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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1403	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus	Rhinolophus malayanus		[MSW3] megaphyllus species group. McFarlane and Blood (1986) suggested that characters used by Lekagul and McNeely (1977) to separate stheno and malayanus may not be reliable, but see Corbet and Hill (1992), Csorba and Jenkins (1998), and Hendrichsen (2001a, b).; [HMW] Rhinolophus malayanus Bonhote in Annandale & H.. Robinson, 1903, “ Biserat,Jalor, ” Thailand . Tentatively included in the megaphyllus group but its placement here is uncertain; further genetic work is needed. Rhinolophus malayanus is generally considered to be close to. stheno , but they do not cluster together in genetic studies. Exact placement of. malayanu s in relation to other Rhinolophus is still uncertain but the most recent genetic study placed it as sister to R. acuminatus , forming a clade sister to much of the megaphyllus , pusillus , philippinensis , and macrotis groups. Monotypic.; [batnames2022]  megaphyllus species group. McFarlane and Blood (1986) suggested that characters used by Lekagul and McNeely (1977) to separate stheno and malayanus may not be reliable, but see Corbet and Hill (1992), Csorba and Jenkins (1998), and Hendrichsen (2001a, b). May contain cryptic diversity; see Chornelia et al. (2022).; [batnames2023]  megaphyllus species group. McFarlane and Blood (1986) suggested that characters used by Lekagul and McNeely (1977) to separate stheno and malayanus may not be reliable, but see Corbet and Hill (1992), Csorba and Jenkins (1998), and Hendrichsen (2001a, b). May contain cryptic diversity; see Chornelia et al. (2022).; [batnames2025_1.7] megaphyllus species group. McFarlane and Blood (1986) suggested that characters used by Lekagul and McNeely (1977) to separate stheno and malayanus may not be reliable, but see Corbet and Hill (1992), Csorba and Jenkins (1998), and Hendrichsen (2001a, b). May contain cryptic diversity; see Chornelia et al. (2022).														malayanus				malayanus	malayanus			malayanus Bonhote, 1903		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	N Malayan horseshoe bat	Thailand – Vietnam, Malaya	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Rhinolophus malayanus	Thailand, Patani, Biserat.	Bonhote	1903	Fasc. Malayenses Zool., 1:15.	Distribution: Ranging from northern Vietnam and Laos through Thailand to Malaya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	N Malayan horseshoe bat	Thailand – Vietnam, Malaya	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Bonhote	1903	In N. Annandale, Fasciculi Malayenses, ZooL, 1:15.		W Malaysia, Thailand, Laos, Vietnam.	Thailand, Patani, Biserat.		BOHOTE	1903	Median anterior nasal swellings definite ly larger than lateral anterior nasal swellings. Size relatively small (forearm length, 40-43 mm; total length of skull, 18-19 mm.	Distribution: Ranging from northern Vietnam and Laos through Thailand to Malaya.	No subspecies.		52	species	R. malayanus	BOHOTE	1903	Rhinolophus	genus	Rhinolophus malayanus				Median anterior nasal swellings definite ly larger than lateral anterior nasal swellings. Size relatively small (forearm length, 40-43 mm; total length of skull, 18-19 mm.	No subspecies.		8. R. malayanus BOHOTE 1903 [ferrumequinum group].	8	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus malayanus	Rhinolophus		malayanus	Bonhote		1903		In N. Annandale, Fasciculi Malayenses, Zool.	1		15		Malayan Horseshoe Bat	Thailand, Jalor, Biserat.	Thailand, Burma, Cambodia, Laos, Vietnam, Peninsular Malaysia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		megaphyllus species group. McFarlane and Blood (1986) suggested that characters used by Lekagul and McNeely (1977) to separate stheno and malayanus may not be reliable, but see Corbet and Hill (1992), Csorba and Jenkins (1998), and Hendrichsen (2001a, b).	885887A2FFD78A31FF57FE9AF6E0DFB1	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	310	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFD08A36FF00FCFEF899D778.xml	Rhinolophus malayanus	Rhinolophidae	Rhinolophus	malayanus	Bonhote in Annandale	1903	Malayan Horseshoe Bat @en | Rhinolophe de Malaisie @fr | Malaya-Hufeisennase @de | Herradura de Malasia @es	Rhinolophus malayanus Bonhote in Annandale & H.. Robinson, 1903, “ Biserat,Jalor, ” Thailand . Tentatively included in the megaphyllus group but its placement here is uncertain; further genetic work is needed. Rhinolophus malayanus is generally considered to be close to. stheno , but they do not cluster together in genetic studies. Exact placement of. malayanu s in relation to other Rhinolophus is still uncertain but the most recent genetic study placed it as sister to R. acuminatus , forming a clade sister to much of the megaphyllus , pusillus , philippinensis , and macrotis groups. Monotypic.	