http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1383	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Rhinolophus clivosus [synonym of]	N/A	Rhinolophus hillorum	Rhinolophus hillorum	Rhinolophus hillorum	Rhinolophus hillorum	Rhinolophus hillorum	Rhinolophus hillorum	Rhinolophus hillorum	Rhinolophus hillorum	Rhinolophus hillorum	Rhinolophus hillorum		[MSW3] ferrumequinum species group. Originally named as a subspecies of clivosus, but apparently distinct; see Cotterill (2002).; [HMW] Rhinolophus clivosus hillorum Koopman, 1989 , “ John Hegbe Farm near Zozoma, ca. 2 mi [= 3- 2 km ] SW Voinjama in Lofa County , extreme northwestern Liberia at ca. 500 m elevation.” Rhinolophus hillorum is in the ferrumequinum species group. It has been included as a subspecies of R clivosus but is now generally recognized as a distinct species. Monotypic.; [batnames2022]  ferrumequinum species group. Originally named as a subspecies of clivosus, but apparently distinct; see Cotterill (2002).; [IUCN] <p>Rhinolophus hillorum was originally recognized as a subspecies of R. clivosus in 1989 (Koopman 1989) and was elevated to species-level in 2002 (Cotterill 2002). There are at least two questionable records from ;Sudan and ;another from ;Sapoba ;Forest Reserve in Nigeria that are not included in this assessment. ;While the specimen from Sudan has characteristics similar to R. hillorum (Fahr 2013), it was reported as R. clivosus keniensis (Koopman 1975). The specimen from Sapoba ;Forest Reserve (Fedden ;and Macleod, 1986 – as ;R. ;clivosus ) is from a lowland habitat ;almost ;400 ;km ;away from the nearest mountain range, leading to calls for re-examination (Fahr 2013). Upon re-examination, the Sapoba specimen, it was re-assigned to R. cf. hillorum as it separated in a PCA of craniodental characteristics from other six specimens collected from Cross River National Park, Nigeria (Tanshi et al . 2021). </p>; [batnames2023]  ferrumequinum species group. Originally named as a subspecies of clivosus, but apparently distinct; see Cotterill (2002).; [batnames2025_1.7] ferrumequinum species group. Originally named as a subspecies of clivosus, but apparently distinct; see Cotterill (2002).														hillorum	<p>Rhinolophus hillorum was originally recognized as a subspecies of R. clivosus in 1989 (Koopman 1989) and was elevated to species-level in 2002 (Cotterill 2002). There are at least two questionable records from ;Sudan and ;another from ;Sapoba ;Forest Reserve in Nigeria that are not included in this assessment. ;While the specimen from Sudan has characteristics similar to R. hillorum (Fahr 2013), it was reported as R. clivosus keniensis (Koopman 1975). The specimen from Sapoba ;Forest Reserve (Fedden ;and Macleod, 1986 – as ;R. ;clivosus ) is from a lowland habitat ;almost ;400 ;km ;away from the nearest mountain range, leading to calls for re-examination (Fahr 2013). Upon re-examination, the Sapoba specimen, it was re-assigned to R. cf. hillorum as it separated in a PCA of craniodental characteristics from other six specimens collected from Cross River National Park, Nigeria (Tanshi et al . 2021). </p>			hillorum	hillorum			hillorum Koopman, 1989						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Rhinolophidae			Rhinolophus hillorum	Rhinolophus		hillorum	Koopman		1989		Amer. Mus. Novit.	2946		4		Upland Horseshoe Bat	Liberia, Lofa County, ca. 2 mi. (3 km) SW Voinjama, near Zozoma, John Hegbe Farm, 8°25'N, 9°35'W, ca. 500 m.	Guinea, Liberia, Nigeria, Cameroon.	IUCN 2003 – not evaluated; not considered in IUCN/SSC Action Plan (2001).		ferrumequinum species group. Originally named as a subspecies of clivosus, but apparently distinct; see Cotterill (2002).	885887A2FFE08A19F8B3FB9AF665D2BD	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Rhinolophidae.pdf.imf	hash://md5/7461ffdaffcf8a29ffccffa1ff85d963	294	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/88/58/87/885887A2FFC08A26F8B1FC3AF3DBD7EB.xml	Rhinolophus hillorum	Rhinolophidae	Rhinolophus	hillorum	Koopman	1989	Upland Horseshoe Bat @en | Rhinolophe de Lofa @fr | Hochland-Hufeisennase @de | Herradura montana @es | Hill's Horseshoe Bat @en	Rhinolophus clivosus hillorum Koopman, 1989 , “ John Hegbe Farm near Zozoma, ca. 2 mi [= 3- 2 km ] SW Voinjama in Lofa County , extreme northwestern Liberia at ca. 500 m elevation.” Rhinolophus hillorum is in the ferrumequinum species group. It has been included as a subspecies of R clivosus but is now generally recognized as a distinct species. Monotypic.	