods. Study of wing morphology showed that the species is suspected to forage in cluttered spaces. It has been recorded feeding largely on Lepidoptera, Coleoptera , Hymenoptera , and Diptera , although it also occasionally takes Orthoptera, Neuroptera, Trichoptera , and Araneae . Presence of Araneae in the diet indicates that the species probably also gleans prey off foliage.	The Yaeyama Horseshoe Bat exhibits restricted seasonal monoestry; females form maternity colonies in May, and give birth to a single young around this time.	Yaeyama Horseshoe Bats are active throughout the year, but probably hibernate through the winter with regular bouts of arousal to forage. They roost in caves, abandoned mines, and bomb shelters during the day and forage by night. Search call shape is FM/CF/ FM with a peak F recorded at 96-98 kHz on Ishigakÿima and 92-93 kHz on Iriomotejima.	The Yaeyama Horseshoe Bat roosts in colonies and females can form maternity colonies with over 1000 individuals while raising young. This species frequently roosts with the Ryukyu Long-fingered Bat (Miniopterusfuscus) in caves.	Not assessed as a separate species on TheIUCNed List, where it is included under the Least Horseshoe Bat (pusiUus) as Least Concern. Listed as endangered in the Japanese Red List. Several caves for breeding and hibernating are known to exist on Ishigakÿima Island. However, in recent years, deforestation, roost destruction and tourism activities are creating serious problems for the island bat population. As a consequence, conservation measures need to be established immediately.	Ando et al. (1980c) | Chen Shiangfan et al. (2006) | Fukui et al. (2009) | Hill & Yoshiyuki (1980) | Hutson, Kingston & Walston (2008) | Li Gang et al. (2006) | Ohdachi et al. (2015) | WuYi , Motokawa et al. (2012) | Yoshiyuki (1989) | Zhang Lin et al. (2018)	https://zenodo.org/record/3750036/files/figure.png	65 . Yaeyama Horseshoe Bat Rhinolophus perditus French: Rhinolophe des Yaeyama / German: Yaeyama-Hufeisennase I Spanish: Herradura de Yaeyama Other common names: Lost Horseshoe Bat, Ryukyu Horseshoe Bat, Yaeyama Little Horseshoe Bat; Imaizumi's Horseshoe Bat ( imaizumii ) Taxonomy. Rhinolophus perditus K. Andersen, 1918 , “ Ishigaki, southern Liu-Kiu [= Ryukyu Islands ] /’ Japan . Rhinolophus perditus is included in the pusillus group. It is occasionally treated as a subspecies or synonym of R. comutus , but it is morphologically distinguishable from R. comutus occurring allopatrically on other Ryukyu Islands, and can also be differentiated based on echolocation. There are differing opinions regarding the taxonomic position of R. imaizumii in relation to R.perditus . The two taxa are morphologically distinguishable based on craniodental morphometries, but no genetic data have yet been shown to differentiate them; they have variously been treated as distinct species or as a single species, R.perditus (including imaizumii as ajunior synonym). It is possible that both may occur on some of the islands, lending to potential further confusion, but this has not yet been proven. Until a focused study using both genetic and morphological data is undertaken, the two taxa are considered subspecies under R.perditus . Two subspecies are tentatively recognized. Subspecies and Distribution. R.p. perditus K. Andersen, 1918 - Yaeyama Is (Ishigakijima and Taketomi), Ryukyu Is. R.p. imaizumii Hill & Yoshiyuki, 1980 — Yaeyama Is (Iriomotejima and Kohamajima), Ryukyu Is. Descriptive notes. Head-body 39-47- 8 mm , tail 18-1-26- 1 mm , ear 17-9-18- 9 mm , hindfoot 8- 4—9 mm , forearm 40- 3—43 mm . The Yaeyama Horseshoe Bat is similar to the Littlejapanese Horseshoe Bat (. cornutus ). Subspecies imaizumii is larger than perditus and the two taxa do not overlap in craniodental morphometries. Pelage is woolly and glossy, dorsal pelage being bright brown (base of hairs light beige), ventral pelage similar but paler