http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1404	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli	Rhinolophus marshalli		[MSW3] philippinensis species group. Reviewed by Yoshiyuki (1990) and Hill and Topál (1990); also see Bates et al. (2001) and Hendrichsen et al. (2001b).; [HMW] Rhinolophus marshalli Thonglongya, 1973 , “ foothills of Khao Soi Dao Tai, Amphoe Pong Nam Ron, Chanthaburi , about lat. 12 ° 55 ’ N ., long. 102 ° 10 ’E ., ” Thailand . Rhinolophus marshalli is the most basal member of the macrotis species group. Monotypic.; [batnames2022]  philippinensis species group. Reviewed by Yoshiyuki (1990) and Hill and Topál (1990); also see Bates et al. (2001) and Hendrichsen et al. (2001b). May contain cryptic diversity; see Chornelia et al. (2022).; [batnames2023]  philippinensis species group. Reviewed by Yoshiyuki (1990) and Hill and Topál (1990); also see Bates et al. (2001) and Hendrichsen et al. (2001b). May contain cryptic diversity; see Chornelia et al. (2022).; [batnames2025_1.7] philippinensis species group. Reviewed by Yoshiyuki (1990) and Hill and Topál (1990); also see Bates et al. (2001) and Hendrichsen et al. (2001b). May contain cryptic diversity; see Chornelia et al. (2022).														marshalli				marshalli 	marshalli 			marshalli Thonglongya, 1973		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Marshall's horseshoe bat	Thailand; ref. 4.57	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinolophus marshalli	Thailand, Chantaburi, Amphoe Pong Nam Ron, foothills of Khao Soi Dao Thai.	Thonglongya	1973	Mammalia, 37:590.	Distribution: Known only from Thailand.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Marshall's horseshoe bat	Thailand, Vietnam, N Malaya; ref. 4.147	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thonglongya	1973	Mammalia, 37:590.		Thailand.	Thailand, Chantaburi, Amphoe Pong Nam Ron, foothills of Khao Soi Dao Thai.		THONGLONGYA	1973	Sella broad and very short with rudi mentary lappets; internarial cup very broad, trap ezoid rather than subcircular and not enclosing base of sella proper. Lancet fairly low but triangu lar. Infraorbital canal short. Median rostral swel lings of skull unusually enlarged. Size medium (forearm length, 45-46 mm).	Distribution: Known only from Thailand.	No subspecies.		58	species	R. marshalli	THONGLONGYA	1973	Rhinolophus	genus	Rhinolophus marshalli				Sella broad and very short with rudi mentary lappets; internarial cup very broad, trap ezoid rather than subcircular and not enclosing base of sella proper. Lancet fairly low but triangu lar. Infraorbital canal short. Median rostral swel lings of skull unusually enlarged. Size medium (forearm length, 45-46 mm).	No subspecies.		49. R. marshalli THONGLONGYA 1973 [luctus group].	49	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus marshalli	Rhinolophus		marshalli	Thonglongya		1973		Mammalia	37		590		Marshall's Horseshoe Bat	Thailand, Chantaburi, Amphoe Pong Nam Ron, foothills of Khao Soi Dao Thai.	Thailand, Burma, Vietnam, Laos, Peninsular Malaysia.	IUCN 2003 – and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		philippinensis species group. Reviewed by Yoshiyuki (1990) and Hill and Topál (1990); also see Bates et al. (2001) and Hendrichsen et al. (2001b).	885887A2FFE98A0EF894F5B7FE60DC11	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	303	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFD78A31FF57FE9AF6E0DFB1.xml	Rhinolophus marshalli	Rhinolophidae	Rhinolophus	marshalli	Thonglongya	1973	Marshall’s Horseshoe Bat @en | Rhinolophe de Marshall @fr | Marshall-Hufeisennase @de | Herradura de Marshall @es	Rhinolophus marshalli Thonglongya, 1973 , “ foothills of Khao Soi Dao Tai, Amphoe Pong Nam Ron, Chanthaburi , about lat. 12 ° 55 ’ N ., long. 102 ° 10 ’E ., ” Thailand . Rhinolophus marshalli is the most basal member of the macrotis species group. Monotypic.	SE China ( S Yunnan and S Guangxi ), E Myanmar , N, C & extreme S Thailand , N Laos , N Vietnam , and N Peninsular Malaysia .	