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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1418	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus lepidus [synonym of]	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi	Rhinolophus osgoodi		[MSW3] pusillus species group. Possibly conspecific with lepidus (Corbet and Hill, 1992), but treated as distinct following Csorba et al. (2003).; [HMW] Rhinolophus osgoodi Sanborn, 1939 , “ Nguluko ( 25° 5’ N . Lat., 100° 15’ E . Long.), north of Likiang , Yunnan , China . ” Rhinolophus osgoodi was formerly in the pusillus group but is in the macrotis group here based on molecular studies and morphological similarities. Rhinolophus osgoodi is sometimes considered a subspecies of R lepidus , but they are morphologically distinct. Recent genetic studies have shown that a clade composed of a paraphyletic assemblage of A siamensis and R macrotis actually represents three distinct species, R osgoodi (specimens originally identified as “ Rhinolophus spi ”), R episcopus , and R siamensis . Until then, R osgoodi was only known from specimens from the type locality. Monotypic.; [batnames2022]  macrotis species group. See Liu et al. (2019) and Csorba et al. (2003); but see Corbet and Hill (1992).; [IUCN] Rhinolophus osgoodi was considered to be conspecific with R. lepidus and belong to pusillus species group (Simmons 2005). Call frequency recorded for R. osgoodi (95.2 ;kHz) is close to that reported in Francis and Habersetzer (1998) for R. lepidus (98 ;kHz) in Malaysia. However, this species is different from the species from the pusillus species group in genetics (unpublished data) and some phenotypic characters, such as the higher anterior swellings and long palatal bridge, which suggested a different taxonomic position (unpublished data, Csorba pers. Comm.).; [batnames2023]  macrotis species group. See Liu et al. (2019) and Csorba et al. (2003); but see Corbet and Hill (1992).; [batnames2025_1.7] macrotisspecies group. See Liu et al. (2019) and Csorba et al. (2003); but see Corbet and Hill (1992).														osgoodi	Rhinolophus osgoodi was considered to be conspecific with R. lepidus and belong to pusillus species group (Simmons 2005). Call frequency recorded for R. osgoodi (95.2 ;kHz) is close to that reported in Francis and Habersetzer (1998) for R. lepidus (98 ;kHz) in Malaysia. However, this species is different from the species from the pusillus species group in genetics (unpublished data) and some phenotypic characters, such as the higher anterior swellings and long palatal bridge, which suggested a different taxonomic position (unpublished data, Csorba pers. Comm.).			osgoodi	osgoodi			osgoodi Sanborn, 1939		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Yunnan, China	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinolophus osgoodi	China, Yunnan, N. of Likiang, Nguluko.	Sanborn	1939	Field Mus. Nat. Hist. Publ. Ser. Zool., 24:40.	Distribution: Known only from Yunnan.					Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Sanborn	1939	Field Mus. Nat. Hist. Publ., Zool. Ser., 24:40.		Yunnan (China).	China, Yunnan, N of Likiang, Nguluko.		SANBORN	1939	Connecting process triangular but somewhat rounded. Lancet not extremely hastate. Size me dium (forearm length, 41 -4 6 mm) but skull rela tively small (greatest length, 16-17 mm; maxilla ry tooth row length, 5.5-5.8 mm). Rostrum rela tively slender.	Distribution: Known only from Yunnan.	No subspecies.		55	species	R. osgoodi	SANBORN	1939	Rhinolophus	genus	Rhinolophus osgoodi				Connecting process triangular but somewhat rounded. Lancet not extremely hastate. Size me dium (forearm length, 41 -4 6 mm) but skull rela tively small (greatest length, 16-17 mm; maxilla ry tooth row length, 5.5-5.8 mm). Rostrum rela tively slender.	