and creamier brown. Ears are moderate-sized. Noseleaf has a slightly concave-sided lancet; connecting process is hom-like, curved anteriorly, narrow, and slender; sella has a broad base with a slightly constricted center and a rounded, anteriorly deflected tip; horseshoe is not very broad (7- 3 mm ), with no lateral leaflets, and a short, narrow median emargination. Lower lip has three mental grooves. Baculum is 4-2-4- 4 mm long and has a very deep dorsal notch on the basal cone with a small ventral notch; shaft is nearly cylindrical and is bent on the distal one-half. Skull is similar to that of the LittleJapanese Horseshoe Bat but can be distinguished based on craniodental measurements; skull is long and narrow, and zygomatic width is slighdy greater than ( imaizumii ) or subequal to ( perditus ) mastoid width; anterior median swellings are small and subcircular; posterior compartments are moderate in size; rostral profile slopes posteriorly ; sagittal crest is weakly to moderately developed; supraorbital crests are weak; frontal depression is shallow. Dental distinctions between subspecies are small but present; imaizumii has a higher crown portion of C1 compared to perditus , this being comparatively stout in perditus , C-M3 tooth row is slightly shorter in perditus but zygomatic width is greater than in imaizumii ; P2 and P3 are within the tooth row in perditus but both are extruded from the tooth row internally in imaizumii . Chromosomal complement has 2n = 62 and FNa = 60. Habitat. Forested habitats. Food and Feeding. The Yaeyama Horseshoe Bat feeds on arthropods. Study of wing morphology showed that the species is suspected to forage in cluttered spaces. It has been recorded feeding largely on Lepidoptera, Coleoptera , Hymenoptera , and Diptera , although it also occasionally takes Orthoptera, Neuroptera, Trichoptera , and Araneae . Presence of Araneae in the diet indicates that the species probably also gleans prey off foliage. Breeding. The Yaeyama Horseshoe Bat exhibits restricted seasonal monoestry; females form maternity colonies in May, and give birth to a single young around this time. Activity patterns. Yaeyama Horseshoe Bats are active throughout the year, but probably hibernate through the winter with regular bouts of arousal to forage. They roost in caves, abandoned mines, and bomb shelters during the day and forage by night. Search call shape is FM/CF/ FM with a peak F recorded at 96-98 kHz on Ishigakÿima and 92-93 kHz on Iriomotejima. Movements, Home range and Social organization. The Yaeyama Horseshoe Bat roosts in colonies and females can form maternity colonies with over 1000 individuals while raising young. This species frequently roosts with the Ryukyu Long-fingered Bat (Miniopterusfuscus) in caves. Status and Conservation. Not assessed as a separate species on TheIUCNed List, where it is included under the Least Horseshoe Bat (pusiUus) as Least Concern. Listed as endangered in the Japanese Red List. Several caves for breeding and hibernating are known to exist on Ishigakÿima Island. However, in recent years, deforestation, roost destruction and tourism activities are creating serious problems for the island bat population. As a consequence, conservation measures need to be established immediately. Bibliography. Ando eta /. (1980c), Chen Shiangfan eta/. (2006), Fukui eta/. (2009), Hill & Yoshiyuki (1980), Hutson, Kingston & Walston (2008), Li Gang eta/. (2006), Ohdachi eta/. (2015), WuYi , Motokawa eta/. (2012), Yoshiyuki (1989), Zhang Lin eta/. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus perditus	Rhinolophus		perditus	Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 377	Yaeyama Horseshoe Bat	None.	Japan, southern Liu-Kiu, Ishigaki	Yaeyama Group (Ryuku Isls)	Not listed.	