Head-body 43-49 mm , tail 16-26 mm , ear 25-30 mm , hindfoot 7-3-8- 1 mm , forearm 38- 9-48 mm ; weight 5-8 g . Dorsal pelage is dark brown, occasionally with reddish tinge; venter is smoky gray or light yellowish brown. No orange morph is known. Males lack axillary tufts. Ears are enormous, with pronounced antitragus. Noseleaf has very short, fleshy lancet, with broadly rounded tip; connecting process is low, rounded, andjoins lancet at base; sella is large, broad, and barely higher than wide, with semicircular upper margin; internarial region is largely expanded laterally, with wing-like structures on edges that are integral with base of sella; and horseshoe is broad at 7-5-9- 2 mm , covers most of upper lip, extends laterally few millimeters beyond muzzle, lacks lateral leaflets, and has deep, narrow median emargination. Lower lip has three medial grooves. Baculum is prominently bent upward, with small basal cone and deep notches; relatively wide ventral notch is deeper than dorsal notch; dorsal knob is low, and ventral depression is small in extension; tip is narrowly rounded with lateral widening and longish dorsal knob. Skull is slender, with long rostrum (zygomatic width is less than mastoid width); median swellings are conspicuously inflated; posterior compartments are underdeveloped and rudimentary; sagittal crest is very low across skull; frontal depression is moderately deep; and supraorbital crests are strong, with sharp ridges. P2 is small to medium in size and in tooth row, separating C1 and P4, and P3 is small and either in tooth row or halfway displaced labially, separating P 2 and P4.	Various forest habitats, including dry deciduous forests, bamboo forests, and lowland moist forests, from sea level to elevations of c . 800 m . Marshall’s Horseshoe Bats tolerate very disturbed habitats.	No information.	Births of Marshall’s Horseshoe Bats occur from late April until early May. Litter size is one. Postnatal growth occurs fast, with mean growth rates of 0-17 g /day until 13 days of age, decreasing gradually until adult size is reached at c .40 days old.	Marshall’s Horseshoe Bat is nocturnal. It roosts primarily in limestone caves but also in rock crevices. Call shape is FM /CF/FM, with peak call frequencies of 43-44-5 kHz in Guangxi , 41-8—42-3 kHz in Yunnan , China , and 44 kHz in Vietnam .	No information.	Classified as Least Concern on The IUCNed List. Marshall’s Horseshoe Bat is considerably uncommon but widespread and resilient in disturbed habitats. Nevertheless, it might be threatened by habitat destruction and roost disturbance from cave tourism.	Bates, Francis & Csorba (2008a) | Csorba et al. (2003) | Francis (2008a) | Jin Longru, Bo Luo et al. (2012) | Liu Ying, Jiang Tinglei et al. (2009) | Thonglongya (1973) | Tu Vuong Tan, Hassanin et al. (2017) | Wu Hui , Yang Qisen et al. (2015) | WuYi et al. (2004) | Zhang Libiao , Jones et al. (2009) | Zhang Libiao , Long Yongcheng et al. (2005) | Zhang Lin et al. (2018)	https://zenodo.org/record/3750000/files/figure.png	48 . Marshall’s Horseshoe Bat Rhinolophus marshalli French: Rhinolophe de Marshall / German : Marshall-Hufeisennase / Spanish : Herradura de Marshall Taxonomy. Rhinolophus marshalli Thonglongya, 1973 , “ foothills of Khao Soi Dao Tai, Amphoe Pong Nam Ron, Chanthaburi , about lat. 12 ° 55 ’ N ., long. 102 ° 10 ’E ., ” Thailand . Rhinolophus marshalli is the most basal member of the macrotis species group. Monotypic. Distribution. SE China ( S Yunnan and S Guangxi ), E Myanmar , N, C & extreme S Thailand , N Laos , N Vietnam , and N Peninsular Malaysia . Descriptive notes. Head-body 43-49 mm , tail 16-26 mm , ear 25-30 mm , hindfoot 7-3-8- 1 mm , forearm 38- 9-48 mm ; weight 5-8 g . Dorsal pelage is dark brown, occasionally with reddish tinge; venter is smoky gray or light yellowish brown. No orange morph is known. Males lack axillary tufts. Ears are enormous, with pronounced antitragus. Noseleaf has very short, fleshy lancet, with broadly rounded tip; connecting process is low, rounded, andjoins lancet at base; sella is large, broad, and barely higher than wide, with semicircular upper margin; internarial region is largely expanded laterally, with wing-like structures on edges that are integral with base of sella; and horseshoe is broad at 7-5-9- 2 mm , covers most of upper lip, extends laterally few millimeters beyond muzzle, lacks lateral leaflets, and has deep, narrow median emargination. Lower lip has three medial grooves. Baculum is prominently bent upward, with