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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1405	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Rhinolophus arcuates [synonym of]	Rhinolophus arcuates mcintyrei	Rhinolophus arcuates mcintyrei	Rhinolophus mcintyrei	Rhinolophus mcintyrei	Rhinolophus mcintyrei	Rhinolophus mcintyrei	Rhinolophus mcintyrei	Rhinolophus mcintyrei	Rhinolophus mcintyrei	Rhinolophus mcintyrei	Rhinolophus mcintyrei		[HMW] Rhinolophus arcuatus mcintyrei Hill & Schütter, 1982 , “ 4 km ENE Telefomin, W[est] Sepik Province , Papuan New Guinea ( 05°06’S , 141°41’ E ).” Rhinolophus mcintyrei is placed within the euryotis group and appears to be sister to a clade that includes R. euryotis , R. tatar , R. arcuatus , and R. inops or. subrufus , based on mitochondrial data. It was originally described as a subspecies of R. arcuatus and has typically been treated thus, but following a recent genetic study by L. E. Patrick and colleagues in 2013, the taxon has been elevated to species status. Monotypic.; [batnames2022] Formerly included in arcuatus but found to be distinct by Patrick et al. 2013.; [MDD2022] split from R. arcuates; [IUCN] <p><span lang="EN-US">Rhinolophus mcintyrei <span lang="EN-US"> <span lang="EN-US">(Hill and Schlitter 1982</span><span lang="EN-US">)<span lang="EN-US"> was originally described by those authors as a subspecies of Rhinolophus arcuatus Peters, 1871. Based on genetic analyses, Patrick and co-workers <span lang="EN-US">(2013<span lang="EN-US">)<span lang="EN-US"> ;suggested that <span lang="EN-US">the <span lang="EN-US">the population ascribed to that taxon</span><span lang="EN-US"> on Papua New Guinea be raised to species status rather than subspecies.<span lang="EN-US">  </span></span></span></span></p>; [batnames2023] Formerly included in arcuatus but found to be distinct by Patrick et al. 2013.; [MDD2023] split from R. arcuates; [MDD2025_2.0] split from R. arcuates; [batnames2025_1.7] Formerly included in arcuatus but found to be distinct by Patrick et al. 2013.; [MDD2025_2.2] split from R. arcuates														mcintyrei	<p><span lang="EN-US">Rhinolophus mcintyrei <span lang="EN-US"> <span lang="EN-US">(Hill and Schlitter 1982</span><span lang="EN-US">)<span lang="EN-US"> was originally described by those authors as a subspecies of Rhinolophus arcuatus Peters, 1871. Based on genetic analyses, Patrick and co-workers <span lang="EN-US">(2013<span lang="EN-US">)<span lang="EN-US"> ;suggested that <span lang="EN-US">the <span lang="EN-US">the population ascribed to that taxon</span><span lang="EN-US"> on Papua New Guinea be raised to species status rather than subspecies.<span lang="EN-US">  </span></span></span></span></p>			mcintyrei 	mcintyrei 			mcintyrei J. Edwards Hill & Schlitter, 1982						N/A																																								NA																											885887A2FFD48A35F8B7FBE7F962D7FC	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	317	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFE98A0EF894F5B7FE60DC11.xml	Rhinolophus mcintyrei	Rhinolophidae	Rhinolophus	mcintyrei	Hill & Schlitter	1982	Mc @en | ntyre’s Horseshoe Bat @en | Rhinolophe de Mc @fr | ntyre @en | Mclntyre-Hufeisennase @de | Herradura de Mc @es	Rhinolophus arcuatus mcintyrei Hill & Schütter, 1982 , “ 4 km ENE Telefomin, W[est] Sepik Province , Papuan New Guinea ( 05°06’S , 141°41’ E ).” Rhinolophus mcintyrei is placed within the euryotis group and appears to be sister to a clade that includes R. euryotis , R. tatar , R. arcuatus , and R. inops or. subrufus , based on mitochondrial data. It was originally described as a subspecies of R. arcuatus and has typically been treated thus, but following a recent genetic study by L. E. Patrick and colleagues in 2013, the taxon has been elevated to species status. Monotypic.	