Endangered	 pusillus species group; distinct from cornutus and imaizumii ; see Wu et al. (2012). See also Yoshiyuki (1989), Csorba et al. (2003), and Sano and Armstrong (2009). Csorba et al. (2019) consider imaizumii as subspecies of perditus , but given the data presented by Wu et al. (2012), we continue to recognize it as a distinct species.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus perditus	23	Yaeyama Horseshoe Bat	Lost Horseshoe Bat|Ryukyu Horseshoe Bat|Yaeyama Little Horseshoe Bat|Imaizumi's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	perditus	K. Andersen	1918	0	Rhinolophus_perditus	Andersen, K. (1918). Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History; Zoology, Botany, and Geology, Ser. 9, 2, 377.	https://www.biodiversitylibrary.org/item/53366#page/409/mode/1up	BM 1905.11.3.15		"Ishigaki, southern Liu-Kiu [= Ryukyu Islands]," Japan.			perditus K. Andersen, 1918	split from R. cornutus; may include imaizumi, but the species is tentatively retained here following Cirranello and Simmons Bat Database	Wu, Y., Motokawa, M., Harada, M., Thong, V. D., Lin, L. K., & Li, Y. C. (2012). Morphometric variation in the pusillus group of the genus Rhinolophus (Mammalia: Chiroptera: Rhinolophidae) in East Asia. Zoological science, 29(6), 396-402.	Japan	Asia	Palearctic	EN	0	0	0	Rhinolophus_perditus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Rhinolophus perditus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	perditus	K. Andersen, 1918		90000000	Rhinolophus perditus	Endangered	B1ab(iii)+2ab(iii)	2020	2018-08-31 00:00:00 UTC	3.1	English	This species is listed as Endangered because its area of occupancy (AOO) is 410 km², estimated by total area of forests in the four islands (531.56 km²) where this species is present. The species also has an extent of occurrence (EOO) smaller than 5,000 km². It occurs in fewer than 5 locations and its range is severely fragmented. The species is dependent on a very specific habitat (karst areas), and there is ongoing decline in the quality of its habitat mainly due to deforestation of the surrounding foraging areas and destruction of its day roosts.	This species roosts in natural caves, abandoned mines, underground culverts and bomb shelters (Sano and Armstrong 2015). Females form maternity colonies from hundreds to over 1,000 individuals in May and give birth to single young (Sano and Armstrong 2015). This species feeds mainly on Lepidoptera, Coleoptera, Hymenoptera and Diptera (Fukui et al. 2009).	The species roosts in natural caves. Roost disturbance from humans/tourists (e.g., some caves have electricity lined in for tourist activities), and the threat of development around caves represent the main threats to the species. An airport has been constructed above several caves used by this species (1,200-2,500 and 990-1,185 individuals during maternity and hibernation periods, respectively between 2002-2006) on the Ishigaki Island. After construction, the number of roosting individuals in these caves have not significantly declined during the maternity season (1,500-2,000 individuals between 2013-2014), but have declined during hibernation periods (about 600-1,000 individuals between 2013-2014) (Kara Karst Scientific Committee 2014, New Ishigaki Airport Bat Monitoring Committee 2015).	Many previously observed colonies in caves have already disappeared. On Ishigaki island, a large maternity colony with 3,000 newborn young has been recorded (Ministry of the Environment 2014). Population size for this species on Ishigaki island in 2004 was estimated to be at least 6,500 (Koyanagi et al.  2013), and between 2004 and 2011 the population size was estimated as 5,000-8,000 individuals (New Ishigaki Airport Bat Monitoring Committee 2015).	Decreasing	This species is known only from the Yaeyama (Sakishima) Islands of Japan, including Iriomote, Ishigaki, Kohama and Taketomi islands (Sano and Armstrong 2015) and is not likely to occur more widely.		