small basal cone and deep notches; relatively wide ventral notch is deeper than dorsal notch; dorsal knob is low, and ventral depression is small in extension; tip is narrowly rounded with lateral widening and longish dorsal knob. Skull is slender, with long rostrum (zygomatic width is less than mastoid width); median swellings are conspicuously inflated; posterior compartments are underdeveloped and rudimentary; sagittal crest is very low across skull; frontal depression is moderately deep; and supraorbital crests are strong, with sharp ridges. P2 is small to medium in size and in tooth row, separating C1 and P4, and P3 is small and either in tooth row or halfway displaced labially, separating P 2 and P4. Habitat. Various forest habitats, including dry deciduous forests, bamboo forests, and lowland moist forests, from sea level to elevations of c . 800 m . Marshall’s Horseshoe Bats tolerate very disturbed habitats. Food and Feeding. No information. Breeding. Births of Marshall’s Horseshoe Bats occur from late April until early May. Litter size is one. Postnatal growth occurs fast, with mean growth rates of 0-17 g /day until 13 days of age, decreasing gradually until adult size is reached at c .40 days old. Activity patterns. Marshall’s Horseshoe Bat is nocturnal. It roosts primarily in limestone caves but also in rock crevices. Call shape is FM /CF/FM, with peak call frequencies of 43-44-5 kHz in Guangxi , 41-8—42-3 kHz in Yunnan , China , and 44 kHz in Vietnam . Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCNed List. Marshall’s Horseshoe Bat is considerably uncommon but widespread and resilient in disturbed habitats. Nevertheless, it might be threatened by habitat destruction and roost disturbance from cave tourism. Bibliography. Bates, Francis & Csorba (2008a), Csorba eta/. (2003), Francis (2008a), Jin Longru, Bo Luo et al. (2012), Liu Ying, Jiang Tinglei et al. (2009), Thonglongya (1973), Tu Vuong Tan, Hassanin et al. (2017), Wu Hui , Yang Qisen et al. (2015), WuYi et al. (2004), Zhang Libiao , Jones et al. (2009), Zhang Libiao , Long Yongcheng et al. (2005), Zhang Lin et al. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus marshalli	Rhinolophus		marshalli	Thonglongya	1973	0	Mammalia	46:50:00	Marshall's Horseshoe Bat	None.	Thailand, Chantaburi, Amphoe Pong Nam Ron, foothills of Khao Soi Dao Thai.	Yunnan Province China, Thailand, Burma, Vietnam, Laos, Peninsular Malaysia.	Not listed.	Least Concern	 philippinensis species group. Reviewed by Yoshiyuki (1990) and Hill and Topál (1990); also see Bates et al. (2001) and Hendrichsen et al. (2001b). May contain cryptic diversity; see Chornelia et al. (2022).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus marshalli	23	Marshall's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	marshalli	Thonglongya	1973	0	Rhinolophus_marshalli	Thonglongya, K. (1973). First record of Rhinolophus paradoxolophus (Bourret, 1951) from Thailand, with the description of a new species of the Rhinolophus philippinensis group (Chiroptera, Rhinolophidae). Mammalia, 37(4), 590.	https://www.degruyter.com/document/doi/10.1515/mamm.1973.37.4.587/html			"foothills of Khao Soi Dao Tai, Amphoe Pong Nam Ron, Chanthaburi, about lat. 12°55'N., long. 102°10'E.," Thailand.	12.917	102.17	marshalli Thonglongya, 1973	NA	NA	China|Myanmar|Thailand|Laos|Vietnam|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_marshalli	0	sciname match	Rhinolophus_marshalli	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19552	Rhinolophus marshalli	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	marshalli	Thonglongya, 1973		20000000	Rhinolophus marshalli	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern as although never abundant, it is a widespread species and it seems able to survive in disturbed areas.	The species is a cave dweller. In Myanmar and Viet Nam it is associated with limestone caves and is tolerant of very disturbed habitat (P. Bates pers. comm.). In Thailand it is also found in rock crevices (Bates et al. 2008). The species forages under canopies of vegetation ranging from plantation to primary forests. Two individuals were collected in a garden in which was growing litchi, jack-fruit, banana and other vegetation in Cat Ba island (Thong 2012).	