Scattered records from SE Guinea , N Liberia , S Nigeria , and W Cameroon with certainty; probably erroneous records from SE South Sudan , N Uganda , and S Kenya .	Head—body c . 64—71 mm , tail 30-41 mm , ear 21—24 mm , hindfoot 12-14 mm , forearm 52-57 mm ; weight 16- 5-25 g . The Upland Horseshoe Bat is large, with medium brown to grayish brown dorsum and paler venter. No orange morph has been reported. Males lack axillary tufts. Ears are short (37-44% of forearm length) and have 11-12 internal folds. Lancet is wide and almost parallel-sided at base; tip of lancet is very narrow, hairy, and slightly rounded; connecting process is narrow, high, rounded, ellipsoid in profile, much higher than sella tip, and liberally covered with hair; sella is naked, nearly parallel-sided (diverging slighdy toward top) and rounded at top; and horseshoe is relatively narrow at 8-3-9- 1 mm , does not completely cover muzzle, lacks lateral leaflets, and has distinct median emargination. Lower lip has single groove. Wings and uropatagium are blackish brown. Skull is exceptionally robust, with thick, broad zygomatic arches (zygomatic width is much larger than mastoid width); rostrum is very broad; nasal swellings are very low; frontal depression is very shallow; supraorbital crest is apparently not well defined; and sagittal crest is well developed anteriorly and moderately defined posteriorly. P2 is absent , allowing C1 and P4 to touch; P is also absent , so P and P4 are in contact; and P2 is one-half to two-thirds height of P4 Dental formula is then 11/2, C 1/1, P 1/2, M 3/3 (x2) = 28.	Montane forests, montane grasslands, dense rainforests, secondary forest, and gallery forests surrounded by savanna in highlands of Guinea , Liberia , and Cameroon and lowland and coastal rainforests surrounding these areas, primarily at elevations of 1400-1950 m .	The Upland Horseshoe Bat is insectivorous and probably occasionally forages by perch-hunting.	A pregnant Upland Horseshoe Bat was captured in lateJune. Litter size is one.	Upland Horseshoe Bats are nocturnal and can enter torpor while roosting during the day. They roost in caves, abandoned mines, and under bridges.	Upland Horseshoe Bats have been recorded roosting in small groups (ten individuals in one group in Liberia ) and sharing roosts with the Angolan Soft-furred Fruit Bat ( Lissonycteris angolensis ), the Bushveld Horseshoe Bat { simulator ), and the Guinean Horseshoe Bat { guineensis ).	Classified as Near Threatened on The IUCNRed List. The Upland Horseshoe Bat is known from very few scattered records and is considered rare, threatened by habitat loss and destruction from agricultural expansion and planned large-scale mining. It is possibly threatened by exploitation for bushmeat.	ACR (2018) | Csorba et al. (2003) | Cotterill (2002a) | Fahr (2013f) | Happold (1987) | Jacobs et al. (2010) | Koopman (1989) | Koopman et al. (1995) | Monadjem, Richards & Denys (2016)	https://zenodo.org/record/3749964/files/figure.png	28 . Upland Horseshoe Bat Rhinolophus hillorum French: Rhinolophe de Lofa / German: Hochland-Hufeisennase / Spanish: Herradura montana Other common names: Hill's Horseshoe Bat Taxonomy. Rhinolophus clivosus hillorum Koopman, 1989 , “ John Hegbe Farm near Zozoma, ca. 2 mi [= 3- 2 km ] SW Voinjama in Lofa County , extreme northwestern Liberia at ca. 500 m elevation.” Rhinolophus hillorum is in the ferrumequinum species group. It has been included as a subspecies of R clivosus but is now generally recognized as a distinct species. Monotypic. Distribution. Scattered records from SE Guinea , N Liberia , S Nigeria , and W Cameroon with certainty; probably erroneous records from SE South Sudan , N Uganda , and S Kenya . Descriptive notes. Head—body c . 64—71 mm , tail 30-41 mm , ear 21—24 mm , hindfoot 12-14 mm , forearm 52-57 mm ; weight 16- 5-25 g . The Upland Horseshoe Bat is large, with medium brown to grayish brown dorsum and paler venter. No orange morph has been reported. Males lack axillary tufts. Ears are short (37-44% of forearm length) and have 11-12 internal folds. Lancet is wide and almost parallel-sided at base; tip of lancet is very narrow, hairy, and slightly rounded; connecting process is narrow, high, rounded, ellipsoid in profile, much higher than sella tip, and liberally covered with hair; sella is naked, nearly parallel-sided (diverging slighdy toward top) and rounded at top; and horseshoe is relatively narrow at 