WC & C Papua New Guinea .	Head-body 48-64 mm , tail 16-23 mm , ear 15-22 mm , hindfoot 8-13 mm , forearm 47- 5-54 mm ; weight 9-15 g . McIntyre’s Horseshoe Bat is generally similar to the Bornean Woolly Horseshoe Bat ( R.proconsulis ), the Poso Horseshoe Bat (. belligeratar), and the Arcuate Horseshoe Bat (. arcuatus ) but is separated by its more expanded intemarial region, shorter and wider sella, and more inflated anterior median rostral swellings. Dorsal pelage is slightly reddish brown or dark brown (hairs whitish at base), whereas ventral pelage is paler. Ears are moderately long. Noseleaf has straightsided lancet with a pointed tip; connecting process is arcuate, forming a semicircle; sella is broadly ovate and wide basally, being slightly narrower than intemarial cup for much of its length before tapering slightly toward rounded tip; horseshoe is relatively wide (8-7-9- 7 mm ), nearly covers the muzzle, and has slight median emargination. Lower lip has three mental grooves. Skull is relatively slender; braincase is elongate and slightly inflated; interorbital region is abruptly narrowed; anterior median swellings are well developed and circular in outline; posterior swellings are less pronounced but still defined; rostral profile is distinctly concave in lateral view; sagittal crest is moderately developed; frontal depression is very shallow; supraorbital crests are slightly developed; canines are relatively slender and slightly curved (smaller and less massive than in the Bornean Woolly Horseshoe Bat). P2 is small and in the tooth row; P 3 is very small and completely extruded from the tooth row. On following pages: 78. New Guinea Broad-eared Horseshoe Bat ( Rhinolophus euryotis ); 79. Sulawesi Broad-eared Horseshoe Bat ( Rhinolophus tatah; 80. Poso Horseshoe Bat ( Rhinolophus belligerator ); 81. Bornean Woolly Horseshoe Bat ( Rhinolophus proconsulis ); 82. Arcuate Horseshoe Bat ( Rhinolophus arcuatus ); 83. Philippine Forest Horseshoe Bat ( Rhinolophus inops ); 84. Small Rufous Horseshoe Bat ( Rhinolophus subrufus ); 85. Large Rufous Horseshoe Bat ( Rhinolophus rufus ); 86. Intermediate Horseshoe Bat ( Rhinolophus affinis ); 87. Homfray's Horseshoe Bat ( Rhinolophus andamanensis ); 88. Croslet Horseshoe Bat ( Rhinolophus coelophyllus ); 89. Shamel's Horseshoe Bat ( Rhinolophus shameli ); 90. Vietnamese Brown Horseshoe Bat ( Rhinolophus microglobosus ); 91. Lesser Brown Horseshoe Bat ( Rhinolophus stheno ).	Typically found in lowland and hill forest habitats, such as primary lowland alluvial and hill forests that exhibit complex four-layered canopies. Recorded at elevations of 270-1600m .	McIntyre’s Horseshoe Bat probably forages using a mixture of aerial-hawking and gleaning; it is a slow, maneuverable flier.	No information.	McIntyre’s Horseshoe Bat is nocturnal and spends the day roosting in limestone caves and abandoned mines. Call shape is FM/CF/FM with a peak F recorded at 70—72 kHz.	McIntyre’s Horseshoe Bats are known to roost in small groups of at least a dozen.	Classified as Data Deficient on The IUCN Red List. Although there is only limited information available on this species’ ecology and threats, McIntyre’s Horseshoe Bat is likely to be threatened by deforestation and other forms of human disturbance, which are common in the lowland forests it inhabits.	