No subspecies.		30. R. osgoodi SANBORN 1939 [pusillus group].	30	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus osgoodi	Rhinolophus		osgoodi	Sanborn		1939		Field Mus. Nat. Hist. Publ., Zool. Ser.	24		40		Osgood's Horseshoe Bat	China, Yunnan, N of Likiang, Nguluko, (27°05'N, 100°15'E).	Yunnan (China).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Data Deficient.		pusillus species group. Possibly conspecific with lepidus (Corbet and Hill, 1992), but treated as distinct following Csorba et al. (2003).	885887A2FFFA8A1CFF01F5D8F291D0BB	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	301	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFD98A3FFF5AF5DDF5B2DB9A.xml	Rhinolophus osgoodi	Rhinolophidae	Rhinolophus	osgoodi	Sanborn	1939	Osgood’s Horseshoe Bat @en | Rhinolophe d'Osgood @fr | Osgood-Hufeisennase @de | Herradura de Osgood @es	Rhinolophus osgoodi Sanborn, 1939 , “ Nguluko ( 25° 5’ N . Lat., 100° 15’ E . Long.), north of Likiang , Yunnan , China . ” Rhinolophus osgoodi was formerly in the pusillus group but is in the macrotis group here based on molecular studies and morphological similarities. Rhinolophus osgoodi is sometimes considered a subspecies of R lepidus , but they are morphologically distinct. Recent genetic studies have shown that a clade composed of a paraphyletic assemblage of A siamensis and R macrotis actually represents three distinct species, R osgoodi (specimens originally identified as “ Rhinolophus spi ”), R episcopus , and R siamensis . Until then, R osgoodi was only known from specimens from the type locality. Monotypic.	S China ( Sichuan , Yunnan , Guizhou , and Hunan ) and N Vietnam .	Tail 15-22 mm , ear 10-14 mm , hindfoot 7-9- 5 mm , forearm 40- 45 mm ; weight 5-6 g . Dorsal pelage is pale brown (hairs have gray bases); venter is paler and grayer. Ears are medium to large in size. There is no known orange morph. Males lack axillary tufts. Noseleaf has relatively short, almost straight-sided lancet with pointed tip; connecting process is high, but more or less pointed; sella is relatively broad, with parallel sides and widely rounded tip; and horseshoe is wide ( c .6- 4 mm ), covers entire muzzle, has very shallow median emargination, and lacks lateral leaflets. Skull is moderately robust; zygomatic width is smaller than mastoid width; anterior median swellings and posterior compartments are moderately developed; rostral profile is sloped backward; sagittal crest is weakly developed; and frontal depression is shallow and bordered by sharp, well-defined supraorbital ridges. P2 is of medium size and in tooth row, separating C1 and P4; P3 is small and completely or partially displaced labially; and P 2 and P4 are very close to one another but do not touch.	Caves surrounded by secondary growth or agricultural fields.	No information.	No information.	Osgood’s Horseshoe Bats are nocturnal. Call shape is FM /CF/FM, with F component at 85—94 kHz.	No information.	Classified as Data Deficient on 77 ie IUCN Red List. Osgood’s Horseshoe Bat was previous known only from Yunnan , but recent genetic analysis has shown that the species is much more widespread than previously thought. However, additional research is needed regarding ecology, distribution, and threats of this species.	