8-3-9- 1 mm , does not completely cover muzzle, lacks lateral leaflets, and has distinct median emargination. Lower lip has single groove. Wings and uropatagium are blackish brown. Skull is exceptionally robust, with thick, broad zygomatic arches (zygomatic width is much larger than mastoid width); rostrum is very broad; nasal swellings are very low; frontal depression is very shallow; supraorbital crest is apparently not well defined; and sagittal crest is well developed anteriorly and moderately defined posteriorly. P2 is absent , allowing C1 and P4 to touch; P is also absent , so P and P4 are in contact; and P2 is one-half to two-thirds height of P4 Dental formula is then 11/2, C 1/1, P 1/2, M 3/3 (x2) = 28. Habitat. Montane forests, montane grasslands, dense rainforests, secondary forest, and gallery forests surrounded by savanna in highlands of Guinea , Liberia , and Cameroon and lowland and coastal rainforests surrounding these areas, primarily at elevations of 1400-1950 m . Food and Feeding. The Upland Horseshoe Bat is insectivorous and probably occasionally forages by perch-hunting. Breeding. A pregnant Upland Horseshoe Bat was captured in lateJune. Litter size is one. Activity patterns. Upland Horseshoe Bats are nocturnal and can enter torpor while roosting during the day. They roost in caves, abandoned mines, and under bridges. Movements, Home range and Social organization. Upland Horseshoe Bats have been recorded roosting in small groups (ten individuals in one group in Liberia ) and sharing roosts with the Angolan Soft-furred Fruit Bat ( Lissonycteris angolensis ), the Bushveld Horseshoe Bat { simulator ), and the Guinean Horseshoe Bat { guineensis ). Status and Conservation. Classified as Near Threatened on The IUCNRed List. The Upland Horseshoe Bat is known from very few scattered records and is considered rare, threatened by habitat loss and destruction from agricultural expansion and planned large-scale mining. It is possibly threatened by exploitation for bushmeat. Bibliography. ACR (2018), Csorba et al. (2003), Cotterill (2002a), Fahr (2013f), Happold (1987), Jacobs et al. (2010), Koopman (1989), Koopman et al. (1995), Monadjem, Richards & Denys (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Rhinolophidae	Rhinolophus hillorum	Rhinolophus		hillorum	Koopman	1989	0	Amer. Mus. Novit.	######	Upland Horseshoe Bat	None.	Liberia, Lofa County, ca. 2 mi. (3 km) SW Voinjama, near Zozoma, John Hegbe Farm, 8°25'N, 9°35'W, ca. 500 m.	Guinea, Liberia, Nigeria, Cameroon.	Not listed.	Near Threatened	 ferrumequinum species group. Originally named as a subspecies of clivosus, but apparently distinct; see Cotterill (2002).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Rhinolophus hillorum	23	Upland Horseshoe Bat	Hill's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	RHINOLOPHIDAE	NA	NA	Rhinolophus	NA	hillorum	Koopman	1989	0	Rhinolophus_clivosus_hillorum	Koopman, K. F. (1989). Systematic notes on Liberian bats. American Museum Novitates, 2946, 4.	http://digitallibrary.amnh.org/bitstream/handle/2246/5100//v2/dspace/ingest/pdfSource/nov/N2946.pdf?sequence=1&isAllowed=y	AMNH 257044		"John Hegbe Farm near Zozoma, ca. 2 mi [= 3.2 km] SW Voinjama in Lofa County, extreme northwestern Liberia at ca. 500 m elevation."			hillorum Koopman, 1989	NA	NA	Guinea|Liberia|Nigeria|Cameroon	Africa	Afrotropic	NT	0	0	0	Rhinolophus_hillorum	0	sciname match	Rhinolophus_hillorum	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	44782	Rhinolophus hillorum	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	RHINOLOPHIDAE	Rhinolophus	hillorum	Koopman, 1989	<p>Rhinolophus hillorum was originally recognized as a subspecies of R. clivosus in 1989 (Koopman 1989) and was elevated to species-level in 2002 (Cotterill 2002). There are at least two questionable records from ;Sudan and ;another from ;Sapoba ;Forest Reserve in Nigeria that are not included in this assessment. ;While the specimen from Sudan has characteristics similar to R. hillorum (Fahr 2013), it was reported as R. clivosus keniensis (Koopman 1975). The specimen from Sapoba ;Forest Reserve (Fedden ;and Macleod, 1986 – as ;R. ;clivosus ) is from a lowland habitat ;almost ;400 ;km ;away from the nearest mountain range, leading to calls for re-examination (Fahr 2013). Upon re-examination, the Sapoba specimen, it was re-assigned to R. cf. hillorum as it separated in a PCA of craniodental characteristics from other six specimens collected from Cross River National Park, Nigeria (Tanshi et al . 