Cave disturbance appears as a major threat to the species. Results from annual surveys and monitoring between 2006 and 2017 in Cat Ba island, northern Vietnam indicated that its colonies are likely declined due to cave visiting and clearance of the foraging habitats.	The largest colony up to ca. 30 individuals of the species was recorded in Cat Ba island, northern Vietnam (Thong et al. unpublished data) although it has usually been found as individuals or small groups, never in large numbers in the remaining distributional range (Bates et al. 2008). It is not clear whether it has declined significantly historically (Bates et al. 2008). Annual surveys and monitoring activities in Cat Ba island indicated that the known colonies are likely to have declined between 2008 and 2018.	Unknown	This species occurs in Myanmar, China (Guangxi, Guizhou, Yunnan), Thailand (Chantaburi, Amphoe Pong Nam Ron, foothills of Khao Soi Dao Thai), Lao PDR, Viet Nam (from northern to south-central regions), and Peninsular Malaysia.		Terrestrial	The species is still not listed as protected in Vietnam. Its roosting sites are found in different areas including the protected and unprotected ones. All known populations are in protected areas in Thailand (Bates et al. 2008).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		marshalli	Thonglongya	1973	0	Mammalia	46:50:00	Marshall's Horseshoe Bat	None.	Thailand, Chantaburi, Amphoe Pong Nam Ron, foothills of Khao Soi Dao Thai.	Yunnan Province China, Thailand, Burma, Vietnam, Laos, Peninsular Malaysia.	Not listed.	Least Concern	 philippinensis species group. Reviewed by Yoshiyuki (1990) and Hill and Topál (1990); also see Bates et al. (2001) and Hendrichsen et al. (2001b). May contain cryptic diversity; see Chornelia et al. (2022).	Rhinolophus marshalli	1004710	23	Marshall's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	marshalli	Thonglongya	1973	0	Rhinolophus_marshalli	Thonglongya, K. (1973). First record of Rhinolophus paradoxolophus (Bourret, 1951) from Thailand, with the description of a new species of the Rhinolophus philippinensis group (Chiroptera, Rhinolophidae). Mammalia, 37(4), 590.	https://www.degruyter.com/document/doi/10.1515/mamm.1973.37.4.587/html			"foothills of Khao Soi Dao Tai, Amphoe Pong Nam Ron, Chanthaburi, about lat. 12°55'N., long. 102°10'E.," Thailand.	12.91667	102.1667	marshalli Thonglongya, 1973	NA	NA				China|Myanmar|Thailand|Laos|Vietnam|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_marshalli	0	sciname match	Rhinolophus_marshalli	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_marshalli	1004710	23	Marshall's Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	marshalli	Thonglongya	0	Rhinolophus marshalli	Thonglongya, K. 1973. First record of _Rhinolophus paradoxolophus_ (Bourret, 1951) from Thailand, with the description of a new species of the _Rhinolophus philippinensis_ group (Chiroptera, Rhinolophidae). Mammalia 37(4):587-597.	https://doi.org/10.1515/mamm.1973.37.4.587	TNRC 54-1669	holotype		"foothills of Khao Soi Dao Tai, Amphoe Pong Nam Ron, Chanthaburi, about lat. 12°55'N., long. 102°10'E.," Thailand.	12.91667	102.1667	NA	NA				China|Myanmar|Thailand|Laos|Vietnam|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Rhinolophus_marshalli	0	sciname match	Rhinolophus_marshalli	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		marshalli	Thonglongya	1973	0	Mammalia	46:50:00	Marshall's Horseshoe Bat	None.	Thailand, Chantaburi, Amphoe Pong Nam Ron, foothills of Khao Soi Dao Thai.	Yunnan Province China, Thailand, Burma, Vietnam, Laos, Peninsular Malaysia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19552/21978274/' target='_blank'>Least Concern</a>	philippinensis species group. Reviewed by Yoshiyuki (1990) and Hill and Topál (1990); also see Bates et al. (2001) and Hendrichsen et al. (2001b). May contain cryptic diversity; see Chornelia et al. (2022).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus marshalli; Rhinolophus marshalli; Rhinolophus marshalli; Rhinolophus marshalli; Rhinolophus marshalli; Rhinolophus marshalli; marshalli; Marshall’s Horseshoe Bat; Rhinolophe de Marshall; Marshall-Hufeisennase; Herradura de Marshall; Marshall's Horseshoe Bat; Marshall's Horseshoe Bat; Marshall's Horseshoe Bat; R. marshalli