Bonaccorso (1998) | Hill & Schütter (1982) | Leary & Pennay (2011) | Patrick & Ruedas (2017b) | Patrick et al. (2013)	https://zenodo.org/record/3750062/files/figure.png	77 . McIntyre’s Horseshoe Bat Rhinolophus mcintyrei French: Rhinolophe de McIntyre/ German: Mclntyre-Hufeisennase / Spanish: Herradura de McIntyre Taxonomy. Rhinolophus arcuatus mcintyrei Hill & Schütter, 1982 , “ 4 km ENE Telefomin, W[est] Sepik Province , Papuan New Guinea ( 05°06’S , 141°41’ E ).” Rhinolophus mcintyrei is placed within the euryotis group and appears to be sister to a clade that includes R. euryotis , R. tatar , R. arcuatus , and R. inops or. subrufus , based on mitochondrial data. It was originally described as a subspecies of R. arcuatus and has typically been treated thus, but following a recent genetic study by L. E. Patrick and colleagues in 2013, the taxon has been elevated to species status. Monotypic. Distribution. WC & C Papua New Guinea . Descriptive notes. Head-body 48-64 mm , tail 16-23 mm , ear 15-22 mm , hindfoot 8-13 mm , forearm 47- 5-54 mm ; weight 9-15 g . McIntyre’s Horseshoe Bat is generally similar to the Bornean Woolly Horseshoe Bat ( R.proconsulis ), the Poso Horseshoe Bat (. belligeratar), and the Arcuate Horseshoe Bat (. arcuatus ) but is separated by its more expanded intemarial region, shorter and wider sella, and more inflated anterior median rostral swellings. Dorsal pelage is slightly reddish brown or dark brown (hairs whitish at base), whereas ventral pelage is paler. Ears are moderately long. Noseleaf has straightsided lancet with a pointed tip; connecting process is arcuate, forming a semicircle; sella is broadly ovate and wide basally, being slightly narrower than intemarial cup for much of its length before tapering slightly toward rounded tip; horseshoe is relatively wide (8-7-9- 7 mm ), nearly covers the muzzle, and has slight median emargination. Lower lip has three mental grooves. Skull is relatively slender; braincase is elongate and slightly inflated; interorbital region is abruptly narrowed; anterior median swellings are well developed and circular in outline; posterior swellings are less pronounced but still defined; rostral profile is distinctly concave in lateral view; sagittal crest is moderately developed; frontal depression is very shallow; supraorbital crests are slightly developed; canines are relatively slender and slightly curved (smaller and less massive than in the Bornean Woolly Horseshoe Bat). P2 is small and in the tooth row; P 3 is very small and completely extruded from the tooth row. On following pages: 78. New Guinea Broad-eared Horseshoe Bat ( Rhinolophus euryotis ); 79. Sulawesi Broad-eared Horseshoe Bat ( Rhinolophus tatah; 80. Poso Horseshoe Bat ( Rhinolophus belligerator ); 81. Bornean Woolly Horseshoe Bat ( Rhinolophus proconsulis ); 82. Arcuate Horseshoe Bat ( Rhinolophus arcuatus ); 83. Philippine Forest Horseshoe Bat ( Rhinolophus inops ); 84. Small Rufous Horseshoe Bat ( Rhinolophus subrufus ); 85. Large Rufous Horseshoe Bat ( Rhinolophus rufus ); 86. Intermediate Horseshoe Bat ( Rhinolophus affinis ); 87. Homfray's Horseshoe Bat ( Rhinolophus andamanensis ); 88. Croslet Horseshoe Bat ( Rhinolophus coelophyllus ); 89. Shamel's Horseshoe Bat ( Rhinolophus shameli ); 90. Vietnamese Brown Horseshoe Bat ( Rhinolophus microglobosus ); 91. Lesser Brown Horseshoe Bat ( Rhinolophus stheno ). Habitat. Typically found in lowland and hill forest habitats, such as primary lowland alluvial and hill forests that exhibit complex four-layered canopies. Recorded at elevations of 270-1600m . Food and Feeding. McIntyre’s Horseshoe Bat probably forages using a mixture of aerial-hawking and gleaning; it is a slow, maneuverable flier. Breeding. No information. Activity patterns. McIntyre’s Horseshoe Bat is nocturnal and spends the day roosting in limestone caves and abandoned mines. Call shape is FM/CF/FM with a peak F recorded at 70—72 kHz. Movements, Home range and Social organization. McIntyre’s Horseshoe Bats are known to roost in small groups of at least a dozen. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Although there is only limited information available on this species’ ecology and threats, McIntyre’s Horseshoe Bat is likely to be threatened by deforestation and other forms of human disturbance, which are common in the lowland forests it inhabits. Bibliography. Bonaccorso (1998), Hill & Schütter (1982), Leary & Pennay (2011), Patrick & Ruedas (2017b), Patrick et al. (2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus mcintyrei	Rhinolophus		mcintyrei	Hill & Schlitter	1982	0	Ann. Carnegie Mus.	58:39:00	McIntyre's Horseshoe Bat	None.	Papua New Guinea, W Sepik Province, 4 km ENE Telefomin	New Guinea	Not listed.	Data Deficient	Formerly included in arcuatus but found to be distinct by Patrick et al. 2013.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus mcintyrei	23	McIntyre's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	mcintyrei	J. Edwards Hill & Schlitter	1982	0	Rhinolophus_arcuatus_mcintyrei	Hill, J. E. & Schiltter, D. A. (1982). A record of Rhinolophus arcuatus (Chiroptera: Rhinolophidae) from New Guinea, with the description of a new subspecies. Annals of the Carnegie Museum, 51, 459.	https://www.biodiversitylibrary.org/page/52374483#page/503/mode/1up	CM 63497		"4 km ENE Telefomin, W[est] Sepik Province, Papuan New Guinea (05°06'S, 141°41'E)."	-5.1	141.68	mcintyrei J. Edwards Hill & Schlitter, 1982	split from R. arcuates	Patrick, L. E., McCulloch, E. S., & Ruedas, L. A. (2013). Systematics and biogeography of the arcuate horseshoe bat species complex (Chiroptera, Rhinolophidae). Zoologica Scripta, 42(6), 553-590.	Papua New Guinea	Oceania	Australasia/Oceania	DD	0	0	0	Rhinolophus_mcintyrei	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Rhinolophus mcintyrei	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	mcintyrei	Hill &; Schlitter 1982	<p><span lang="EN-US">Rhinolophus mcintyrei <span lang="EN-US"> <span lang="EN-US">(Hill and Schlitter 1982</span><span lang="EN-US">)<span lang="EN-US"> was originally described by those authors as a subspecies of Rhinolophus arcuatus Peters, 1871. Based on genetic analyses, Patrick and co-workers <span lang="EN-US">(2013<span lang="EN-US">)<span lang="EN-US"> ;suggested that <span lang="EN-US">the <span lang="EN-US">the population ascribed to that taxon</span><span lang="EN-US"> on Papua New Guinea be raised to species status rather than subspecies.<span lang="EN-US">  </span></span></span></span></p>	80000000	Rhinolophus mcintyrei	Data Deficient		2017	2016-06-25 00:00:00 UTC	3.1	English	<span lang="EN-US">Little is currently known about this species’ population size, distribution or threats, therefore it is listed as Data Deficient (DD).	<p><span lang="EN-US">This species roosts in limestone caves, in groups with other members of its species <span lang="EN-US">(Bonaccorso 1998)<span lang="EN-US">. It is a gleaning insectivore capable of slow, manoeuvrable flight. Most specimens have been captured in the caves in which they roost <span lang="EN-US">(Flannery and Seri 1990,<span lang="EN-US"> Hill and Schlitter 1982<span lang="EN-US">)<span lang="EN-US">, however, additional individuals have been captured with mist nets in primary lowland alluvial and hill forests exhibiting complex, four layer canopies <span lang="EN-US">(Leary and Pennay 2011<span lang="EN-US">)<span lang="EN-US">.