2021). </p>	10000000	Rhinolophus hillorum	Near Threatened		2010	2009-09-24 00:00:00 UTC	3.1	English	<p>Listed as Near Threatened since although its Extent of Occurrence is probably less than 20,000 km<sup>2</sup> and its habitat is declining from the loss of montane habitat, especially in northern Liberia and southern Guinea, making the species close to qualifying for Vulnerable.</p>	Populations of this species have been recorded from caves in both montane and lowland tropical moist forest. It is not known if the species can persist in disturbed habitats.	Colonies are threatened by general deforestation, often resulting from logging and mining operations, and overharvesting for the bushmeat trade.	The species appears to have a small population, but further research is needed to confirm this.	Decreasing	This species is largely endemic to West and Central Africa, with a single questionable record from northern Uganda. It has been recorded from a few small, and disjunct, colonies in Liberia (including Voinjama; Tokadeh; and the Wonegizi Mountains), Guinea, Nigeria (Sapoba Forest Reserve) and Cameroon (including Lake Manenguba and the Bali Forest Reserve).		Terrestrial	There appear to be no direct conservation measures in place. It is not known if the species is present in any protected areas, but it has been recorded from the Sapoba Forest Reserve in Nigeria and the Bali Forest Reserve in Cameroon (Cotterill 2002). There is a need to identify and protect important areas for this species. Further research is needed into the species distribution, including the location of any additional colonies.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Rhinolophidae	Rhinolophus		hillorum	Koopman	1989	0	Amer. Mus. Novit.	########	Upland Horseshoe Bat	None.	Liberia, Lofa County, ca. 2 mi. (3 km) SW Voinjama, near Zozoma, John Hegbe Farm, 8°25'N, 9°35'W, ca. 500 m.	Guinea, Liberia, Nigeria, Cameroon.	Not listed.	Near Threatened	 ferrumequinum species group. Originally named as a subspecies of clivosus, but apparently distinct; see Cotterill (2002).	Rhinolophus hillorum	1004692	23	Upland Horseshoe Bat	Hill's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Rhinolophidae	NA	NA	Rhinolophus	NA	hillorum	Koopman	1989	0	Rhinolophus_clivosus_hillorum	Koopman, K. F. (1989). Systematic notes on Liberian bats. American Museum Novitates, 2946, 4.	http://digitallibrary.amnh.org/bitstream/handle/2246/5100//v2/dspace/ingest/pdfSource/nov/N2946.pdf?sequence=1&isAllowed=y	AMNH 257044		"John Hegbe Farm near Zozoma, ca. 2 mi [= 3.2 km] SW Voinjama in Lofa County, extreme northwestern Liberia at ca. 500 m elevation."			hillorum Koopman, 1989	NA	NA				Guinea|Liberia|Nigeria|Cameroon	Africa	Afrotropic	NT	0	0	0	Rhinolophus_hillorum	0	sciname match	Rhinolophus_hillorum	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Rhinolophus_hillorum	1004692	23	Upland Horseshoe Bat	Hill's Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Rhinolophidae	NA	NA	Rhinolophus	NA	hillorum	Koopman	0	Rhinolophus clivosus hillorum	Koopman, K.F. 1989-06-28. Systematic notes on Liberian bats. American Museum Novitates 2946:1-11.	https://www.biodiversitylibrary.org/page/62290607	AMNH M-257044	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-257044	"John Hegbe Farm near Zozoma, ca. 2 mi [= 3.2 km] SW Voinjama in Lofa County, extreme northwestern Liberia at ca. 500 m elevation."	8.416667	-9.58333	NA	NA				Guinea|Liberia|Nigeria|Cameroon	Africa	Afrotropic	VU	0	0	0	Rhinolophus_hillorum	0	sciname match	Rhinolophus_hillorum	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Rhinolophidae	Rhinolophus		hillorum	Koopman	1989	0	Amer. Mus. Novit.	########	Upland Horseshoe Bat	None.	Liberia, Lofa County, ca. 2 mi. (3 km) SW Voinjama, near Zozoma, John Hegbe Farm, 8°25'N, 9°35'W, ca. 500 m.	Guinea, Liberia, Nigeria, Cameroon.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/44782/203829273/' target='_blank'>Vulnerable</a>	ferrumequinum species group. Originally named as a subspecies of clivosus, but apparently distinct; see Cotterill (2002).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Rhinolophus hillorum; Rhinolophus hillorum; Rhinolophus hillorum; Rhinolophus hillorum; Rhinolophus hillorum; Rhinolophus hillorum; hillorum; Upland Horseshoe Bat; Rhinolophe de Lofa; Hochland-Hufeisennase; Herradura montana; Hill's Horseshoe Bat; Upland Horseshoe Bat; Hill's Horseshoe Bat; Upland Horseshoe Bat; Upland Horseshoe Bat; R. hillorum