Csorba et al. (2003) | LiuTong et al. (2019) | Smith & XieYan (2008) | Smith, Johnston, Jones & Rossiter (2008 a) | Zhang Libiao , Jones et al. (2009)	https://zenodo.org/record/3749990/files/figure.png	43 . Osgood’s Horseshoe Bat Rhinolophus osgoodi French: Rhinolophe d'Osgood / German: Osgood-Hufeisennase / Spanish: Herradura de Osgood Taxonomy. Rhinolophus osgoodi Sanborn, 1939 , “ Nguluko ( 25° 5’ N . Lat., 100° 15’ E . Long.), north of Likiang , Yunnan , China . ” Rhinolophus osgoodi was formerly in the pusillus group but is in the macrotis group here based on molecular studies and morphological similarities. Rhinolophus osgoodi is sometimes considered a subspecies of R lepidus , but they are morphologically distinct. Recent genetic studies have shown that a clade composed of a paraphyletic assemblage of A siamensis and R macrotis actually represents three distinct species, R osgoodi (specimens originally identified as “ Rhinolophus spi ”), R episcopus , and R siamensis . Until then, R osgoodi was only known from specimens from the type locality. Monotypic. Distribution. S China ( Sichuan , Yunnan , Guizhou , and Hunan ) and N Vietnam . Descriptive notes. Tail 15-22 mm , ear 10-14 mm , hindfoot 7-9- 5 mm , forearm 40- 45 mm ; weight 5-6 g . Dorsal pelage is pale brown (hairs have gray bases); venter is paler and grayer. Ears are medium to large in size. There is no known orange morph. Males lack axillary tufts. Noseleaf has relatively short, almost straight-sided lancet with pointed tip; connecting process is high, but more or less pointed; sella is relatively broad, with parallel sides and widely rounded tip; and horseshoe is wide ( c .6- 4 mm ), covers entire muzzle, has very shallow median emargination, and lacks lateral leaflets. Skull is moderately robust; zygomatic width is smaller than mastoid width; anterior median swellings and posterior compartments are moderately developed; rostral profile is sloped backward; sagittal crest is weakly developed; and frontal depression is shallow and bordered by sharp, well-defined supraorbital ridges. P2 is of medium size and in tooth row, separating C1 and P4; P3 is small and completely or partially displaced labially; and P 2 and P4 are very close to one another but do not touch. Habitat. Caves surrounded by secondary growth or agricultural fields. Food and Feeding. No information. Breeding. No information. Activity patterns. Osgood’s Horseshoe Bats are nocturnal. Call shape is FM /CF/FM, with F component at 85—94 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on 77 ie IUCN Red List. Osgood’s Horseshoe Bat was previous known only from Yunnan , but recent genetic analysis has shown that the species is much more widespread than previously thought. However, additional research is needed regarding ecology, distribution, and threats of this species. Bibliography. Csorba et al. (2003), LiuTong et al. (2019), Smith & XieYan (2008), Smith, Johnston, Jones & Rossiter (2008 a ), Zhang Libiao , Jones et al. (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus osgoodi	Rhinolophus		osgoodi	Sanborn	1939	0	Field Mus. Nat. Hist. Publ., Zool. Ser.	######	Osgood's Horseshoe Bat	None.	China, Yunnan, N of Likiang, Nguluko, (27°05'N, 100°15'E).	Yunnan (China).	Not listed.	Least Concern	 macrotis species group. See Liu et al. (2019) and Csorba et al. (2003); but see Corbet and Hill (1992).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus osgoodi	23	Osgood's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	osgoodi	Sanborn	1939	0	Rhinolophus_osgoodi	Sanborn, C. C. (1939). Eight new bats of the genus Rhinolophus. Field Museum of Natural History Zoological Series, 24(5), 40.	https://www.biodiversitylibrary.org/item/20729#page/10/mode/1up	FMNH 33296		"Nguluko (25° 5' N. Lat., 100° 15' E. Long.), north of Likiang, Yunnan, China."	25.083	100.25	osgoodi Sanborn, 1939	NA	NA	China|Vietnam	Asia	Palearctic	LC	0	0	0	Rhinolophus_osgoodi	0	sciname match	Rhinolophus_osgoodi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19557	Rhinolophus osgoodi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	osgoodi	Sanborn, 1939	Rhinolophus osgoodi was considered to be conspecific with R. lepidus and belong to pusillus species group (Simmons 2005). Call frequency recorded for R. osgoodi (95.2 ;kHz) is close to that reported in Francis and Habersetzer (1998) for R. lepidus (98 ;kHz) in Malaysia. However, this species is different from the species from the pusillus species group in genetics (unpublished data) and some phenotypic characters, such as the higher anterior swellings and long palatal bridge, which suggested a different taxonomic position (unpublished data, Csorba pers. Comm.).	