</span></span></span></span></span></p>	<p><span lang="EN-US">Given the general dearth of information for this species, specific threats are unknown at this time. However, based on its lowland forest habitat, its roosting requirements, and foraging habits, deforestation and human disturbance of caves could be very real threats to the on-going survival of this species and should be minimized <span lang="EN-US">(Wiles and Brooke 2009)<span lang="EN-US">.</span></p>	<p><span lang="EN-US">Currently, this species is known from approximately twelve localities. Population sizes and trends are unknown.</p>	Unknown	<p><span lang="EN-US">This species is currently known from the provinces of Eastern Highlands, Sandaun, Southern Highlands, and Western <span lang="EN-US">(Bonaccorso 1998)<span lang="EN-US">. The elevation of known localities range from 270 to 1,600 m asl ;<span lang="EN-US">(Bonaccorso 1998, Leary and Pennay 2011).<span lang="EN-US"></span></span></p>		Terrestrial	No conservation actions are known to be in place for this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		mcintyrei	Hill & Schlitter	1982	0	Ann. Carnegie Mus.	58:39:00	McIntyre's Horseshoe Bat	None.	Papua New Guinea, W Sepik Province, 4 km ENE Telefomin	New Guinea	Not listed.	Data Deficient	Formerly included in arcuatus but found to be distinct by Patrick et al. 2013.	Rhinolophus mcintyrei	1004711	23	McIntyre's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	mcintyrei	J. Edwards Hill & Schlitter	1982	0	Rhinolophus_arcuatus_mcintyrei	Hill, J. E. & Schiltter, D. A. (1982). A record of Rhinolophus arcuatus (Chiroptera: Rhinolophidae) from New Guinea, with the description of a new subspecies. Annals of the Carnegie Museum, 51, 459.	https://www.biodiversitylibrary.org/page/52374483#page/503/mode/1up	CM 63497		"4 km ENE Telefomin, W[est] Sepik Province, Papuan New Guinea (05°06'S, 141°41'E)."	-5.1	141.6833	mcintyrei J. Edwards Hill & Schlitter, 1982	split from R. arcuates	Patrick, L. E., McCulloch, E. S., & Ruedas, L. A. (2013). Systematics and biogeography of the arcuate horseshoe bat species complex (Chiroptera, Rhinolophidae). Zoologica Scripta, 42(6), 553-590.				Papua New Guinea	Oceania	Australasia/Oceania	DD	0	0	0	Rhinolophus_mcintyrei	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_mcintyrei	1004711	23	McIntyre's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	mcintyrei	J. Edwards Hill & Schlitter	0	Rhinolophus arcuatus mcintyrei	Hill, J.E. and Schlitter, D.A. 1982-12-03. A record of _Rhinolophus arcuatus_ (Chiroptera: Rhinolophidae) from New Guinea, with the description of a new subspecies. Annals of Carnegie Museum 51(23):455-464.	https://www.biodiversitylibrary.org/page/52374487	CM 63497	holotype		"4 km ENE Telefomin, W[est] Sepik Province, Papuan New Guinea (05°06'S, 141°41'E)."	-5.1	141.6833	split from R. arcuates	Patrick, L. E., McCulloch, E. S., & Ruedas, L. A. (2013). Systematics and biogeography of the arcuate horseshoe bat species complex (Chiroptera, Rhinolophidae). Zoologica Scripta, 42(6), 553-590.				Papua New Guinea	Oceania (Continent)	Australasia	DD	0	0	0	Rhinolophus_mcintyrei	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		mcintyrei	Hill & Schlitter	1982	0	Ann. Carnegie Mus.	58:39:00	McIntyre's Horseshoe Bat	None.	Papua New Guinea, W Sepik Province, 4 km ENE Telefomin	New Guinea	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84372245/84372277/' target='_blank'>Data Deficient</a>	Formerly included in arcuatus but found to be distinct by Patrick et al. 2013.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus mcintyrei; Rhinolophus mcintyrei; Rhinolophus mcintyrei; Rhinolophus mcintyrei; Rhinolophus mcintyrei; mcintyrei; Mc; ntyre’s Horseshoe Bat; Rhinolophe de Mc; ntyre; Mclntyre-Hufeisennase; Herradura de Mc; McIntyre's Horseshoe Bat; McIntyre's Horseshoe Bat; R. mcintyrei