Myanmar , Thailand , Laos , Vietnam , Cambodia , and N Peninsular Malaysia along with Tarutao and Langkawi Is.			No information.	Pregnant females were captured in late February in Cambodia .	The Malayan Horseshoe Bat is nocturnal, roosting in limestone caves during the day. Torpid individuals have been captured in day roosts during July, in western Thailand . Call shape is FM/CF/FM with peak F recorded at 72—80 kHz in northern Thailand , 82—86 in central Thailand , 83—91 kHz in southern Thailand , 75— 82 kHz in Cambodia , 76-79 kHz in Myanmar , and 77—80 kHz in Laos .	Malayan Horseshoe Bats roost in colonies, with up to 3000 individuals recorded in one cave in western Thailand .	Classified as Least Concern on The IUCN Red List. There do not seem to be any major threats currently affecting this widespread species.	Bates, Bumrungsri & Csorba (2008b) | Csorba et al. (2003) | Francis (2008a) | Harada, Minezawa et al. (1982) | Kruskop (2013a) | Soisook (2008) | Soisook et al. (2008) | Stoffberg et al. (2010) | Zhang Lin et al. (2018)	https://zenodo.org/record/3750028/files/figure.png	61 . Malayan Horseshoe Bat Rhinolophus malayanus French: Rhinolophe de Malaisie / German: Malaya-Hufeisennase / Spanish: Herradura de Malasia Taxonomy. Rhinolophus malayanus Bonhote in Annandale & H.. Robinson, 1903, “ Biserat,Jalor, ” Thailand . Tentatively included in the megaphyllus group but its placement here is uncertain; further genetic work is needed. Rhinolophus malayanus is generally considered to be close to. stheno , but they do not cluster together in genetic studies. Exact placement of. malayanu s in relation to other Rhinolophus is still uncertain but the most recent genetic study placed it as sister to R. acuminatus , forming a clade sister to much of the megaphyllus , pusillus , philippinensis , and macrotis groups. Monotypic. Distribution. Myanmar , Thailand , Laos , Vietnam , Cambodia , and N Peninsular Malaysia along with Tarutao and Langkawi Is. Descriptive notes. Head-body 36-1-55- 8 mm , tail 15-26- 9 mm , ear 12-7-19- 3 mm , hindfoot 5-8—9- 6 mm , forearm 38- 2—44 mm ; weight 4-5—9- 7 g . Dorsal pelage is either brown (brown morph) or bright cinnamon (orange morph), whereas ventral pelage is either buff or whitish. Ears are of medium size. Noseleaf has triangular lancet with concave or nearly parallel lateral margins and elongated tip; connecting process is rounded; sella is relatively wide with broadly rounded top, and lateral margins are almost parallel; horseshoe is relatively broad (6- 6-9 mm ) and nearly completely covers muzzle, has moderately developed lateral leaflets, and a shallow but present median emargination. Lower lip has three mental grooves. Skull is moderately robust and zygomatic width is only slightly larger than mastoid width; anterior median swellings are well developed and wider than they are long; posterior compartments are well inflated; sagittal crest is rather weak; frontal depression is relatively elongate but shallow; supraorbital crests are more or less distinct P2 is of medium size, with distinct cusp, and is within tooth row, separating C1 and P4; P 3 is small and slightly to almost completely extrudes labially from tooth row, which allows P2 and P4 to touch. Chromosomal complement has 2n = 62 and FNa = 60. Habitat Mainly recorded from areas with limestone karst and sandstone mountains for roosting. Malayan Horseshoe Bats have been recorded in lowland mixed deciduous forest, hill evergreen forest, highly disturbed dry dipterocarp forest, and agricultural areas, indicating that the species is resilient in response to disturbance. It is found at elevations from sea level up to c. 1400 m . Food and Feeding. No information. Breeding. Pregnant females were captured in late February in Cambodia . Activity patterns. The Malayan Horseshoe Bat is nocturnal, roosting in limestone caves during the day. Torpid individuals have been captured in day roosts during July, in western Thailand . Call shape is FM/CF/FM with peak F recorded at 72—80 kHz in northern Thailand , 82—86 in central Thailand , 83—91 kHz in southern Thailand , 75— 82 kHz in Cambodia , 76-79 kHz in Myanmar , and 77—80 kHz in Laos . Movements, Home range and Social organization. Malayan Horseshoe Bats roost in colonies, with up to 3000 individuals recorded in one cave in western Thailand . Status and Conservation. Classified as Least Concern on The IUCN Red List. There do not seem to be any major threats currently affecting this widespread species. Bibliography. Bates, Bumrungsri & Csorba (2008b), Csorba eta/. (2003), Francis (2008a), Harada, Minezawa et al. (1982), Kruskop (2013a), Soisook (2008), Soisook et al. (2008), Stoffberg et al. (2010), Zhang Lin et al. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus malayanus	Rhinolophus		malayanus	Bonhote	1903	0	In N. Annandale, Fasciculi Malayenses, Zool.	1:15	Malayan Horseshoe Bat	None.	