Terrestrial	Near the airport, an artificial cave has been constructed as an alternative roost, but only few individuals used it so far. The species is listed as Vulnerable (VU) in the Japanese Red List (Ministry of the Environment 2014).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		perditus	Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 377	Yaeyama Horseshoe Bat	None.	Japan, southern Liu-Kiu, Ishigaki	Yaeyama Group (Ryuku Isls)	Not listed.	Endangered	 pusillus species group; distinct from cornutus and imaizumii ; see Wu et al. (2012). See also Yoshiyuki (1989), Csorba et al. (2003), and Sano and Armstrong (2009). Csorba et al. (2019) consider imaizumii as subspecies of perditus , but given the data presented by Wu et al. (2012), we continue to recognize it as a distinct species.	Rhinolophus perditus	1004725	23	Yaeyama Horseshoe Bat	Lost Horseshoe Bat|Ryukyu Horseshoe Bat|Yaeyama Little Horseshoe Bat|Imaizumi's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	perditus	K. Andersen	1918	0	Rhinolophus_perditus	Andersen, K. (1918). Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History; Zoology, Botany, and Geology, Ser. 9, 2, 377.	https://www.biodiversitylibrary.org/item/53366#page/409/mode/1up	BM 1905.11.3.15		"Ishigaki, southern Liu-Kiu [= Ryukyu Islands]," Japan.			perditus K. Andersen, 1918	split from R. cornutus; may include imaizumi, but the species is tentatively retained here following Cirranello and Simmons Bat Database	Wu, Y., Motokawa, M., Harada, M., Thong, V. D., Lin, L. K., & Li, Y. C. (2012). Morphometric variation in the pusillus group of the genus Rhinolophus (Mammalia: Chiroptera: Rhinolophidae) in East Asia. Zoological science, 29(6), 396-402.				Japan	Asia	Palearctic	EN	0	0	0	Rhinolophus_perditus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_perditus	1004725	23	Yaeyama Horseshoe Bat	Lost Horseshoe Bat|Ryukyu Horseshoe Bat|Yaeyama Little Horseshoe Bat|Imaizumi's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	perditus	Andersen	0	Rhinolophus perditus	Andersen, K.C. 1918-10-01. Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History (9)2(10):374-384.	https://www.biodiversitylibrary.org/page/22103259	BMNH:Mamm:1905.11.3.15	holotype	https://data.nhm.ac.uk/object/7e9ae616-80e3-40e8-a76d-620f514bd499	"Ishigaki, southern Liu-Kiu [= Ryukyu Islands]," Japan.			split from R. cornutus; may include imaizumi, but the species is tentatively retained here following Cirranello and Simmons Bat Database	Wu, Y., Motokawa, M., Harada, M., Thong, V. D., Lin, L. K., & Li, Y. C. (2012). Morphometric variation in the pusillus group of the genus Rhinolophus (Mammalia: Chiroptera: Rhinolophidae) in East Asia. Zoological science, 29(6), 396-402.				Japan	Asia	Palearctic	EN	0	0	0	Rhinolophus_perditus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		perditus	K. Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 377	Yaeyama Horseshoe Bat	None.	Japan, southern Liu-Kiu, Ishigaki	Yaeyama Group (Ryuku Isls)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85707170/85707174/' target='_blank'>Endangered</a>	pusillus species group; distinct from cornutus and imaizumii; see Wu et al. (2012). See also Yoshiyuki (1989), Csorba et al. (2003), and Sano and Armstrong (2009). Burgin (2019) considers imaizumii as subspecies of perditus, but given the data presented by Wu et al. (2012), we continue to recognize it as a distinct species.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus imaizumii; Rhinolophus perditus; Rhinolophus perditus; Rhinolophus perditus; Rhinolophus perditus; Rhinolophus perditus; perditus; imaizumii; perditus; Yaeyama Horseshoe Bat; Rhinolophe des Yaeyama; Yaeyama-Hufeisennase; Herradura de Yaeyama; Spanish; Lost Horseshoe Bat; Ryukyu Horseshoe Bat; Yaeyama Little Horseshoe Bat; maizumi's Horseshoe Bat (imaizumii); Yaeyama Horseshoe Bat; Lost Horseshoe Bat; Ryukyu Horseshoe Bat; Yaeyama Little Horseshoe Bat; Imaizumi's Horseshoe Bat; Imaizumi's Horseshoe Bat; Yaeyama Horseshoe Bat; R. perditus