20000000	Rhinolophus osgoodi	Least Concern		2020	2018-09-04 00:00:00 UTC	3.1	English	Assessed as Least Concern because it occurs over a broad area. The global population is assumed to be larger and no declines have been observed.	Data on the species ecology and habitat use is limited. This species usually roosts in natural caves.	The threats to this species are not known.	The species global population is assumed to be stable.	Stable	This species has a wide distribution in Southern China, including the type locality, Nguluko, Yunnan (Sanborn 1939), and other localities in Yunnan, Guizhou and Hunan provinces (Sun et al. pers. comm.). It has been also discovered in Hoang Lien National Park of Vietnam.		Terrestrial	The species is known to be present in Hoang Lien National Park of Vietnam.Conservation and research actions are needed including protecting and managing known roosts, clarifying the species taxonomy, and research into the species population, life history, ecology and threats.	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		osgoodi	Sanborn	1939	0	Field Mus. Nat. Hist. Publ., Zool. Ser.	24:40:00	Osgood's Horseshoe Bat	None.	China, Yunnan, N of Likiang, Nguluko, (27°05'N, 100°15'E).	Yunnan (China).	Not listed.	Least Concern	 macrotis species group. See Liu et al. (2019) and Csorba et al. (2003); but see Corbet and Hill (1992).	Rhinolophus osgoodi	1004723	23	Osgood's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	osgoodi	Sanborn	1939	0	Rhinolophus_osgoodi	Sanborn, C. C. (1939). Eight new bats of the genus Rhinolophus. Field Museum of Natural History Zoological Series, 24(5), 40.	https://www.biodiversitylibrary.org/item/20729#page/10/mode/1up	FMNH 33296		"Nguluko (25° 5' N. Lat., 100° 15' E. Long.), north of Likiang, Yunnan, China."	25.08333	100.25	osgoodi Sanborn, 1939	NA	NA				China|Vietnam	Asia	Palearctic	LC	0	0	0	Rhinolophus_osgoodi	0	sciname match	Rhinolophus_osgoodi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_osgoodi	1004723	23	Osgood's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	osgoodi	Sanborn	0	Rhinolophus osgoodi	Sanborn, C.C. 1939-09-19. Eight new bats of the genus _Rhinolophus_. Field Museum of Natural History, Zoological Series 24(5):37-43.	https://www.biodiversitylibrary.org/page/2744785	FMNH:Mamm:33296	holotype	http://portal.vertnet.org/o/fmnh/mammals?id=76a671a2-aed5-4cb7-a048-534bb83d513a	"Nguluko (25° 5' N. Lat., 100° 15' E. Long.), north of Likiang, Yunnan, China."	27.08333	100.25	NA	NA				China|Vietnam	Asia	Palearctic	LC	0	0	0	Rhinolophus_osgoodi	0	sciname match	Rhinolophus_osgoodi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		osgoodi	Sanborn	1939	0	Field Mus. Nat. Hist. Publ., Zool. Ser.	24:40:00	Osgood's Horseshoe Bat	None.	China, Yunnan, N of Likiang, Nguluko, (27°05'N, 100°15'E).	Yunnan (China).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19557/21992735/' target='_blank'>Least Concern</a>	macrotisspecies group. See Liu et al. (2019) and Csorba et al. (2003); but see Corbet and Hill (1992).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus osgoodi; Rhinolophus osgoodi; Rhinolophus osgoodi; Rhinolophus osgoodi; Rhinolophus osgoodi; Rhinolophus osgoodi; osgoodi; Osgood’s Horseshoe Bat; Rhinolophe d'Osgood; Osgood-Hufeisennase; Herradura de Osgood; Osgood's Horseshoe Bat; Osgood's Horseshoe Bat; Osgood's Horseshoe Bat; R. osgoodi