Thailand, Jalor, Biserat.	Yunnan (China), Thailand, Burma, Cambodia, Laos, Vietnam, Peninsular Malaysia.	Not listed.	Least Concern	 megaphyllus species group. McFarlane and Blood (1986) suggested that characters used by Lekagul and McNeely (1977) to separate stheno and malayanus may not be reliable, but see Corbet and Hill (1992), Csorba and Jenkins (1998), and Hendrichsen (2001a, b). May contain cryptic diversity; see Chornelia et al. (2022).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus malayanus	23	Malayan Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	malayanus	Bonhote	1903	0						"Biserat, Jalor," Thailand.			malayanus Bonhote, 1903	NA	NA	China|Myanmar|Thailand|Laos|Vietnam|Cambodia|Malaysia	Asia	Indomalaya	LC	0	0	0	Rhinolophus_malayanus	0	sciname match	Rhinolophus_malayanus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	19551	Rhinolophus malayanus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	malayanus	Bonhote, 1903		20000000	Rhinolophus malayanus	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Confirmed as Least Concern because it is widespread and common, there are no major threats, and there is no reason to believe the population is in decline.	In eastern Myanmar it is associated with limestone caves, and found in agricultural areas and secondary forest (P. Bates pers. comm.). In Cambodia it is found in caves (not necessarily limestone) in secondary forest and degraded habitat (G. Csorba pers. comm.). This species is tolerant to some degree of habitat disturbance.	There are no major threats to the whole population of this species, but as it is associated with cave, limestone mining and roost disturbance in much of region, e.g. in Myanmar, may affect some local populations.	The species is common across its range, with largest population size of 3,000 individuals was reported from a cave in Thailand (Soisook et al. 2008).	Stable	This species is found in Myanmar, China (Yunnan), Thailand, Cambodia, Lao PDR, Viet Nam, and Peninsular Malaysia.		Terrestrial	The species is found in a number of protected areas.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		malayanus	Bonhote	1903	0	In N. Annandale, Fasciculi Malayenses, Zool.	1:15	Malayan Horseshoe Bat	None.	Thailand, Jalor, Biserat.	Yunnan (China), Thailand, Burma, Cambodia, Laos, Vietnam, Peninsular Malaysia.	Not listed.	Least Concern	 megaphyllus species group. McFarlane and Blood (1986) suggested that characters used by Lekagul and McNeely (1977) to separate stheno and malayanus may not be reliable, but see Corbet and Hill (1992), Csorba and Jenkins (1998), and Hendrichsen (2001a, b). May contain cryptic diversity; see Chornelia et al. (2022).	Rhinolophus malayanus	1004709	23	Malayan Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	malayanus	Bonhote	1903	0						"Biserat, Jalor," Thailand.			malayanus Bonhote, 1903	NA	NA				China|Myanmar|Thailand|Laos|Vietnam|Cambodia|Malaysia	Asia	Indomalaya	LC	0	0	0	Rhinolophus_malayanus	0	sciname match	Rhinolophus_malayanus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_malayanus	1004709	23	Malayan Horseshoe Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	malayanus	Bonhote	0	Rhinolophus malayanus	Bonhote, J.L.J. 1903. Report on the mammals. Pp. 1â€“45 in Annandale, N. and Robinson, H.C. (eds.). Fasciculi Malayenses. Anthropological and Zoological Results of an Expedition to Perak and the Siamese Malay States, 1901-1902. Zoology. Part I. The University Press of Liverpool, Liverpool.	https://www.biodiversitylibrary.org/page/56338932	BMNH:Mamm:1903.2.6.83	holotype	https://data.nhm.ac.uk/object/970e3413-6b83-4cfb-a5ad-d37c63427dff	"Biserat, Jalor," Thailand.			NA	NA				China|Myanmar|Thailand|Laos|Vietnam|Cambodia|Malaysia	Asia	Indomalaya	LC	0	0	0	Rhinolophus_malayanus	0	sciname match	Rhinolophus_malayanus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		malayanus	Bonhote	1903	0	In N. Annandale, Fasciculi Malayenses, Zool.	1:15	Malayan Horseshoe Bat	None.	Thailand, Jalor, Biserat.	Yunnan (China), Thailand, Burma, Cambodia, Laos, Vietnam, Peninsular Malaysia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/19551/21978424/' target='_blank'>Least Concern</a>	megaphyllus species group. McFarlane and Blood (1986) suggested that characters used by Lekagul and McNeely (1977) to separate stheno and malayanus may not be reliable, but see Corbet and Hill (1992), Csorba and Jenkins (1998), and Hendrichsen (2001a, b). May contain cryptic diversity; see Chornelia et al. (2022).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus malayanus; Rhinolophus malayanus; Rhinolophus malayanus; Rhinolophus malayanus; Rhinolophus malayanus; Rhinolophus malayanus; malayanus; Malayan Horseshoe Bat; Rhinolophe de Malaisie; Malaya-Hufeisennase; Herradura de Malasia; Malayan Horseshoe Bat; Malayan Horseshoe Bat; Malayan Horseshoe Bat; R. malayanus
